Invreiella Suárez, 1966

Williams, Kevin A., 2025, Color syndrome terminology for velvet ant females (Hymenoptera: Mutillidae and Myrmosidae), Zootaxa 5684 (1), pp. 1-238 : 89-95

Invreiella Suárez, 1966 —The species were keyed by Waldren et al. (2020).

Fig. 1160— Invreiella acuminata Waldren in Waldren et al., 2020: Code 3/2, length 9.3 mm, Mexico (NA), No further data, NHMUK. Holotype. Published in Waldren et al. (2020).

Fig. 1161— Invreiella australis Waldren in Waldren et al., 2020: Code 1/1, length 10.9 mm, Mexico (NT), Chiapas, Villa Flores (16.32 -93.34), DGMC. Holotype. Published in Waldren et al. (2020).

Fig. 1162— Invreiella bimaculata Waldren in Waldren et al., 2020: Code 1/1, length 9.3 mm, Mexico (NA), Sinaloa, Microondas El Indo (23.66 -106.56), UNAM. Holotype. Published in Waldren et al. (2020).

Fig. 1163— Invreiella breviclypeata Waldren in Waldren et al., 2020: Code 1/3, length 10.4 mm, Mexico (NA), Sinaloa, 5 mi West El Palmito (23.56 -105.92), DGMC. Holotype. Published in Waldren et al. (2020).

Fig. 1164— Invreiella cardinalis (Gerstaecker, 1874) : Code 1/3, length 11 mm, Mexico (NT), Veracruz, Xalapa (19.54 -96.91), ZMHB. Holotype. Published in Waldren et al. (2020).

Fig. 1165— Invreiella cephalargia (Mickel, 1924) : Code 3/2, length 10 mm, USA (NA), Arizona, Leslie Canyon (31.58 -109.51), DGMC. Determined by D.G. Manley. Published in Waldren et al. (2020) .

Fig. 1166— Invreiella chihuahuensis Waldren in Waldren et al., 2020: Code 1/3, length 8 mm, Mexico (NA), Chihuahua, Ciudad Juarez (31.69 -106.42), DGMC. Holotype. Published in Waldren et al. (2020).

Fig. 1167— Invreiella cuernavaca Waldren in Waldren et al., 2020: Code 1/2, length 10.6 mm, Mexico (NA), Morelos, Cuernavaca (18.92 -99.22), CASC. Holotype. Published in Waldren et al. (2020).

Fig. 1168— Invreiella erythrocephala Waldren in Waldren et al., 2020: Code 1/3, length 11.2 mm, Mexico (NT), Morelos, 11 km NW Tepexco (18.70 -98.77), CASC. Holotype. Published in Waldren et al. (2020).

Fig. 1169— Invreiella manleyi Waldren in Waldren et al., 2020: Code 1/3, length 7.5 mm, USA (NA), Arizona, 15 mi West Portal (31.93 -109.39), DGMC. Holotype. Published in Waldren et al. (2020).

Fig. 1170— Invreiella mesomexicana Waldren in Waldren et al., 2020: Code 1/2, length 12.8 mm, Mexico (NT), Guanajuato, Roque , North of Celaya (20.58 -100.84), EMUS. Holotype. Published in Waldren et al. (2020).

Fig. 1171— Invreiella satrapa (Gerstaecker, 1874) : Code 1/3, length 11 mm, Mexico (NA), Durango, 31.7 mi North Vitoria de Durango (24.40 -104.70), CSCA. Determined by G.C. Waldren. Published in Waldren et al. (2020) .

Fig. 1172— Invreiella suarezi Waldren in Waldren et al., 2020: Code 3/2, length 9.9 mm, Mexico (NA), Guerrero, Atlixtac km 39 Chilapa-Tlapa (17.60 -99.02), UNAM. Holotype. Published in Waldren et al. (2020).

Fig. 1173— Invreiella tequila Waldren in Waldren et al., 2020: Code 3/2, length 12.5 mm, Mexico (NT), Jalisco, Tequila (20.88 -103.83), FSCA. Holotype. Published in Waldren et al. (2020).

Kevinilla Bartholomay & Cambra in Bartholomay et al., 2024—The species were keyed by Bartholomay et al. (2024).

Fig. 1174— Kevinilla bicarinata (Cambra & Quintero, 2008): Code 3/2, length 7.5 mm, Unknown (NT), Further data not available. Published in Bartholomay et al., 2024.

Fig. 1175— Kevinilla bimaculata (Cambra & Quintero, 2008) : Code 2/1, length 6.7 mm, Panama (NT), Further data not available. Published in Bartholomay et al., 2024.

Fig. 1176— Kevinilla hansoni (Cambra & Quintero, 2008): Code 2/1, length 7.2 mm, Panama (NT), Panama, 9.5 km North El Llano (9.30 -78.98), CSCA. Determined by KAW. Photo by KAW at USU.

Fig. 1177— Kevinilla ludovica (Cameron, 1895): Code 2/1, length 4 mm, Costa Rica (NT), Cartago, Tuis (9.86 -83.57), FSCA. Determined by KAW. Photo by KAW at DPI .

Fig. 1178— Kevinilla cf. ludovica (Cameron, 1895): Code 1/1, length 4 mm, Mexico (NT), Guerrero (16.81 - 98.69), CSCA. Determined by KAW. Photo by KAW at DPI .

Lophomutilla Mickel, 1952 —Keys to the species were written by Fritz (1990) and Fritz & Pagliano (1993); an unpublished thesis by Cambra (1996) also has a good key.

Fig. 1179— Lophomutilla calva Fritz, 1990 : Code 3/1, length 9.4 mm, Trinidad (NT), Talparo (10.51 -61.27), FSCA. Determined by KAW. Photo by KAW at USU .

Fig. 1180— Lophomutilla caribe Casal, 1963 : Code 4/1, length 11 mm, Brazil (NT), Maranhão, Açailândia (-4.95 -47.50), MPEG. Determined by KAW. Photo by KAW at UFPR .

Fig. 1181— Lophomutilla chrysomalla (Gerstaecker, 1874) : Code 2/1, length 8 mm, Brazil (NT), Paraná, Chácara São Roque (-24.93 -51.64), DZUP. Determined by KAW. Photo by KAW at UFPR.

Fig. 1182— Lophomutilla corupa Casal, 1968 : Code 2/1, length 5 mm, Brazil (NT), Paraná, Parque Estadual de Vila Velha (-25.22 -50.04), DZUP. Determined by KAW. Photo by KAW at UFPR.

Fig. 1183— Lophomutilla inca Fritz & Pagliano, 1993 : Code 1/1, length 5.3 mm, Peru (NT), Further data not available, EMUS. Determined by KAW. Photo by KAW at CDFA.

Fig. 1184— Lophomutilla lenkoi Suárez, 1962 : Code 2/1, length 7.5 mm, Brazil (NT), Paraná, Mananciais da Serra, Piraquara (-25.23 -49.98), DZUP. Determined by KAW. Photo by KAW at UFPR.

Fig. 1185— Lophomutilla mickeli Suárez, 1962 : Code 2/1, length 8 mm, Brazil (NT), Paraná, Parque Estadual de Vila Velha (-25.22 -50.04), DZUP. Determined by KAW. Photo by KAW at UFPR.

Fig. 1186— Lophomutilla obscura Fritz & Pagliano, 1993 : Code 3/2, length 6 mm, Brazil ** (NT), Mato Grosso, Rio Manso (-14.87 -55.80), DZUP. Determined by KAW. Photo by KAW at UFPR.

Fig. 1187— Lophomutilla prionophora jaragua Casal, 1961 : Code 3/1, length 8 mm, Brazil (NT), Paraná (-23.89 -52.00), DZUP. Determined by KAW. Photo by KAW at UFPR.

Fig. 1188— Lophomutilla prionophora prionophora (Burmeister, 1854) : Code 3/1, length 10 mm, Brazil (NT), Minas Gerais, 12 km N Bandeira, Reserva Biodiversita (-15.78 -40.52), DZUP. Determined by KAW. Photo by KAW at UFPR.

Fig. 1189— Lophomutilla seabrai Casal, 1968 : Code 2/1, length 5 mm, Brazil (NT), Rio de Janeiro, 6 km NW de Itatiaia, PN Itatiaia, Hotel Simon (-22.43 -44.60), DZUP. Determined by KAW. Photo by KAW at UFPR.

Fig. 1190— Lophomutilla staphylomma (Gerstaecker, 1874) : Code 2/1, length 5 mm, Brazil (NT), Rio de Janeiro, 6 km NW de Itatiaia, PN Itatiaia, Hotel Simon (-22.43 -44.60), DZUP. Determined by KAW. Photo by KAW at UFPR. Fig. 1191— Lophomutilla triguttata Mickel, 1952 : Code 4/1, length 7.4 mm, Peru ** (NT), Further data not available, EMUS. Determined by KAW. Photo by KAW at CDFA.

Fig. 1192— Lophomutilla tuberculata (Fabricius, 1804) : Code 1/1, length 7.9 mm, Guyana (NT), Essequibo

( 5.92 -58.60), ZMUC. Holotype. Photo posted online by NHMUK: https://collections.snm.ku.dk/en/object/ NHMD49771 , accessed on May 5, 2025.

Fig. 1193— Lophomutilla urca Casal, 1969 : Code 2/1, length 7 mm, Brazil (NT), Rio de Janeiro, Floresta da Tijuca (-22.95 -43.27), AMNH. Determined by M.A. Fritz. Photo by KAW at CDFA .

Fig. 1194— Lophomutilla vina Fritz & Pagliano, 1993 : Code 3/2, length 4.6 mm, Bolivia ** (NT), Further data not available, EMUS. Determined by KAW. Photo by KAW at CDFA.

Fig. 1195— Lophomutilla yala Casal, 1968 : Code 2/1, length 6 mm, Brazil (NT), Rio de Janeiro, Santa Teresa (- 19.94 -40.60), MZSP. Determined by KAW. Photo by G. Furman at UFPR .

Fig. 1196— Lophomutilla cf. braulioi (FPIP025) Fritz & Pagliano, 1993: Code 1/1, length 4.5 mm, Brazil (NT), Pará, Goianésia, Faz.Rio Capim (-3.84 -49.09), CEPLAC. Determined by KAW. Photo by KAW at UFPR.

Fig. 1197— Lophomutilla cf. obscura (FPIP027) Fritz & Pagliano, 1993: Code 3/2, length 4.5 mm, Brazil (NT), Paraná, Parque Estadual de Vila Velha (-25.23 -50.00), DZUP. Determined by KAW. Photo by KAW at UFPR. Fig. 1198— Lophomutilla cf. triguttata Mickel, 1952 : Code 3/1, length 8 mm, Suriname (NT), Brokopondo (5.03 -54.99), RMNH. Determined by KAW. Photo by KAW at DPI.

Fig. 1199— Lophomutilla aff. denticulata (Smith, 1855) : Code 3/2, length 7 mm, Brazil (NT), Pará, Belém, Conjunto Panorama XXI (-1.38 -48.43), MPEG. Determined by KAW. Photo by KAW at UFPR.

Fig. 1200— Lophomutilla sp. nov. 1 (FPIP023): Code 2/1, length 5 mm, Brazil (NT), Paraná, Parque Estadual de Vila Velha (-25.22 -50.04), DZUP. Determined by KAW. Photo by KAW at UFPR.

Fig. 1201— Lophomutilla sp. nov. 2 (FPIP024): Code 3/2, length 4.5 mm, Brazil (NT), Rondônia, Itapuã do Oeste, FLONA do Jamari (-9.26 -62.91), DZUP. Determined by KAW. Photo by KAW at UFPR.

Fig. 1202— Lophomutilla sp. nov. 3 (FPIP032): Code 2/1, length 6 mm, Brazil (NT), Amazonas, Tefé (-3.35 - 64.71), DZUP. Determined by KAW. Photo by KAW at UFPR.

Fig. 1203— Lophomutilla sp. nov. 4 (FPIP033): Code 2/1, length 6 mm, Brazil (NT), Rio de Janeiro, 6 km NW de Itatiaia, PN Itatiaia Hotel Simon (-22.49 -44.56), DZUP. Determined by KAW. Photo by KAW at UFPR.

Fig. 1204— Lophomutilla sp. nov. 5 (FPIP039): Code 2/1, length 8 mm, Brazil (NT), São Paulo, Campos do Jordão (-22.74 -45.59), MZSP. Determined by KAW. Photo by G. Furman at UFPR .

Fig. 1205— Lophomutilla sp. nov. 6 (FPIP040): Code 3/2, length 8 mm, Brazil (NT), Santa Catarina, Rio Vermelho (-26.37 -49.29), MZSP. Determined by KAW. Photo by G. Furman at UFPR .

Fig. 1206— Lophomutilla sp. nov. 7 (FPIP043): Code 3/2, length 5 mm, Brazil (NT), Amazonas, Rio Negro, Tapuruquara (-0.49 -65.08), MZSP. Determined by KAW. Photo by G. Furman at UFPR.

Fig. 1207— Lophomutilla sp. nov. 8 (FPIP055): Code 1/2, length 4 mm, Brazil (NT), Minas Gerais, Bom Despacho (-19.74 -45.25), CEPLAC. Determined by KAW. Photo by KAW at UFPR .

Fig. 1208— Lophomutilla sp. nov. 9 (FPIP056): Code 3/2, length 5.5 mm, Brazil (NT), Mato Grosso, 30 km North Uirapuru, Chapada dos Parecis (-12.97 -60.03), EMUS. Determined by KAW. Photo by KAW at UFPR.

Fig. 1209— Lophomutilla sp. nov. 10 Code 1/1, length 4.2 mm, Trinidad (NT), Arima Ward, Aripo Valley (10.67 -61.23), FSCA. Determined by KAW. Photo by KAW at USU .

Fig. 1210— Lophomutilla sp. nov. 11 Code 1/1, length 4.2 mm, Costa Rica (NT), Cartago, Tuis (9.86 -83.57), FSCA. Determined by KAW. Photo by KAW at USU .

Lophostigma Mickel, 1952 —The Central American species were keyed by Cambra & Quintero (1996); many South American species were included in Mickel’s (1952) key.

Fig. 1211— Lophostigma acanthophora (Dalla Torre, 1897) : Code 3/1, length 8 mm, Unknown (NT), Further data not available, Collection unknown. Identifier unknown. Photo by R.A. Cambra in Pagliano et al. (2020).

Fig. 1212— Lophostigma alopha Mickel, 1952 : Code 2/1, length 8 mm, Brazil ** (NT), Amazonas, Ilha de Curari- Varzea (-3.34 -60.10), EMUS. Determined by KAW. Photo by KAW at UFPR.

Fig. 1213— Lophostigma cayennensis (André, 1906): Code 2/1, length 7.5 mm, Brazil ** (NT), Amazonas, Manaus, Reserva Ducke (-2.93 -59.97), MPEG. Determined by KAW. Photo by KAW at UFPR.

Fig. 1214— Lophostigma cincta 1 (Buysson, 1892): Code 3/1, length 7.8 mm, Costa Rica ** (NT), Heredia (10.33 -84.08), CSCA. Determined by KAW. Photo by KAW at USU .

Fig. 1215— Lophostigma cincta 2 (Buysson, 1892): Code 3/2, length 5.6 mm, Costa Rica (NT), Guanacaste, Finca Montezuma (10.66 -85.08), CSCA. Determined by KAW. Photo by KAW at USU .

Fig. 1216— Lophostigma grisseli Cambra & Quintero, 1996 : Code 3/1, length 7.9 mm, Mexico (NT), Yucatan, 8 mi East Merida (20.91 -89.47), SEMC. Determined by KAW. Photo by KAW at CDFA.

Fig. 1217— Lophostigma iracunda (FPIP051) (Cresson, 1902): Code 3/1, length 9 mm, Brazil (NT), Pará, Rio Xingú, A2 Igarapé di Maria (-2.88 -52.01), MPEG. Determined by KAW. Photo by KAW at UFPR.

Fig. 1218— Lophostigma seabrai Casal, 1963 : Code 2/1, length 7 mm, Brazil (NT), Rio de Janeiro (-22.73 - 43.23), DZUP. Determined by KAW. Photo by KAW at UFPR.

Fig. 1219— Lophostigma simoni (Buysson, 1892) : Code 3/1, length 5.3 mm, Trinidad ** (NT), Simla Res.St. (10.69 -61.29), EMUS. Determined by KAW. Photo by KAW at USU.

Fig. 1220— Lophostigma subgracilis (Cameron, 1895) : Code 1/2, length 8.2 mm, Costa Rica ** (NT), Cartago, 3 km East Turrialba, CATIE (9.89 -83.66), CSCA. Determined by KAW. Photo by KAW at USU.

Fig. 1221— Lophostigma cf. alopha Mickel, 1952 : Code 2/1, length 4 mm, Brazil (NT), Pará, Belém, Mocambo (-1.46 -48.47), MPEG. Determined by KAW. Photo by KAW at UFPR.

Fig. 1222— Lophostigma cf. caenodonta (Cameron, 1912) : Code 2/1, length 6.5 mm, Brazil (NT), Pará, Benevides (-1.35 -48.24), MPEG. Determined by KAW. Photo by KAW at UFPR.

Fig. 1223— Lophostigma sp. nov. 1 (FPIP021): Code 3/1, length 9 mm, Brazil (NT), Amazonas, Estirão do Equador, Rio Javari (-6.60 -73.30), DZUP. Determined by KAW. Photo by KAW at UFPR.

Fig. 1224— Lophostigma sp. nov. 2: Code 3/2, length 5 mm, Brazil (NT), Mato Grosso dp Sul, Campo Grande (- 20.40 -54.60), DZUP. Determined by KAW. Photo by KAW at UFPR.

Lynchiatilla Casal, 1963 — The best available keys were written by Casal (1963b) and Bergamaschi et al. (2012). Fig. 1225— Lynchiatilla armanda (Smith, 1879) : Code 2/1, length 11 mm, Brazil (NT), Rio de Janeiro, Petrópolis (-22.51 -43.18), MNRJ. Determined by KAW. Photo by KAW at UFPR.

Fig. 1226— Lynchiatilla bubekii (Dalla Torre, 1897) : Code 1/1, length 8 mm, Argentina (NT), Granja [La Granja, Cordoba] (-31.02 -64.27), UMSP. Determined by KAW. Photo by KAW at CDFA.

Fig. 1227— Lynchiatilla chayera Casal, 1963 : Code 1/1, length 8.1 mm, Argentina (NT), Huerta Grande (-31.08 -64.49), UMSP. Determined by KAW. Photo by KAW at CDFA.

Fig. 1228— Lynchiatilla hoplites (Gerstaecker, 1874) : Code 2/1, length 7 mm, Paraguay (NT), Paraguarí, East Ybycuí (-26.02 -57.02), CSCA. Determined by KAW. Photo by KAW at DPI .

Fig. 1229— Lynchiatilla leguera Casal, 1963 : Code 1/1, length 6 mm, Argentina (NT), Jujuy, Huacalera , 17 km North Tilcara (-23.44 -65.35), CSCA. Determined by D.J. Brothers. Photo by KAW at DPI .

Fig. 1230— Lynchiatilla parana Cambra, 2012 : Code 2/1, length 6.9 mm, Brazil (NT), Paraná, Parque Estadual de Vila Velha (-25.23 -50.00), DZUP. Determined by KAW. Photo by KAW at UFPR.

Fig. 1231— Lynchiatilla seabrai Casal, 1963 : Code 1/2, length 4.2 mm, Brazil (NT), São Paulo, Bragança Paulista.Haras Danúbio (-22.95 -46.54), MZSP. Determined by KAW. Photo by G. Furman at UFPR .

Fig. 1232— Lynchiatilla serrana Casal, 1963 : Code 3/2, length 6.5 mm, Argentina (NT), Huerta Grande (-31.08 -64.49), UMSP. Determined by KAW. Photo by KAW at CDFA.

Fig. 1233— Lynchiatilla silvai Casal, 1963 : Code 3/2, length 5 mm, Brazil (NT), Paraná, Parque Estadual de Vila Velha (-25.23 -50.00), DZUP. Determined by KAW. Photo by KAW at UFPR.

Fig. 1234— Lynchiatilla tacana Casal, 1963 : Code 1/2, length 7.5 mm, Argentina (NT), Tucumán, Rio Nio (-26.42 -64.93), FSCA. Determined by KAW. Photo by KAW at DPI .

Fig. 1235— Lynchiatilla cf. armanda (Smith, 1879) : Code 2/1, length 9 mm, Brazil (NT), Rio de Janeiro, 6 km NW de Itatiaia, PN Itatiaia, Hotel Simon (-22.43 -44.60), DZUP. Determined by KAW. Photo by KAW at UFPR. Fig. 1236— Lynchiatilla cf. parana Cambra, 2012 : Code 2/1, length 7.5 mm, Brazil (NT), São Paulo, Campos do Jordão, Parque Estadual (-22.73 -45.58), MZSP. Determined by KAW. Photo by G. Furman at UFPR.

Fig. 1237— Lynchiatilla cf. silvai Casal, 1963 : Code 2/1, length 4 mm, Brazil (NT), Minas Gerais, Caratinga (- 19.75 -41.83), DZUP. Determined by KAW. Photo by KAW at UFPR.

Mickelia Suárez, 1966 —There is no key to the species, but Suárez (1966) mentioned differences between the species. Fig. 1238— Mickelia cressoni Suárez, 1966 : Code 3/1, length 15 mm, Brazil (NT), Minas Gerais, 10 km south of Janaúba, Poções de Santa Cruz (-15.90 -43.34), DZUP. Determined by KAW. Photo by KAW at UFPR.

Fig. 1239— Mickelia sp. nov.: Code 4/1, length 14 mm, Brazil (NT), Further data not available, MPEG. Determined by KAW. Photo by KAW at UFPR.