Empis (Enoplempis) yakimensis, Sinclair & Brooks & Cumming, 2025

Sinclair, Bradley J., Brooks, Scott E. & Cumming, Jeffrey M., 2025, Revision of the western Nearctic species of Empis subgenus Enoplempis (Diptera: Empididae), Zootaxa 5615 (1), pp. 1-200 : 182-188

Empis (Enoplempis) yakimensis sp. nov.

urn:lsid:zoobank.org:act:802274CD-A7DE-4072-A5A2-7D62DC413651

( Figs 308 , 349 , 352 )

Type material. HOLOTYPE ♂, labelled: “[ USA] WASHINGTON: Yakima / Co., 8 mi SW Tieton / RS, Snoqualmie NP/ Bear Crk [46°42′N 120°45′W], VI-(11-12)-73”; “ D. Corredor / Collector”; “ HOLOTYPE / Empis (Enoplempis) / yakimensis Sinclair ,/ Brooks & Cumming [red label]” [dissected] ( WSU).

Diagnosis. This species is distinguished from other species of the subgenus by the grey body and males are dichoptic with hindleg unmodified, slender hind tarsomere 1, hind tibia without long anterior setae ( Fig. 349 ), cercus with straight dorsal margin, and phallus with swollen basal third, strongly bent at apical third, apex not expanded with subapical roughened surface ( Fig. 352 ).

Description. Wing length 4.7 mm. Male. Similar to E. (En.) californica sp. nov., except as follows: head dichoptic. Frons slightly narrower than width of anterior ocellus. Antenna with postpedicel long and tapered, 5× longer than basal width.

Thorax with scutum with fine biserial acr; dc slightly longer, stronger than acr anteriorly, increasing in length and stronger posteriorly; 2 posterior npl, with 1–2 shorter anterior npl.

Legs long, slender, yellow, apical tarsi darker; coxae without dense greyish pruinescence. Femora without white ventral pile. Fore femur with anteroventral and posteroventral row of setae, shorter than half width of femur. Fore tibia with 1–2 erect anterodorsal setae; anterior face with short appressed setae. Fore tarsomere 1 slender [? flattened, difficult to determine]; without long dorsal setae; tarsomeres 1–5 with biserial row of spine-like ventral setae; tarsomeres 1–4 with apicolateral pair of spine-like setae, longer than width of segment. Mid femur similar to fore femur. Mid tibia without erect setae. Mid tarsomere 1 with stronger ventral spine-like setae, otherwise similar to fore tarsus. Hind femur slender, without modified setae and processes ( Fig. 349 ); row of fine anteroventral and posteroventral setae, length shorter than width of femur. Hind tibia without processes or modified setae; 2– 4 anterodorsal and posterodorsal setae ( Fig. 348 ). Hind tarsus slender with several long preapical dorsal setae, otherwise similar to mid tarsus.

Wing with R 4+5 unbranched on left wing, R 4 short stub on right wing.

Terminalia ( Fig. 352 ): cercus straight along dorsal margin, tapered on apical half; setae shorter than width of cercus. Subepandrial process extending on either side of phallus. Epandrial lamella elongate, longer than height; linear with posterior margin slightly upcurved, rounded, bearing long setae ventrally, shorter than width of epandrium. Hypandrium narrow, strap-like; without setae. Phallus with basal third strongly expanded, tapered apically, with strong bend near apical third; apical third straight, apex emerging beyond cerci; apex narrow with elongate opening;

roughened surface subapically; ejaculatory apodeme more than half-length of epandrium, Y-shaped, with lateral apodemes positioned near ventral margin with short medial apodeme.

Female. Unknown.

Geographical distribution and seasonal occurrence ( Fig. 308 ). Empis (En.) yakimensis sp. nov. is known only from the holotype collected in Yakima County, Washington in June .

Etymology. The species name is in reference to the type locality, Yakima County, Washington.

Nuptial gift presentation. Form unknown.

Key to males of Empis (Enoplempis) from the Rocky Mountains westward

1 Entirely yellow, including thorax ( Figs 107 , 144 , 291 )........................................................ 2

- At least thorax dark, usually with grey pruinescence (e.g., Figs 41 , 62 , 143 , 164 , 179 , 299 , 332 )....................... 9

2 Hind tibia geniculate at base; male hindlegs asymmetrical ( Figs 210–214, 216, 217 ) [ mira group]..................... 3

- Hind tibia not geniculate at base; male hindlegs symmetrical ( Figs 108, 109 , 145–147 , 291 ).......................... 7

3 Scutum usually with single median brown vitta (sometimes not clearly visible due to preservation); right hind femur with two elongate posteroventral tufts, as long as width of femur ( Fig. 213 )..... E. (En.) williamturneri Sinclair, Brooks & Cumming - Scutum usually with or without several vittae; right hind femur with one elongate posteroventral tuft as long as width of femur ( Fig. 212 )........................................................................................... 4

4 Scutum with row of dark spots at base of the dorsocentral setae................................................. 5

- Scutum with dark or pale vitta at base of the dorsocentral setae................................................. 6

5 Right hind femur with anteroventral digitiform lobe on apical third, clothed in short setae ( Fig. 211 ); right hind tibia with subapical swelling directed vertically, with black thorn-like setae ( Figs 211, 212 )................... E. (En.) mira (Bigot)

- Right hind femur without anteroventral digitiform lobe ( Fig. 214 ); right hind tibia with subapical swelling directed horizontally, without black thorn-like setae ( Fig. 214 ).............................. E. (En.) submira Sinclair, Brooks & Cumming

6 Right hind femur without anteroventral setae opposite subapical swelling on tibia ( Fig. 217 ); mid femur with long posteroventral setae at most slightly longer than width of femur....................... E. (En.) winkleri Sinclair, Brooks & Cumming

- Right hind femur with long anteroventral setae opposite subapical swelling on tibia ( Fig. 216 ); mid femur with long posteroventral setae more than twice width of femur ( Fig. 215 )..................... E. (En.) macdonaldi Sinclair, Brooks & Cumming

7 Hind femur and tibia without processes, setal combs or rows of modified setae ( Fig. 291 ).......... E. (En.) cerina sp. nov.

- Hind femur and tibia with processes, setal combs and/or rows of modified setae ( Figs 108, 109 , 145–147 )............... 8

8 Hind femur without anteroventral paint brush-like process; hind tibia with subbasal swelling and bend, bearing anteroventral lobe without setae, slightly arched with flattened anterior face; long dorsal setae evenly distributed along entire hind tibia (Figs 145–147) [ delumbis group, in part].......................................... E. (En.) delumbis Melander [in part]

- Hind femur with anteroventral paint brush-like process; hind tibia without subbasal swelling, straight, bearing anteroventral lobe with setae; long dorsal setae confined mostly to apical half ( Figs 107–109 ) [ canaster group, in part]................................................................................................... E. (En.) safra sp. nov.

9 Hind femur and tibia without processes, setal combs or rows of modified setae; hind femur at most with slender setulae beneath ( Figs 179–182 , 286, 288 ).............................................................................. 10

- Hind femur and tibia with processes, setal combs and/or rows of modified setae (e.g., Figs 16 , 72 , 134 , 156 , 178 , 189 , 205 , 219 , 231 , 258 , 334 )....................................................................................... 21

10 Face and scape with numerous setae, longer than scape; palpus with ventral setae longer than palpus ( Figs 179 , 181 ) [ geneatis group, in part]....................................................................................... 11

- Face bare; scape and palpus without elongate setae.......................................................... 13

11 Very large species, wing length more than 6 mm; cercus broadly rounded dorsally ( Fig. 187 )...................................................................................................... E. (En.) subgeneatis sp. nov.

- Smaller species, wing length 5 mm or less; cercus prolonged dorsally or projecting horizontally ( Figs 185 , 186 )......... 12

12 Hairy species ( Fig. 181 ); cercus broader than epandrium, prolonged dorsally ( Fig. 185 )........ E. (En.) geneatis (Melander)

- Shorter haired species ( Fig. 179 ); cercus narrower than epandrium, projecting horizontally ( Fig. 186 ).................................................................................................... E. (En.) oharai sp. nov.

13 Hind tibia with more than 6 pairs of dorsal setae ( Figs 286, 288, 290 , 319 , 331 )................................... 14

- Hind tibia with 4–6 pairs of dorsal setae ( Figs 304 , 320 , 348, 349 )............................................. 18

14 Eyes holoptic, frons narrower than width of anterior ocellus ( Fig. 319 ).......................................... 15

- Eyes dichoptic, frons as wide or wider than width of anterior ocellus............................................ 17

15 Cercus with deep cleft near mid-length; phallus sinuous on apical half ( Figs 294, 295 )....... E. (En.) blodgettensis sp. nov.

- Cercus with dorsal margin straight; phallus gradually arched on apical half ( Figs 325 , 338, 339 )...................... 16

16 Apex of phallus expanded, with roughened surface ( Fig. 339 )........................... E. (En.) oregonensis sp. nov.

- Apex of phallus slender, with subapical roughened surface, opposite cerci ( Fig. 325 )........... E. (En.) holoptica sp. nov.

17 Hind tibia with long, slender anterior setae ( Fig. 290 ); cercus expanded dorsally, with sloping posterior face and posteroventral elongation ( Fig. 296 )............................................................ E. (En.) californica sp. nov.

- Hind tibia without long, slender anterior setae ( Fig. 286 ); cercus not greatly expanded dorsally, posterior face projecting and tapered ( Fig. 292 )................................................................... E. (En.) aphela sp. nov.

18 Eyes dichoptic, narrowly separated by less than width of anterior ocellus................... E. (En.) yakimensis sp. nov.

- Eyes holoptic, closely approximated..................................................................... 19

19 Epandrium strongly, narrowly upturned apically; phallus with broad apex, with roughened surface encircling tip ( Fig. 306 ).............................................................................. E. (En.) delnortensis sp. nov.

- Epandrium broadly upturned apically; phallus with narrow apex, with subapical roughened surface ( Figs 326, 327 , 340, 341 ) .................................................................................................. 20

20 Cercus with truncate apex; phallus without distinct upcurved lip at edge of opening ( Figs 340, 341 ).................................................................................................... E. (En.) susanae sp. nov.

- Cercus with tapered apex; phallus with distinct upcurved lip at edge of opening ( Figs 326, 327 )... E. (En.) manning sp. nov.

21 Hind femur and tibia with dense rows of preapical setae, lacking processes and lobes ( Figs 219, 220 , 343–347 ).......... 22

- Hind femur and tibia with setose processes and lobes (e.g., Figs 17 , 46 , 69 , 74 , 106 , 137 , 146 , 168 , 178 , 189 , 205 , 228 , 258 , 334 )............................................................................................... 24

22 Hind femur with preapical ventral subtriangular swelling ( Fig. 220 ); subepandrium without ventral process ( Sinclair et al. 2013, fig. 13B) [ nodipoplitea group].............................................. E. (En.) nodipoplitea Steyskal

- Hind femur without preapical swelling ( Figs 344–347 ); subepandrium with ventral process ( Figs 350, 351 )............. 23

23 Hind tibia with numerous long dorsal setae, length nearly twice width of tibia; hind tibia with sub-basal setal rows tightly appressed, longer than width of tibia ( Figs 343, 344 ); phallus strongly attenuated near mid-length ( Fig. 350 )..................................................................................... E. (En.) richardvockerothi sp. nov.

- Hind tibia with sparse dorsal setae, length less than twice width of tibia; hind tibia with sub-basal setae in tuft, shorter than width of tibia ( Fig. 346 ); phallus gradually tapered beyond mid-length ( Fig. 351 )............. E. (En.) sierraensis sp. nov.

24 Hind femur with elongate, bare sickle-shaped anteroventral process ( Figs 189–191 , 195, 196 , 202–205 ) [ manca group]... 25

- Hind femur without elongate sickle-shaped anteroventral process (e.g., Figs 30 , 43 , 58 , 95 , 117 , 146 , 155 , 165 , 222, 225 , 258 , 300 )............................................................................................... 28

25 Sickle-shaped process straight, base arising obliquely from hind femur ( Figs 190, 191 ); hind tibia usually with small curvature and anteroventral swelling opposite sickle-shaped process ( Figs 189–191 ); apex of phallus slender, without sculpturing ( Fig. 197 )............................................................................... E. (En.) bova sp. nov.

- Sickle-shaped process strongly curved, base arising perpendicular to hind femur; hind tibia usually without small curvature and anteroventral swelling opposite sickle-shaped process ( Figs 195, 196 , 202–205 ); apex of phallus usually expanded, with sculpturing ( Figs 198–200 , 206 )......................................................................... 26

26 Hind femur with anteroventral and posteroventral rows of short, thickened black setae proximal to sickle-shaped process ( Figs 195, 196 )........................................................................ E. (En.) manca Coquillett

- Hind femur without anteroventral and posteroventral rows of short, thickened black setae proximal to sickle-shaped process ( Figs 202–205 )...................................................................................... 27

27 Hind femur with short row of posteroventral setae, opposite sickle-shaped process ( Fig. 204 ); sickle-shaped process greatly prolonged and slender apically ( Figs 202–204 )......................................... E. (En.) submanca sp. nov.

- Hind femur without short row of posteroventral setae, opposite sickle-shaped process ( Fig. 205 , upper leg); sickle-shaped process not greatly prolonged and slender apically ( Fig. 205 )............................... E. (En.) wilderae sp. nov.

28 Abdomen mostly shiny, especially on apical segments ( Figs 15, 22 , 29 , 62, 63 , 73 , 131 , 176 , 240 ).................... 29

- Abdomen pruinescent; at most only shiny bands on anterior or posterior margin of tergites (doubtful cases, where abdomen appears partially shiny, will key both ways) (e.g., Figs 41 , 55 , 71 , 104 , 154 , 227 , 299 , 321 , 332 )...................... 41

29 Scutum with 4 shiny vittae ( Fig. 239 ); male terminalia narrow and standing erect ( Fig. 240 ) [ turgida group]............ 30

- Scutum without shiny vittae ( Figs 21 , 28 , 32 , 66 , 137 , 176 ); male terminalia directed horizontally ( Figs 15, 22 , 29 , 63 , 73 , 164 ).............................................................................................. 31

30 Tergite 7 with pair of rounded, membranous lateral lobes ( Fig. 243 ); subepandrial process long, extending beyond posterior margin of epandrium ( Fig. 244 ); phallus with 4–5 lateral teeth-like marginal projections in posterior view ( Fig. 245 )........................................................................................ E. (En.) turgida sp. nov.

- Tergite 7 without pair of rounded, membranous lateral lobes; subepandrial process very short, not extending beyond posterior margin of epandrium ( Fig. 247 ); phallus with more than 10 lateral teeth-like marginal projections in posterior view ( Fig. 248 )............................................................................... E. (En.) woodleyi sp. nov.

31 Cercus held upright, perpendicular to epandrium ( Figs 15, 22 , 29 ) [ arrecta group]................................. 32

- Cercus not held upright, parallel to epandrium ( Figs 63 , cf. 71, 73, 164)......................................... 35

32 Scutum with three brownish vittae ( Fig. 28 ); hind tibia with single narrow process ( Figs 15, 16 , 29–31 ); hind tibia distinctly broadened apically ( Figs 16 , 29–31 )..................................................................... 33

- Scutum without three brownish vittae ( Figs 21 , 32 ); hind tibia with pair of narrow processes ( Figs 17–19 , 34 ); hind tibia not distinctly broadened apically ( Figs 17 , 33 )................................................................ 34

33 Distal dorsal setae of hind tibia subequal in length to distal width of tibia ( Fig. 16 ); posteroventral preapical row of thickened setae on hind femur with proximal setae evenly spaced and distal setae slightly shorter than proximal setae ( Fig. 16 )....................................................................................... E. (En.) arrecta sp. nov.

- Distal dorsal setae of hind tibia twice width of distal half of tibia ( Figs 30, 31 ); posteroventral preapical row of thickened setae on hind femur with proximal setae closely approximated and distal setae ca half length of proximal setae ( Fig. 31 )..................................................................................... E. (En.) occidentalis sp. nov.

34 Hind femur with posterior row of 5–6 short, thickened preapical setae ( Figs 17, 19, 20 ); preapical posteroventral lobe of hind femur without narrow lateral prolongation ( Figs 18, 19 ); male cercus with elongate ventrally projecting posterior lobe, hypandrium with long setae ( Fig. 26 ).................................................... E. (En.) diokra sp. nov.

- Hind femur with posterior row of 3–4 short, thickened preapical setae ( Figs 33, 34 ); preapical posteroventral lobe of hind femur with narrow lateral prolongation ( Fig. 34 ); male cercus with short ventrally projecting posterior lobe, hypandrium without long setae ( Fig. 38 )................................................................... E. (En.) shoshona sp. nov.

35 Frons very narrow, eyes closely approximated (narrower than anterior ocellus) or holoptic ( Figs 139 , 176 ).............. 36

- Frons at least as wide as anterior ocellus ( Fig. 66 )........................................................... 39

36 Eyes with upper facets enlarged ( Fig. 177 ); hind tibia somewhat clavate ( Fig. 176 ); hind tarsomere 1 expanded, nearly as wide as apex of hind tibia [ geneatis group]............................................ E. (En.) fisheri sp. nov. [in part]

- Eyes without upper facets enlarged; hind tibia not clavate ( Figs 137 , 164 ); hind tarsomere 1 not as wide as apex of hind tibia. .................................................................................................. 37

37 Hind femur without row of black anteroventral setae proximal to preapical region ( Fig. 135 ) [clava group, in part]................................................................................ E. (En.) mariposa sp. nov. [in part]

- Hind femur with row of black anteroventral setae proximal to preapical region ( Figs 138 , 165 )....................... 38

38 Hind femur with tight row of short, black anteroventral setae proximal to preapical region ( Fig. 165 ); hind tibia with long dorsal setae, length more than twice width of tibia [ delumbis group, in part]................. E. (En.) spiculata sp. nov. [in part]

- Hind femur with row of stout, long anteroventral setae on apical third, length longer than width of femur ( Fig. 138 , lower leg); hind tibia with short dorsal setae, less than twice width of tibia [clava group, in part].... E. (En.) tularensis sp. nov. [in part]

39 Hind femur with posterior preapical setae greatly thickened and lengthened, longer than apical width of femur ( Figs 133, 134 ) [clava group, in part].......................................................... E. (En.) clava sp. nov. [in part]

- Hind femur with posterior preapical setae not greatly thickened and lengthened, not longer than apical width of femur ( Figs 70 , 75 ) [ bigoti group, in part].............................................................................. 40

40 Hind tibia with long, narrow, bare anteroventral pointed lobe ( Figs 67–70 )............ E. (En.) multifaria sp. nov. [in part]

- Hind tibia with broad, anteroventral anvil-shaped setose lobe ( Figs 74, 75 )................... E. (En.) podagra Melander

41 Abdomen grey pruinescent with narrow shiny bands on tergites (sometimes bands partially hidden beneath folds ( Figs 41 , 55 , 71 , 251 )............................................................................................ 42

- Abdomen grey to yellowish, without shiny bands margins on tergites, although some species with pale, dull tergal margins (e.g., Figs 94, 97 , 104 , 154, 157 , 164, 167 , 227 , 277 , 299 )......................................................... 49

42 Hind femur with preapical row of black posterior setae, often short and thickened and arranged horizontally ( Figs 44 , 47 , 70 , 72 , 254 , 270 )........................................................................................ 43

- Hind femur without preapical row of black posterior setae ( Fig. 58 ) [ bigoti group, in part].... E. (En.) montgomeryi sp. nov.

43 Frons narrowed below ocellar triangle, narrower than width of anterior ocellus ( Fig. 66 ) [ bigoti group, in part]................................................................................... E. (En.) multifaria sp. nov. [in part]

- Frons parallel-sided, wider than anterior ocellus ( Fig. 42 ).................................................... 44

44 Wing length 6 mm or longer [ valentis group, in part]........................................................ 45

- Wing length less than 6 mm ............................................................................ 46

45 Scutellum usually with 6 or more setae of various sizes; wing without tuft of basal costal setae distinct from distal row of costal setae; occiput with multiple rows of setae, rather dense, especially ventrally ( Fig. 252 ); large species, wing length generally greater than 7 mm ................................................................. E. (En.) aeripes Melander

- Scutellum with 4 strong setae; wing with tuft of black basal costal setae distinct from distal row of costal setae; occiput with setae confined to two rows, very sparse ventrally ( Fig. 268 ); smaller species, wing length generally less than 7 mm .............................................................................. E. (En.) mixopolia Melander [in part]

46 Wing with tuft of black basal costal setae distinct from distal row of costal setae [ valentis group, in part]....................................................................................... E. (En.) mixopolia Melander [in part]

- Wing with basal costal setae not forming tuft distinct from distal row of costal setae [ bigoti group, in part]............. 47

47 Median ventral lobe of hind femur low, divided apically into pair of lobes ( Figs 46, 47 ); cercus tapered posteriorly ( Fig. 53 )................................................................................. E. (En.) bigoti Melander

- Median ventral lobe of hind femur oval, inflated, rarely shallowly divided apically into pair of low lobes ( Figs 43, 44 , 72 ); cercus rounded posteriorly ( Figs 52 , 78 ).................................................................. 48

48 Lower lobe on hind tibia with setae along posterior margin shorter than length of lobe ( Figs 43, 44 ); phallus with apical section abruptly bent, without rounded lobe at apex of basal section of phallus ( Fig. 52 )............ E. (En.) aerobatica Melander

- Lower lobe on hind tibia with setae along posterior margin, longer than length of lobe ( Fig. 72 ); phallus with apical section gently arched, with rounded lobe at apex of basal section of phallus ( Figs 78, 79 )........... E. (En.) paraerobatica sp. nov.

49 Hind femur with preapical row of black (rarely pale) posterior or posteroventral setae, often short and thickened and arranged horizontally (e.g., Figs 133, 134 , 138 , 147 , 156 , 166 , 272 , 311 ), occasionally seta(e) with expanded tip ( Fig. 301 )........ 50

- Hind femur without preapical row of posterior or posteroventral setae (e.g., Figs 87 , 106 , 114 , 159 , 223, 226 , 232 ), occasionally with cluster of posteroventral setae distal to preapical swelling ( Figs 232 , 313 ).................................... 68

50 Hind femur with long, obliquely directed, digitiform anteroventral process ( Figs 255 , 258, 260 , 271, 273 , 278 ) [ valentis group, in part]............................................................................................ 51

- Hind femur without long, obliquely directed, digitiform anteroventral process ( Figs 50 , 122 , 132 , 135, 138 , 146 , 155 , 165, 168 , 178 , 300 , 310 )....................................................................................... 57

51 Halter with pale, yellowish knob ( Fig. 259 )................................................................ 52

- Halter with dark knob ( Fig. 277 )........................................................................ 56

52 Anteroventral digitiform process of hind femur with short lobe attached at base (viewed ventrally) ( Figs 260 , 273, 274 )... 53

- Anteroventral digitiform process of hind femur without short lobe attached at base ( Figs 255, 256 , 267, 272 )............ 54

53 Basal lobe of anteroventral digitiform process of hind femur with several thin apical setae ( Fig. 260 ).................................................................................................. E. (En.) flavobulla sp. nov.

- Basal lobe of anteroventral digitiform process of hind femur with apical brush of black, thick setae ( Figs 273 ).......................................................................................... E. (En.) subaeripes sp. nov.

54 Hind femur with pair of short lobes medial to digitiform anteroventral process ( Fig. 256 )................................................................................................ E. (En.) disconvenita Melander stat. rev.

- Hind femur with one lobe medial to digitiform anteroventral process ( Figs 267, 272 )............................... 55

55 Lobe medial to digitiform anteroventral process slightly arched apically ( Fig. 272 )........... E. (En.) paraeripes sp. nov.

- Lobe medial to digitiform anteroventral process strongly arched, sickle-shaped ( Fig. 267 ).......... E. (En.) fresno sp. nov.

56 Acrostichals usually uniserial; scutellum with 4 primary setae; wing length shorter than 7 mm; anteroventral process on hind femur with separate inner medial knob ( Fig. 258 )..................................... E. (En.) dolabraria Melander

- Acrostichals biserial; scutellum with 6 or more setae; wing length greater than 7 mm; anteroventral process on hind femur with inner knob at base, not separated ( Figs 278, 279 )....................................... E. (En.) valentis Coquillett

57 Eyes with upper facets enlarged ( Fig. 177 ); hind tibia somewhat clavate; hind tarsomere 1 expanded, nearly as wide as apex of hind tibia ( Fig. 176 ) [ geneatis group, in part]...................................... E. (En.) fisheri sp. nov. [in part]

- Eyes with upper facets not enlarged; hind tibia not clavate; hind tarsomere 1 not expanded as wide as apex of hind tibia ( Figs 48 , 121 , 131 , 137 , 145 , 154 , 167 , 299 , 309 )................................................................ 58

58 Hind femur with row of pale or black anteroventral setae proximal to preapical region ( Figs 132, 133 , 138 , 165 , 300 , 310 )... .................................................................................................. 59

- Hind femur without row of pale or black anteroventral setae proximal to preapical region ( Figs 50 , 122 , 135 , 146 , 155 , 168 ). .................................................................................................. 63

59 Hind femur with pair of nipple-like posteroventral lobes, proximal to row of preapical posteroventral setae ( Fig. 311 )...................................................................................... E. (En.) dubitata sp. nov.

- Hind femur without pair of nipple-like posteroventral lobes, proximal to row of preapical posteroventral setae ( Figs 133 , 166 , 301 )............................................................................................... 60

60 Hind femur with row of stout, long anteroventral setae on apical third, length longer than width of femur ( Fig. 138 )............................................................................... E. (En.) tularensis sp. nov. [in part]

- Hind femur with row of anteroventral setae shorter than width of femur ( Figs 132 , 165 , 300 )........................ 61

61 Hind femur with row of 5–6 thickened posterior setae, shorter than width of femur [ delumbis group, in part] ( Fig. 166 )............................................................................. E. (En.) spiculata sp. nov. [in part]

- Hind femur with 1–2 very elongate thickened posterior setae, longer than width of femur ( Figs 133, 134 , 301 ).......... 62

62 Abdomen with modified tergites 6 and 7 ( Figs 302, 303 ); outstanding posterior seta on hind femur with expanded tip ( Fig. 301 ); hind femur with anteroventral row of dark, thickened setae and sparse dark ventral setae, shorter than width of femur ( Fig. 300 )............................................................................ E. (En.) culcitella sp. nov.

- Abdomen without modified tergites; outstanding posterior setae on hind femur tapered apically ( Figs 133, 134 ); hind femur with anteroventral row of fine, pale setae and dense pale ventral setae, nearly as long as width of femur ( Figs 133, 134 ) [clava group, in part]................................................................ E. (En.) clava sp. nov. [in part]

63 Hind tibia with dorsal setae shorter than width of tibia ( Fig. 137 ) [clava group, in part]...................................................................................................... E. (En.) mariposa sp. nov. [in part]

- Hind tibia with long, sometimes dense dorsal setae, longer than width of tibia ( Figs 48 , 121 , 145 , 154 , 167 )............. 64

64 Hind tibia with dorsal setae three times or more width of tibia ( Figs 154 , 167 ) [ delumbis group, in part]................ 65

- Hind tibia with dorsal setae less than three times width of tibia ( Figs 48 , 121 , 145 )................................ 66

65 Hind femur with 3 closely approximated preapical ventral protuberances ( Figs 155, 156 ); hypandrial setae slender, not longer than length of hypandrium ( Fig. 160 ); abdominal marginal setae dark........................ E. (En.) floricola sp. nov.

- Hind femur with 2 closely approximated preapical ventral protuberances and third offset protuberance ( Figs 168, 169 ); hypandrial setae thickened, longer than length of hypandrium ( Fig. 172 ); abdominal marginal setae pale............................................................................................. E. (En.) tripenicillata sp. nov.

66 Scutum with 3 brown vittae (viewed anteriorly), contrasting with grey pruinescence, with rows of setae running within vittae ( Fig. 49 ); apical third of hind femur with setose knob-like anteroventral lobe ( Fig. 50 ) [ bigoti group, in part].............................................................................................. E. (En.) collinsi sp. nov.

- Scutum with 2 or 4 dark vittae, with rows of setae running between vittae; apical third of hind femur without anteroventral lobe ( Figs 122 , 146 )...................................................................................... 67

67 Hind tibia with basal anteroventral lobe without setae, slightly arched with flattened anterior face ( Fig. 146 ) [ delumbis group, in part]................................................................ E. (En.) delumbis Melander [in part]

- Hind tibia with basal anteroventral lobe with apical setae, knob-like and rounded ( Fig. 122 ) [ canaster group, in part]................................................................................... E. (En.) subcanaster sp. nov.

68 Hind tibia with elongate, forked anteroventral process, fringed with dark setae ( Fig. 334 ).......... E. (En.) poplitea Loew

- Hind tibia without elongate, forked anteroventral process (e.g., Figs 86 , 99 , 105 , 158 , 177 , 231 , 322 ).................. 69

69 Hind femur with 1–2 paint brush-like preapical ventral lobes ( Figs 83, 86, 89 , 95, 98 , 113, 117 , 322 ).................. 70

- Hind femur without paint brush-like preapical ventral lobes ( Figs 105 , 125, 126 , 158 , 222, 225 , 228, 231 , 312 ).......... 77

70 Hind tibia with single basal process ( Figs 86, 89 ) [ canaster group, in part]....................................... 71

- Hind tibia with 2 or more basal processes ( Figs 83 , 95, 98 , 113, 117 , 322 )........................................ 72

71 Hind tibia with long, narrow posteroventral process ( Fig. 86 ); hind tibia with long dorsal setae, twice width of tibia ( Fig. 85 )............................................................................... E. (En.) canaster Melander

- Hind tibia with short, broad, triad posteroventral process ( Fig. 89 ); hind tibia with shorter dorsal setae, only slightly longer than width of tibia ( Fig. 88 )............................................................. E. (En.) falcata Melander

72 Hind femur with preapical, knob-like posterior lobe ( Fig. 99 ); hypandrium with pair of very long, strong, closely approximated setae ( Fig. 102 ) [ canaster group, in part].............................................. E. (En.) contrasta sp. nov.

- Hind femur without preapical knob-like posterior lobe ( Figs 96 , 84 , 114, 118 , 323 ); hypandrium with numerous slender setae or without setae ( Figs 93 , 101 , 119, 120 , 328 ).............................................................. 73

73 Hind femur without curved, bare posteroventral process ( Fig. 323 ); epandrium narrow and prolonged posteriorly ( Fig. 328 )................................................................................. E. (En.) missoula sp. nov.

- Hind femur with curved, bare posteroventral process(es) ( Figs 84 , 96 , 114, 118 ); epandrium round and truncate posteriorly ( Figs 93 , 101 , 119, 120 ) [ canaster group, in part]........................................................... 74

74 Hypandrium without setae ( Fig. 119 ).............................................. E. (En.) sagehenensis sp. nov.

- Hypandrium with setae ( Figs 93 , 101 , 120 )................................................................ 75

75 Hind femur with single curved, bare posteroventral process ( Figs 84 , 96 )........................................ 76

- Hind femur with 3 variously shaped, bare posteroventral processes ( Fig. 118 )............... E. (En.) spokanensis sp. nov.

76 Hind femur with inner ventral paint brush-like process ( Figs 95, 96 )......................... E. (En.) chillcotti sp. nov.

- Hind femur with inner ventral process reduced to unmounted appressed narrow cluster of setae ( Figs 83, 84 )........................................................................................... E. (En.) arizonensis sp. nov.

77 Holoptic ( Fig. 157 ) [ delumbis group, in part]............................................ E. (En.) sphaera sp. nov.

- Dichoptic to closely approximated....................................................................... 78

78 Hind femur with cluster of slender, black posteroventral setae distal to preapical swelling ( Figs 223, 226 , 229, 232 ) [ teres group]............................................................................................. 79

- Hind femur without cluster of slender, black posteroventral setae distal to preapical swelling ( Figs 106 , 126 , 313 )........ 82

79 Fore tarsomere 1 with long dorsal setae, more than twice width of tarsus ( Figs 227, 230 )............................ 80

- Fore tarsomere 1 with dorsal setae, less than twice width of tarsus ( Fig. 221 )..................................... 81

80 Hind femur with preapical ventral whip-like tuft of setae arising from short knob-like lobe ( Figs 228, 229 ); dorsal margin of cercus with angled projection ( Fig. 235 )...................................................... E. teres Melander

- Hind femur with preapical ventral whip-like tuft of setae arising from round, inflated lobe ( Figs 231, 232 ); dorsal margin of cercus with straight dorsal margin ( Fig. 236 )........................................ E. (En.) undulasetosa sp. nov.

81 Hind tibia with single basal lobe ( Fig. 222 )................................................ E. (En.) scotti sp. nov.

- Hind tibia with two basal lobes ( Fig. 225 )............................................... E. (En.) scutica sp. nov.

82 Anterobasal process of hind tibia broad with setal fringe on anterior and posterior sides ( Figs 105, 106 ) [ canaster group, in part]........................................................................... E. (En.) fitzgeraldi sp. nov.

- Anterobasal process of hind tibia narrow, with setae confined mostly to apex ( Figs 125, 126 , 312, 313 )................ 83

83 Hind femur with pair of elongate tufts of appressed preapical setae (anteroventral and posteroventral), longer than apical width of femur ( Figs 312, 313 ); pregenital segments modified ( Figs 314, 315 ); subepandrial process present ( Fig. 317 )........................................................................................... E. (En.) genista sp. nov.

- Hind femur with pair of shorter tufts of setae, shorter than apical width of femur ( Fig. 126 ); pregenital segments unmodified; subepandrial process absent ( Fig. 127 ) [ canaster group, in part]...................................... E. (En.) sp. A

Figures

FIGURES 307–308. Known geographical distribution of the E. (Enoplempis) unplaced species. 307, E. (En.) aphela sp. nov., E. (En.) blodgettensis sp. nov. and E. (En.) californica sp. nov.; 308, E. (En.) cerina sp. nov., E. (En.) delnortensis sp. nov., E. (En.) dubitata sp. nov., E. (En.) susanae sp. nov. and E. (En.) yakimensis sp. nov.

FIGURES 342–349. Male habitus and hindleg of the E. (Enoplempis) unplaced species. 342, E. (En.) richardvockerothi sp. nov., habitus (holotype), lateral view; 343, same, hind tibia, lateral view; 344, same, hind femur tibia joint, anterior view; 345, same, hind femur tibia joint, posterior view; 346, E. (En.) sierraensis sp. nov., hind tibia (holotype), lateral view; 347, same, hind femur tibia joint (holotype), anterior view; 348, E. (En.) susanae sp. nov., hind tibia (holotype), lateral view; 349, E. (En.) yakimensis sp. nov., hind tibia (holotype), lateral view.

FIGURES 350–352. Male terminalia of the E. (Enoplempis) unplaced species, lateral view. 350, E. (En.) richardvockerothi sp. nov.; 351, E. (En.) sierraensis sp. nov. (holotype); 352, E. (En.) yakimensis sp. nov. (holotype).

FIGURES 104–109. Male habitus and hindleg of the E. (Enoplempis) canaster species group. 104, E. (En.) fitzgeraldi sp. nov., habitus, lateral view; 105, same, hind femur tibia joint, anterior view; 106, same, hind femur tibia joint, posterior view; 107, E. (En.) safra sp. nov., habitus (holotype), lateral view; 108, same, hind femur tibia joint, anterior view; 109, same, hind femur tibia joint, posterior view.

FIGURES 143–147. Male habitus and hindleg of E. (Enoplempis) delumbis Melander. 143, habitus, dark form, lateral view; 144, habitus, pale form, lateral view; 145, hindleg, with tarsal spine-like setae, anterior view; 146, hind femur tibia joint, anterior view; 147, hind femur tibia joint, posterior view.

FIGURES 286–291. Male habitus and hindleg of the E. (Enoplempis) unplaced species. 286, E. (En.) aphela sp. nov., hindleg, lateral view; 287, E. (En.) blodgettensis sp. nov., habitus (holotype), lateral view; 288, same, hindleg, lateral view; 289, same, foreleg, lateral view; 290, E. (En.) californica sp. nov., hindleg, lateral view; 291, E. (En.) cerina sp. nov., habitus (holotype), lateral view.

FIGURES 41–45. Male habitus, thorax, hindleg and female habitus of E. (Enoplempis) aerobatica Melander. 41, male habitus, lateral view; 42, thorax, dorsal view; 43, hind femur tibia joint, anterior view; 44, hind femur tibia joint, posterior view; 45, female habitus, lateral view.

FIGURES 62–70. Habitus, head, thorax, abdomen and hindleg of E. (Enoplempis) multifaria sp. nov. 62, male habitus, lateral view; 63, male abdomen and terminalia, lateral view; 64, female abdomen, lateral view; 65, female abdomen, dorsal view; 66, male head and thorax, dorsal view; 67, apex of male hind femur and hind tibia, anterior view; 68, male hindleg, anterior view; 69, hind femur tibia joint, anterior view; 70, hind femur tibia joint, posterior view.

FIGURES 164–169. Male habitus and hindleg of the E. (Enoplempis) delumbis species group. 164, E. (En.) spiculata sp. nov., habitus, lateral view; 165, same, hind femur tibia joint, anterior view; 166, same, hind femur tibia joint, posterior view; 167, E. (En.) tripenicillata sp. nov., habitus (holotype), lateral view; 168, same, hind femur tibia joint, anterior view; 169, same, hind femur tibia joint, posterior view.

FIGURES 176–180. Male habitus, head and hindleg of the E. (Enoplempis) geneatis species group. 176, E. (En.) fisheri sp. nov., habitus, dorsolateral view; 177, same, head (holotype), lateral view; 178, same, hind femur tibia joint, anterior view; 179, E. (En.) oharai sp. nov., habitus, lateral view; 180, same, hindleg, lateral view.

FIGURES 299–304. Male habitus, postabdomen and hindleg of the E. (Enoplempis) unplaced species. 299, E. (En.) culcitella sp. nov., habitus, lateral view; 300, same, hind femur tibia joint, anterior view; 301, same, hind femur tibia joint, posterior view; 302, same, postabdomen, lateral view; 303, same, postabdomen, dorsal view; 304, E. (En.) delnortensis sp. nov., habitus (holotype), lateral view.

FIGURES 331–335. Male habitus, female abdomen and male hindleg of the E. (Enoplempis) unplaced species. 331, E. (En.) oregonensis sp. nov., habitus (holotype), lateral view; 332, E. (En.) poplitea Loew, habitus, lateral view; 333, same, female abdomen; 334, same, hind femur tibia joint, anterior view; 335, same, hind femur tibia joint, posterior view.

FIGURES 210–217. Male hindlegs of the E. (Enoplempis) mira species group. 210, E. (En.) mira (Bigot), left hind femur tibia joint, anterior view; 211, same, right hind femur tibia joint, anterior view; 212, same, right hind femur tibia joint, posterior view; 213, E. (En.) williamturneri Sinclair, Brooks & Cumming, right hind femur tibia joint, posterior view; 214, E. (En.) submira Sinclair, Brooks & Cumming, right hind femur tibia joint (holotype), anterior view; 215, E. (En.) macdonaldi Sinclair, Brooks & Cumming, midleg (holotype), posterior view; 216, same, right hind femur tibia joint (holotype), anterior view; 217, E. (En.) winkleri Sinclair, Brooks & Cumming, right hind femur tibia joint, anterior view.

FIGURES 15–22. Male thorax, abdomen and terminalia, and hindleg of the E. (Enoplempis) arrecta species group. 15, E. (En.) arrecta sp. nov., abdomen, terminalia and hindlegs, lateral view; 16, same, hindlegs, anterior and posterior views; 17, E. (En.) diokra sp. nov., hindlegs, anterior and posterior views; 18, same, hind femur tibia joint, anterior view; 19, same, hind femur tibia joint, posterior view; 20, same, apex of hind femur, dorsal view; 21, same, thorax, dorsal view; 22, same, abdomen and terminalia, lateral view.

FIGURES 71–75. Male habitus, abdomen, terminalia and hindleg of the E. (Enoplempis) bigoti species group. 71, E. (En.) paraerobatica sp. nov., male abdomen and terminalia (holotype), lateral view; 72, same, hind femur tibia joint, anterior and posterior views; 73, E. (En.) podagra Melander, male habitus, lateral view; 74, same, hind femur tibia joint, anterior view; 75, same, hind femur tibia joint, posterior view.

FIGURES 131–134. Holotype male habitus and hindleg of E. (Enoplempis) clava sp. nov. 131, habitus, lateral view; 132, hindleg, anterior view; 133, hind femur tibia joint, posterior view; 134, hind femora, dorsal view.

FIGURES 154–159. Male habitus and hindleg of the E. (Enoplempis) delumbis species group. 154, E. (En.) floricola sp. nov., habitus, lateral view; 155, same, hind femur tibia joint, anterior view; 156, same, hind femur tibia joint, posterior view; 157, E. (En.) sphaera sp. nov., habitus, lateral view; 158, same, hind femur tibia joint, anterior view; 159, same, hind femur tibia joint, posterior view.

FIGURES 188–191. Male habitus and hindleg of E. (Enoplempis) bova sp. nov. 188, habitus, lateral view; 189, hindleg (holotype), anterior view; 190, hind femur tibia joint (holotype), anterior view; 191, hind femur tibia joint (holotype), posterior view.

FIGURES 201–205. Male habitus and hindleg of the E. (Enoplempis) manca species group. 201, E. (En.) submanca sp. nov., habitus, lateral view; 202, same, hindleg, anterior view; 203, same, hind femur tibia joint, anterior view; 204, same, hind femur tibia joint, posterior view; 205, E. (En.) wilderae sp. nov., hindleg (holotype), anterior and posterior views.

FIGURES 218–220. Male habitus and hindleg of E. (Enoplempis) nodipoplitea Steyskal. 218, habitus, lateral view; 219, hindleg, anterior view; 220, hind femur tibia joint, anterior view.

FIGURES 227–232. Male habitus and hindleg of the E. (Enoplempis) teres species group. 227, E. (En.) teres Melander, habitus, lateral view; 228, same, hind femur tibia joint, anterior view; 229, same, hind femur tibia joint, posterior view; 230, E. (En.) undulasetosa sp. nov., habitus, lateral view; 231, same, hind femur tibia joint, anterior view; 232, same, hind femur tibia joint, posterior view.

FIGURES 257–261. Male habitus and hindleg of the E. (Enoplempis) valentis species group. 257, E. (En.) dolabraria Melander, hindlegs, lateral view; 258, same, hind femur tibia joint, anterior and posterior views; 259, E. (En.) flavobulla sp. nov., habitus, lateral view; 260, same, hind femur tibia joint, oblique anterior view [arrow pointing to basal lobe]; 261, same, hind femur tibia joint, posterior view.

FIGURES 181–182. Male habitus and hindleg of E. (Enoplempis) geneatis (Melander). 181, habitus, lateral view; 182, hind femur tibia joint, anterior view.

FIGURES 186–187. Male terminalia of the E. (Enoplempis) geneatis species group, lateral view. 186, E. (En.) oharai sp. nov.; 187, E. (En.) subgeneatis sp. nov.

FIGURES 183–185. Male terminalia of the E. (Enoplempis) geneatis species group, lateral view. 183, E. (En.) fisheri sp. nov.; 184, same, phallus; 185, E. (En.) geneatis (Melander).

FIGURES 319–323. Male habitus and hindleg of the E. (Enoplempis) unplaced species. 319, E. (En.) holoptica sp. nov., habitus, lateral view; 320, E. (En.) manning sp. nov., hindleg (holotype), lateral view; 321, E. (En.) missoula sp. nov., habitus, lateral view; 322, same, hind femur tibia joint, anterior view; 323, same, hind femur tibia joint, posterior view.

FIGURES 292–295. Male terminalia of the E. (Enoplempis) unplaced species, lateral view. 292, E. (En.) aphela sp. nov.; 293, same, phallus; 294, E. (En.) blodgettensis sp. nov.; 295, same, phallus.

FIGURES 324–325. Male terminalia of the E. (Enoplempis) unplaced species, lateral view. 324, E. (En.) dubitata sp. nov.; 325, E. (En.) holoptica sp. nov.

FIGURES 338–341. Male terminalia of the E. (Enoplempis) unplaced species, lateral view. 338, E. (En.) oregonensis sp. nov.; 339, same, phallus; 340, E. (En.) susanae sp. nov.; 341, same, phallus.

FIGURES 296–298. Male terminalia of the E. (Enoplempis) unplaced species, lateral view. 296, E. (En.) californica sp. nov.; 297, E. (En.) cerina sp. nov.; 298, same, phallus.

FIGURES 305–306. Male terminalia of the E. (Enoplempis) unplaced species, lateral view. 305, E. (En.) culcitella sp. nov.; 306, E. (En.) delnortensis sp. nov.

FIGURES 326–328. Male terminalia of the E. (Enoplempis) unplaced species, lateral view. 326, E. (En.) manning sp. nov. (holotype); 327, same, phallus (holotype); 328, E. (En.) missoula sp. nov.

FIGURES 46–51. Male habitus, thorax and hindleg of the E. (Enoplempis) bigoti species group. 46, E. (En.) bigoti Melander, hind femur tibia joint, anterior view; 47, same, hind femur tibia joint, posterior view; 48, E. (En.) collinsi sp. nov., male habitus (holotype), lateral view; 49, same, head and thorax, dorsal view; 50, same, hind femur tibia joint, anterior view; 51, same, hind femur tibia joint, posterior view.

FIGURES 135–139. Male head, thorax and hindleg of the E. (Enoplempis) clava species group. 135, E. (En.) mariposa sp. nov., hind femur tibia joint (holotype), anterior view; 136, same, hind femur (holotype), dorsal view; 137, same, hindleg (holotype), posterior view; 138, E. (En.) tularensis sp. nov., hindlegs, lateral view; 139, same, thorax, dorsal view.

FIGURES 194–196. Male habitus and hindleg of E. (Enoplempis) manca Coquillett. 194, habitus, lateral view; 195, hind femur tibia joint, anterior view; 196, hind femur tibia joint, posterior view.

FIGURES 28–31. Male head, thorax, abdomen and terminalia, and hindleg of E. (Enoplempis) occidentalis sp. nov. 28, head and thorax, dorsal view; 29, abdomen, terminalia and hindlegs (holotype), lateral view; 30, hindleg, anterior view; 31, hindleg, posterior view.

FIGURES 55–58. Abdomen and hindleg of E. (Enoplempis) montgomeryi sp. nov. 55, male abdomen, dorsal view; 56, hindleg, anterior view; 57, hind femur tibia joint, anterior view; 58, hind femur tibia joint, posterior view.

FIGURES 94–99. Male habitus and hindleg of the E. (Enoplempis) canaster species group. 94, E. (En.) chillcotti sp. nov., habitus (holotype), lateral view; 95, same, hind femur tibia joint, anterior view; 96, same, hind femur tibia joint, posterior view; 97, E. (En.) contrasta sp. nov., habitus (holotype), lateral view; 98, same, hind femur tibia joint, anterior view; 99, same, hind femur tibia joint, posterior view.

FIGURES 112–118. Male habitus and hindleg of the E. (Enoplempis) canaster species group. 112, E. (En.) sagehenensis sp. nov., hind tibia (holotype), lateral view; 113, same, hind femur tibia joint (holotype), anterior view; 114, same, hind femur tibia joint (holotype), posterior view; 115, E. (En.) spokanensis sp. nov., hind tibia, lateral view; 116, habitus (holotype), lateral view; 117, same, hind femur tibia joint, anterior view; 118, same, hind femur tibia joint, posterior view.

FIGURES 221–226. Male habitus and hindleg of the E. (Enoplempis) teres species group. 221, E. (En.) scotti sp. nov., habitus (holotype), lateral view; 222, same, hind femur tibia joint (holotype), anterior view; 223, same, hind femur tibia joint (holotype), posterior view; 224, E. (En.) scutica sp. nov., hindleg, with tarsal spine-like setae, anterior view; 225, same, hind femur tibia joint, anterior view; 226, same, hind femur tibia joint, posterior view.

FIGURES 197–200. Male terminalia of the E. (Enoplempis) manca species group, lateral view. 197, E. (En.) bova sp. nov.; 198, E. (En.) manca Coquillett; 199, E. (En.) submanca sp. nov.; 200, same, phallus.

FIGURES 206–207. Male terminalia of the E. (Enoplempis) manca and teres species groups, lateral view. 206, E. (En.) wilderae sp. nov.; 207, E. (En.) scotti sp. nov.

FIGURES 239–242. Male thorax, abdomen and hindleg of E. (Enoplempis) turgida sp. nov. 239, thorax, dorsal view; 240, abdomen and hindleg, lateral view; 241, hind femur tibia joint, anterior view; 242, hind femur tibia joint, posterior view.

FIGURES 32–34. Holotype male head, thorax and hindleg of E. (Enoplempis) shoshona sp. nov. 32, head and thorax, dorsal view; 33, hindlegs, lateral view; 34, hind femur tibia joint, anterior and posterior views.

FIGURES 243–248. Male postabdomen and terminalia of the E. (Enoplempis) turgida species group. 243, E. (En.) turgida sp. nov., postabdomen, dorsal view; 244, same, terminalia, lateral view; 245, same, phallus, lateral and posterior views; 246, E. (En.) woodleyi sp. nov. (holotype), tergite 7, dorsal view; 247, same, terminalia (holotype), lateral view; 248, same, phallus (holotype), posterior view.

FIGURES 23–27. Male terminalia of the E. (Enoplempis) arrecta species group. 23, E. (En.) arrecta sp. nov., lateral view; 24, same, phallus, lateral view; 25, same, phallus, posterior view; 26, E. (En.) diokra sp. nov., lateral view; 27, same, phallus, lateral view. Abbreviations: cerc—cercus; ej apod—ejaculatory apodeme; epand—epandrium; hypd—hypandrium; sbepand proc—subepandrial process.

FIGURES 35–39. Male terminalia of the E. (Enoplempis) arrecta species group. 35, E. (En.) occidentalis sp. nov., lateral view; 36, same, phallus, lateral view; 37, same, phallus, posterior view; 38, E. (En.) shoshona sp. nov. (holotype), lateral view; 39, same, phallus (holotype), lateral view.

FIGURES 251–256. Male habitus, head and hindleg of the E. (Enoplempis) valentis species group. 251, E. (En.) aeripes Melander, habitus, lateral view; 252, same, head, posterolateral view; 253, same, hind femur tibia joint, anterior view; 254, same, hind femur tibia joint, posterior view; 255, E. (En.) disconvenita Melander, hind femur tibia joint, anterior view; 256, same, hind femur tibia joint, posterior view.

FIGURES 277–279. Male habitus and hindleg of E. (Enoplempis) valentis Coquillett. 277, habitus, lateral view; 278, hind femur tibia joint, anterior view; 279, hind femur tibia joint, posterior view.

FIGURES 266–274. Male head and hindleg of the E. (Enoplempis) valentis species group. 266, E. (En.) fresno sp. nov., hind femur tibia joint (holotype), anterior view; 267, same, hind femur tibia joint (holotype), posterior view; 268, E. (En.) mixopolia Melander, head, lateral view; 269, same, hind femur tibia joint, anterior view; 270, same, hind femur tibia joint, posterior view; 271, E. (En.) paraeripes sp. nov., hind femur tibia joint, anterior view; 272, same, hind femur tibia joint, posterior view; 273, E. (En.) subaeripes sp. nov., hind femur tibia joint (holotype), oblique anterior view [arrow pointing to basal lobe]; 274, same, hind femur tibia joint (holotype), posterior view.

FIGURES 52–53. Male terminalia of the E. (Enoplempis) bigoti species group, lateral view. 52, E. (En.) aerobatica Melander; 53, E. (En.) bigoti Melander.

FIGURES 76–79. Male terminalia of the E. (Enoplempis) bigoti species group, lateral view. 76, E. (En.) multifaria sp. nov.; 77, same, phallus; 78, E. (En.) paraerobatica sp. nov.; 79, same, phallus.

FIGURES 309–315. Male postabdomen and hindleg of the E. (Enoplempis) unplaced species. 309, E. (En.) dubitata sp. nov., hindleg, anterior view; 310, same, hind femur tibia joint, anterior view; 311, same, hind femur tibia joint, posterior view; 312, E. (En.) genista sp. nov., hind femur tibia joint, anterior view; 313, same, hind femur tibia joint, posterior view; 314, same, postabdomen (holotype), lateral view; 315, same, postabdomen (holotype), dorsal view.

FIGURES 82–90. Male hindleg of the E. (Enoplempis) canaster species group. 82, E. (En.) arizonensis sp. nov., hindleg, anterior view; 83, same, hind femur tibia joint, anterior view; 84, same, hind femur tibia joint, posterior view; 85, E. (En.) canaster Melander, hind tibia, anterior view; 86, same, hind femur tibia joint, anterior view; 87, same, hind femur tibia joint, posterior view; 88, E. (En.) falcata Melander, hind femur tibia joint (holotype), anterior view; 89, same, hind femur tibia joint (holotype), anterior view; 90, same, hind femur tibia joint (holotype), posterior view.

FIGURES 121–126. Male habitus and hindleg of the E. (Enoplempis) canaster species group. 121, E. (En.) subcanaster sp. nov., habitus, lateral view; 122, same, hind femur tibia joint, anterior view; 123, same, hind femur tibia joint, posterior view; 124, E. (En.) sp. A, hindleg, anterior view; 125, same, hind femur tibia joint, anterior view; 126, same, hind femur tibia joint, posterior view.

FIGURES 160–163. Male terminalia of the E. (Enoplempis) delumbis species group, lateral view. 160, E. (En.) floricola sp. nov.; 161, same, phallus; 162, E. (En.) sphaera sp. nov.; 163, same, phallus.

FIGURES 170–173. Male terminalia of the E. (Enoplempis) delumbis species group, lateral view. 170, E. (En.) spiculata sp. nov.; 171, same, phallus; 172, E. (En.) tripenicillata sp. nov.; 173, same, phallus.

FIGURES 102–103. Male terminalia of the E. (Enoplempis) canaster species group, lateral view. 102, E. (En.) contrasta sp. nov.; 103, E. (En.) falcata Melander.

FIGURES 91–93. Male terminalia of the E. (Enoplempis) bigoti and E. (En.) canaster species groups, lateral view. 91, E. (En.) podagra Melander; 92, same, phallus; 93, E. (En.) arizonensis sp. nov.

FIGURES 100–101. Male terminalia of the E. (Enoplempis) canaster species group, lateral view. 100, E. (En.) canaster Melander; 101, E. (En.) chillcotti sp. nov.

FIGURES 119–120. Male terminalia of the E. (Enoplempis) canaster species group, lateral view. 119, E. (En.) sagehenensis sp. nov.; 120, E. (En.) spokanensis sp. nov.

FIGURES 233–236. Male terminalia of the E. (Enoplempis) teres species group, lateral view. 233, E. (En.) scutica sp. nov.; 234, same, phallus; 235, E. (En.) teres Melander; 236, E. (En.) undulasetosa sp. nov.

FIGURES 316–318. Male postabdomen and terminalia of E. (Enoplempis) genista sp. nov. 316, postabdominal tergites 7–8, dorsal view; 317, terminalia, lateral view; 318, phallus, lateral view.

FIGURES 127–128. Male terminalia of the E. (Enoplempis) canaster species group, lateral view. 127, E. (En.) sp. A; 128, E. (En.) subcanaster sp. nov.