Anchusa L.
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publication ID |
https://doi.org/10.5281/zenodo.305475 |
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persistent identifier |
https://treatment.plazi.org/id/03C287E6-FF09-559E-EDB1-665FF3B41982 |
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treatment provided by |
Plazi |
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scientific name |
Anchusa L. |
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21. Anchusa L. View in CoL 1
(incl. Lycopsis L. )
Annual, biennial or perennial herbs, rarely woody at the base. Flowers in axillary and terminal cymes, bracteate usually throughout. Calyx lobed from | to almost to the base. Corolla purple, blue, yellow or white, with cylindrical, straight or curved tube and rotate to campanulate limb divided into equal or unequal lobes, with 5 ovate or oblong, papillose or hairy scales in throat. Stamens included or slightly exserted, variously inserted. Style included; stigma capitate. Nutlets ovoid to reniform or hemispherical, erect or oblique, reticulate or rugose, usually more or less tuberculate, with a thickened collar-like ring at the base.
Literature: M. Guçuleac, Bui. Fac. §ti. Cernäufi 1: 72-123 (1927); Feddes Repert. 26: 286-322 (1929). W. Greuter, Candollea 20: 192-210 (1965).
1 Corolla yellow
2 Bracts exceeding flowers; nutlets 4-5 x 2-3 mm, erect
21. aegyptiaca 2 Bracts shorter than flowers; nutlets c. 2 x 4 mm, oblique
3 Leaves mostly more than 10 mm wide 10. officinalis 3 Leaves mostly less than 10 mm wide 3. ochroleuca 1 Corolla blue, violet, purplish or whitish
4 Corolla zygomorphic, the tube curved, the limb oblique, with 5 unequal lobes
5 Filaments inserted at or below middle of corolla-tube;
cymes bracteate throughout 22. arvensis 5 Filaments inserted above middle of corolla-tube; cymes bracteate only at base
6 Nutlets densely rugulose between the primary reticulations; lobes of style obliquely truncate 23. eretica 6 Nutlets smooth between the primary reticulations; lobes of style ± acute 24. variegata 4 Corolla actinomorphic
7 Corolla-tube much shorter than limb
8 Annual; inflorescence simple or once branched 16. macedonica 8 Perennial; inflorescence paniculate
9 Caespitose; leaves up to 7 mm wide 20. serpentinicola 9 Not caespitose; most leaves at least 10 mm wide 19. barrelieri 7 Corolla-tube equalling or longer than limb
10 Calyx divided more or less to the base
11 Nutlets longer than wide
12 Nutlets c. 4 mm; corolla-limb 5-7 mm in diameter
13. thessala 12 Nutlets at least 6 mm; corolla-limb (8-)10-15 mm in diameter 11. azurea 11 Nutlets wider than long, or hemispherical
13 Caespitose perennial with stems not more than 10 cm
4. cespitosa 13 Annual, biennial or perennial, not caespitose, with stems more than 10 cm
14 Style 2-3 times as long as calyx; corolla-scales with subglobose papillae 17. stylosa 14 Style less than twice as long as calyx; corolla-scales hairy or with cylindrical papillae
15 Corolla-tube 12-15 mm, about twice as long as calyx
15. macrosyrinx 15 Corolla-tube less than 10 mm, not more than l i times as long as calyx
16 Pedicels equalling or exceeding calyx at anthesis
18. spruneri 16 Pedicels shorter than calyx at anthesis
17 Cymes capitate, with lower bracts much exceeding calyx; nutlets ±hemispherical 12. aggregata 17 Cymes ± elongate, with bracts not exceeding calyx; nutlets obliquely ovoid
18 Annual; cymes with c. 5 flowers 14. pusilla
18 Usually perennial; cymes with more than 5
flowers 10. officinalis 10 Calyx lobed to not more than j
19 Calyx-teeth acute
20 Most hairs on leaves tubercle-based 7. granatensis 20 Most hairs on leaves not tubercle-based 10. officinalis 19 Calyx-teeth obtuse
21 Bracts equalling or exceeding calyx 9. crispa 21 Bracts shorter than calyx
22 Stamens inserted at top of corolla-tube and overlapping scales
23 Plant with strongly tubercle-based setae 8. calcarea 23 Plant pubescent and with setae which are mostly not tubercle-based 5. undulata 22 Stamens inserted below top of tube and not or only slightly overlapping scales
24 Plant subglabrous or with setae less than 1 mm;
nutlets subreniform 2. gmelinii 24 Plant with hairs or setae more than 1 mm; nutlets obliquely ovoid
25 Plant with dense, stout, patent, white, mostly tubercle-based setae 6. sartorii 25 Plant pubescent and with setae which are mostly not conspicuously white or tubercle-based
26 Stamens inserted at middle of tube and not nearly reaching scales 5. undulata 26 Stamens inserted above middle of tube and reaching base of scales 1. leptophylla
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