Leonurus cardiaca, L.

Tutin, T. G., Heywood, V. H., Burges, N. A., Moore, D. M., Valentine, D. H., Walters, S. M. & Webb, D. A., 1972, Flora Europaea. Volume 3. Diapensiacea to Myoporaceae, Cambridge: Cambridge University Press : 149

1. L. cardiaca L. , Sp. Pl. 584 (1753)

(incl. L. glaucescens Ledeb. , L. quinquelobatus Usteri , L. tataricus L. , L. villosus Desf. ex Sprengel ).

Glabrous to villous perennial 30-200 cm. Leaves 3-1 2x2-4 cm, the lower digitately 5- to 7-lobed, the lobes toothed or shallowly lobed, the upper shallowly to deeply 3-fid. Calyx 3-5-8 mm, the tube very prominently 5-veined; teeth almost as long as tube, the lower deflexed. Corolla 8-12 mm, distinctly exceeding calyx, white or pale pink, sometimes with purple spots; upper lip densely villous on back; tube with a ring of hairs inside. Nutlets hispid at apex. I n — 18. M ost o f Europe, but absent from the extreme north, the islands, and much o f the Mediterranean region. Al Au Be Bu Cz Da Fe Ga Ge G r He Ho Hs Hu It Ju N o Po Rm Rs (N, B, C, W, K, E) Su Tu [Br].

Plants in the eastern part of the range usually have densely hairy stems with patent hairs 1-2 mm, leaves and bracts deeply divided and calyx 4-5-5(-8) mm, and are often regarded as a separate species or subspecies ( subsp. villosus (Desf. ex Sprengel) Hyl. , Uppsala Univ. Ârsskr. 1945(7): 273 (1945)); those in the central and western part usually have glabrous or sparsely hairy stems with recurved hairs 0-5 mm, leaves and bracts shallowly lobed and calyx 3-5-4(-5) mm. Sparsely hairy variants, however, are sometimes found in E. Europe and occasionally plants with a mixture of patent and recurved hairs occur.