Euura vaga ( Fabricius, 1781 )

Prous, Marko, Liston, Andrew, Monckton, Spencer K., Kramp, Katja, Vårdal, Hege, Vikberg, Veli, Heibo, Erik & Mutanen, Marko, 2025, West Palaearctic species of Euura Newman, 1837 (Hymenoptera, Tenthredinidae), European Journal of Taxonomy 977, pp. 1-377 : 230-232

303 Euura vaga ( Fabricius, 1781)

Figs 146 , 542

Tenthredo vaga Fabricius, 1781: 415 . Syntypes (3 specimens in NHMD according to Zimsen 1964), sex not stated, not examined. Type locality: England.

Tenthredo flaviventris Gmelin, 1790: 2667 . Syntypes, sex not stated, not located. Type locality: Europe. Synonymy by Konow (1904c). Primary homonym of Tenthredo flaviventris Retzius, 1783 [ Neurotoma saltuum ( Linnaeus, 1758) ].

Nematus leucogaster Hartig, 1840: 23 . Lectotype designated below. Synonymy by Konow (1904c).

Nematus hypoleucus Förster, 1854a: 304–305 . Lectotype designated below. Synonymy by Konow (1904c).

Nematus micraulius Förster, 1854b: 428–429 . Lectotype designated below. Synonymy by Konow (1904c).

Nematus punctipleuris Thomson, 1863: 628 . Lectotype designated below. Synonymy by Konow (1904c).

Nematus punctulatus Thomson, 1863: 626–627 . Lectotype designated below. Synonymy by Konow (1904c).

Nematus inconspicuus W.F. Kirby, 1882: 141 . Holotype ♀ (BMNH?), not examined. Type locality: New York. Synonymy by Benson (1962).

Pachynematus corticosus MacGillivray, 1901: 584–585 . Holotype ♀ (INHS), not examined. Type locality: USA, New York, Saranac Inn.

Holcocneme ulbrichti Enslin, 1910: 316–317 . Holotype ♀, not found in ZSM. Type locality: Germany, Nordrhein-Westfalen, Krefeld. Synonymy by Enslin (1915).

Pachynematus roscidus MacGillivray, 1921b: 31 . Holotype ♀ (INHS), not examined. Type locality: USA, Maine, Orono.

Pachynematus rugosulus Lindqvist, 1959b: 71 . Syn. nov.

Diagnosis

The female is most similar to E. obducta from which it differs in usually having a completely pale metafemur (extensively black in E. obducta ), more triangular valvula 3 in dorsal view, and apically curved basal part of lamnium (more or less straight in E. obducta ). Males, which are extremely rare in Europe, are best recognised by their penis valves. Abdomen sometimes also dorsally nearly completely pale (terga 3–10) (ZMUO.062019). Pterostigma varies in European specimens from yellowish to dark brown, in North America often (usually?) black.

Type material examined

Lectotype Nematus leucogaster , here designated

GERMANY • ♀; northern Germany [according to Hartig 1840: 22]; ZSM, GBIF-GISHym3334 .

Lectotype Nematus hypoleucus , here designated

GERMANY – Nordrhein-Westfalen • ♀; Aachen area ; 50.77° N, 6.09° E; ZSM, GBIF-GISHym3297 .

Lectotype Nematus micraulius , here designated

GERMANY – Nordrhein-Westfalen • ♂; Aachen area ; 50.77° N, 6.09° E; ZSM, GBIF-GISHym3352 .

Notes

Penis valves lost, but identified by Conde as Pristiphora vaga .

Lectotype Nematus punctipleuris , here designated

SWEDEN • ♀; Norlandia [Lapland, probably Sweden]; MZLU, MZLU2017272 .

Paralectotype Nematus punctipleuris

SWEDEN • ♀; Norlandia [Lapland, probably Sweden]; MZLU, MZLU2017273 .

Lectotype Nematus punctulatus , here designated SWEDEN • ♀; Torne Lappmark ; MZLU, MZLU2017265

Paralectotypes Nematus punctulatus

SWEDEN • 2 ♀♀; Torne Lappmark ; MZLU, MZLU 2017266 , MZLU 2017269 • 2 ♀♀; [labels illegible]; MZLU, MZLU 2017267 , MZLU 2017268 .

Holotype Pachynematus rugosulus

NORWAY – Finnmark • ♀; Jotkajavrek ; 69.77° N, 23.85° E; Jul. 1924; Expedition A. Strand leg.; ZMUN, NHMO:1000201217 .

Notes

Basal quarter to third of metafemur brown, mesepisternum matt. Best morphological match: ZMUO.041887.

Host plants

Carex spp. More than one generation per year in suitable conditions ( Lorenz & Kraus 1957; Kangas 1985).

Genetics

COI

Based on 29 specimens, maximum within-species distance is 3.5% and the nearest neighbour, diverging by a minimum of 6.26%, is Euura miliaris . Including 8 specimens from North America does not change the results.

Nuclear

Based on 8 specimens, maximum within-species distance is 0.61% (1.02% based on haplotypes of individual females). The nearest neighbour, diverging by a minimum of 2.93%, is Euura fuscomaculata . When a specimen from North America (DEI-GISHym15293) is included, maximum within-species distance is 0.8% (1.02% based on haplotypes of individual females).

Distribution and material examined

Holarctic. Specimens studied are from Canada, Estonia, Finland, France, Germany, Iceland, Norway, Russia, Slovakia, Sweden, United Kingdom, and the USA.

Figures

Figs 146–149. Lancets of Euura Newman, 1837. 146. vaga (Fabricius, 1781) Prep.344.OR. 147. epilosa (Lindqvist, 1970), Pteronidea epilosa Lindqvist, 1970, holotype, ♀ https://id.luomus.fi/GL.5142. 148. epilosa DEI-GISHym80684. 149. fahraei (Thomson, 1863) PR.568VV.

Figs 534–545. Penis valves of Euura Newman, 1837 (534–543, 545) and Pristiphora Latreille, 1810 (544). 534. hyperborea (Thomson, 1871) DEI-GISHym31674. 535. imperfecta (Zaddach, 1876) ZMUO.030832. 536. itoi (Okutani, 1955) DEI-GISHym13850. 537. poecilonota (Zaddach, 1876) DEI- GISHym12527. 538. pumilio (Konow, 1903) DEI-GISHym31087. 539. salicis (Linnaeus, 1758) DEI- GISHym17954. 540. spiraeae (Zaddach, 1883) DEI-GISHym14075 (Switzerland). 541. spiraeae DEI- GISHym1916 (China). 542. vaga (Fabricius, 1781) DEI-GISHym13874. 543. telos Liston & Prous nom. nov. DEI-GISHym80794. 544. Pristiphora biscalis (Förster, 1854), Nematus meridionalis André, 1880, lectotype DEI-GISHym89376. 545. fagi Zaddach, 1876 DEI-GISHym13853.