Trigonostemon beccarii Ridl.
publication ID |
https://doi.org/10.3767/blumea.2018.62.03.04 |
persistent identifier |
https://treatment.plazi.org/id/03CA2B5B-F74B-7356-FFDC-F9E7FB1E9DF9 |
treatment provided by |
Felipe |
scientific name |
Trigonostemon beccarii Ridl. |
status |
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4. Trigonostemon beccarii Ridl. View in CoL — Fig. 3 View Fig ; Map 3
Trigonostemon beccarii Ridl.(1925) 89;Jabl.(1963) 165; Airy Shaw (1981) 353. — Type: Beccari PS 965 (K, L), Sumatra, Padang, Sungei Buluh .
Trigonostemon longisepalus Ridl.(1925) 89;Jabl.(1963) 165. — Type: Brooks 8274 (holo K), Sumatra, Lubok Tandai .
Shrubs; flowering branches 1.6–4.5 mm diam, sometimes hollow. Outer bark 0.1–0.2 mm thick, brownish to greyish, smooth; inner bark 0.1–0.2 mm thick, dark reddish brown; wood white to pale reddish brown. Stipules falcate, 0.5–0.7 mm long, glabrous. Leaves: petiole 0.25–1.5 cm long, grooved above, glabrous to glabrescent; blade oblong to oblanceolate, 8–28 by 1.5–7 cm, base cuneate, adaxially 2 glands present but caducous, margin slightly distantly serrate, apex acuminate, upper side dark greenish, lower side pale green, both sides glabrous; midrib slightly raised on both sides, nerves 10–14 pairs, very slightly raised beneath, often obscure, small nerves reticulate, obscure. Inflorescences bisexual, axillary to subterminal, often 1 pistillate flower on the top and few short scorpioid cymes (glomerules) of staminate flowers along the
Map 3 Distribution of Trigonostemon balgooyi R.Y.Yu & Welzen (●); T. beccarii Ridl. (■); T. calcicolus (R.I.Milne) R.Y.Yu & Welzen (★); T. diffusus Merr. (▲); T. dipteranthus Airy Shaw (▼); T. filiformis Quisumb. (◆).
rachis below; rachis 3–12 cm long, 0.5–0.7 mm diam, glabrous; bracts elliptic to lanceolate, 0.7–1.5 by 0.3–0.6 mm, glabrous to pubescent, apex acuminate to rounded. Staminate flowers c. 4.5 mm diam; pedicel c. 3.5 mm long, c. 0.3 mm diam, glabrescent; sepals elliptic, c. 1 by 0.7 mm, apex rounded, slightly ciliate on margin, glabrescent outside; petals elliptic, 1.5–2.2 by 1.2–1.5 mm, margin entire, apex rounded, glabrous outside, rough and papillose inside; disc lobes semi-orbicular to nearly lanceolate, c. 0.5 by 0.5 mm, apex rounded, glabrous; stamens 5, androphore c. 0.5 mm long, 5-cleft at the top, anthers ovoid to ellipsoid, c. 0.6 mm long, free, thecae slightly divaricate at apex, connectives apically with numerous droplets (expanded cells) with secretion. Pistillate flowers 2.5–3.5 mm diam; pedicel slightly thickened towards apex, c. 6 mm long, apically 0.7–1 mm diam when flowering, accrescent to 2.5 cm long and apically 2 mm diam when fruiting, glabrous to glabrescent; sepals ovate, 1.4–1.7 by 1–1.5 mm, apex acute, slightly pubescent to glabrescent outside; petals (remnants) nearly elliptic, c. 2.1 by 0.8 mm, glabrous outside, rough and scurfy inside; disc lobes as staminate flowers; ovary 1–1.5 mm diam, densely pubescent; styles indiscernible; stigmas completely bifid, each arm c. 1 mm long, thickened at base, slender and often bent upward. Fruits c. 1 cm diam, appressed pubescent; sepals persistent but not accrescent; wall c. 0.5 mm thick, exocarp partly detaching; columella c. 6.5 mm long. Seeds 4.2–5.5 mm diam, pale brown when dry, hilum heart-shaped or rhombic, c. 1.2 by 1–1.2 mm.
Distribution — Sumatra (endemic).
Habitat & Ecology — Flowering and fruiting: September.
Note — A distinct species with short petiole, inflorescences of short glomerules along an erect rachis and 5 stamens in the staminate flowers. Ridley (1925) described T. beccari and T. longisepalus as separate species but he did admit that these two species were possibly based on two sexes of one plant. Our recent collection R. Y. Yu 169 (L), from the Bogor Botanical Gardens (XI.B.XVII.270-270a, originally from Sumatra), bears both the same staminate and pistillate structures as described in Ridley (1925), and thereby confirms the conspecific status of T. beccari and T. longisepalus .
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