Zaptyx, PILSBRY, 1900

Motochin, Risho, Wang, Min & Ueshima, Rei, 2017, Molecular phylogeny, frequent parallel evolution and new system of Japanese clausiliid land snails (Gastropoda: Stylommatophora), Zoological Journal of the Linnean Society 181, pp. 795-845 : 833

GENUS ZAPTYX PILSBRY, 1900

(= CLADE D)

Type species: Clausilia hirasei Pilsbry, 1900

Synonyms: Diceratoptyx Pilsbry, 1905 ; Hemizaptyx Pilsbry, 1905 ; Heterozaptyx Pilsbry, 1906 ; Idiozaptyx Pilsbry, 1909 ; Oligozaptyx Pilsbry, 1905 ; Metazaptyx Pilsbry, 1905 ; Paganizaptyx Azuma, 1982 ; Parazaptyx Pilsbry, 1905 ; Pulchraptyx Minato, 1981 ; Selenoptyx Pilsbry, 1908 ; Stereozaptyx Pilsbry, 1901 ; Thaumatoptyx Pilsbry, 1908 ; Tyrannozaptyx Käufel, 1930 ; Zaptychopsis Ehrmann, 1927 .

Diagnosis: Shell mostly small and slender, but becoming large only in a few subgenera. Plicae and clausilium located laterally to ventrally. Sutural plica present or absent. Lunella always present, strongly developed. Upper palatal plica usually present, mostly attached to the lunella, variable in the length. Presence of unique deletions and substitution in the hyper variable region of ITS2 ( Fig. 5) that are shared only with Tauphaedusa is a molecular diagnostic character of the genus.

Morphology: Apex pointed. Lower palatal plica frequently absent. Middle palatal plica absent. Superior lamella usually developed, rarely reduced. Inferior lamella variable, spirally ascending and appearing at the aperture, or almost straightly ascending and receding within the aperture. Clausilium wide or narrow. Diverticulum of bursa copulatrix developed. See also Diagnosis.