Alopoglossus carinicaudatus, (COPE, 1876)

Ribeiro-Júnior, Marco Antonio, Choueri, Erik, Lobos, Simon, Venegas, Pablo, Torres-Carvajal, Omar & Werneck, Fernanda, 2019, Eight in one: morphological and molecular analyses reveal cryptic diversity in Amazonian alopoglossid lizards (Squamata: Gymnophthalmoidea), Zoological Journal of the Linnean Society 190, pp. 227-270 : 253-254

ALOPOGLOSSUS CARINICAUDATUS ( COPE, 1876)

[ FIGS 2–4 ( A. ANGULATUS ‘ ECUADOR’), 7I, J, 14, 15;

TABLES 1 AND 2]

Lepidosoma carinicaudatum Cope (1876: 160) .

Leposoma carinicaudatum O’Shaughnessy (1881: 233) .

Alopoglossus carinicaudatus Boulenger (1885: 384) ; Ruthven (1924: 3); Ruibal (1952: 508, part).

Pantodactylus carinicaudatus Burt & Burt (1931: 358) . Alopoglossus angulatus Köhler et al. (2012: 176 , part); Ribeiro-Júnior (2018: 32, part).

Holotype: ANSP 11371 , male, Marañon, Orton Expedition. Type locality restricted by Ruibal (1952) to Valley of the Marañon River , Peru, but here corrected and restricted to Santa Rosa de Otas , Department of Napo, Ecuador (upper Napo River ; 0°58 ′ 41.2 ″ S, 77°27 ′ 4.4 ″ W; see comments in Remarks below) ( Figs 14, 15).

Material examined: Six referred specimens for morphological analyses from Ecuador and Peru and 17 samples for molecular analyses from Ecuador (see Supporting Information; Appendix S2, Table S1).

Diagnosis

Alopoglossus carinicaudatus is distinguished from all other species of Alopoglossus by the combination of the following characters: (1) scales on the sides of the neck non-granular, keeled, imbricate (at least medial and posterior ones phylloid), in 11 or 12 transverse rows; (2) three pairs of chin shields; (3) third pair of chin shields irregularly quadrangular (anterior and posterior margins almost parallel), with straight posterior margins, separated from gulars by a row of small scales; (4) absence of a pair of medial enlarged pregular scales; (5) scales on gular region subequal in shape and size: lateral, anterior, medial and posterior ones strongly keeled, pointed, phylloid; anteriormost transverse row feebly keeled; (6) ventral scales mucronate, imbricate, with bluntly pointed posterior margins, varying from strongly keeled on anterior and lateral surfaces to feebly keeled on medial and posterior surfaces; and (7) total number of femoral pores in males 22–26.