Trigonostemon villosus var. cordatus

Yu, R. - Y. & Welzen, P. C. van, 2018, A taxonomic revision of Trigonostemon (Euphorbiaceae) in Malesia, Blumea 62 (3), pp. 179-229 : 220

3. Leaves pubescent above. — Sabah ..... c. var. cordatus a. var. villosus — Fig. 16 ; Map 13

Leaves: petiole 3–8 cm long; blade oblong, 12–18 by 3.5–5.5 cm, glabrous to glabrescent (rarely villose) above and villose beneath. Inflorescences bisexual or unisexual (often because staminate flowers fallen), thyrsoid, 6–16 cm long, one pistillate flower and a few staminate flowers per node, often more than 4 pistillate flowers present at one time. Staminate flowers: sepals often without a subapical gland; petals elliptic, honey mark unseen. Pistillate flowers: sepals lanceolate, apex acuminate, margin often serrate, always accrescent when fruiting; ovary villose. Fruits villose.

Distribution — Malay Peninsula (endemic).

Habitat & Ecology — Understorey in primary forests to logged-over areas, often on hill sides or along rivers, growing in swamps forests, on peaty soil. Flowering: January, May, June, October, November; fruiting: January, April to June, October, November.

Figures

Fig. 16 Trigonostemon villosus Hook.f.var.villosus (a, c, f), Forest Research Institute Malaysia,Kepong, Malay Peninsula and Trigonostemon villosus Hook.f. var. borneensis (Merr.) Airy Shaw (b, d, e, g–m), Segaliud Lokan Forest Reserve, Sabah, Malaysia. a, b. Growing habit; c, d. top view of the tree; e. bark and reddish exude; f, g. pistillate inflorescence; h, i. staminate flower; j, k. pistillate flower; l, m. fruit. — Photos by Ren-Yong Yu.