Skorikovia Ovchinnikov, 2002

Williams, Kevin A., 2025, Color syndrome terminology for velvet ant females (Hymenoptera: Mutillidae and Myrmosidae), Zootaxa 5684 (1), pp. 1-238 : 168-175

Skorikovia Ovchinnikov, 2002 —The species were keyed by Lelej (2024b).

Fig. 2320— Skorikovia caucasica (Lelej, 1985) : Code 1/1, length 7.2 mm, Azerbaijan (PA), Ordubad (38.90 46.02). Holotype. Published in Lelej (2024).

Fig. 2321— Skorikovia elongata (Radoszkowski, 1885) : Code 1/1, length 7.5 mm, Tajikistan (PA), 40 km South Dushanbe, Chimbulak Vill. (43.13 77.08), CSCA. Determined by KAW. Photo by KAW at DPI .

Fig. 2322— Skorikovia maxim Lelej, 2024 : Code 1/1, length 11.5 mm, Unknown (PA), Further data not available. Paratype. Published in Lelej (2024).

Fig. 2323— Skorikovia radosdkovskii (Skorikov, 1935) : Code 3/2, length 6.8 mm, Russia (PA), Dagestan, Petrovkport, Makhachakala (42.98 47.45). Determined by A.S. Lelej. Published in Lelej (2024).

Fig. 2324— Skorikovia serta (Radoszkowski, 1885) : Code 3/2, length 8.4 mm, Russia (PA), Orenburg (51.83 55.20). Lectotype. Published in Lelej (2024).

Fig. 2325— Skorikovia trinotata (Costa, 1858): Code 2/1, length 6 mm, Italy (PA), Aosta, Ozein (45.68 7.24), FSCA. Determined by M. Romano. Photo by KAW at DPI.

Smicromyrme Thomson, 1870 —The most complete key available was written by Lelej (1985), other regional keys are also available: China (Chen 1957), Pacific Islands (Mickel 1935), Italy (Pagliano & Strumia 2007), Central Europe (Lelej & Schmid-Eger 2005), the Iberian Peninsula (Matias 2024), and Africa (Bischoff 1920). Until recently, the gender of this genus name was considered to be masculine, but is now recognized to be feminine (Brothers et al. 2025, in press).

Fig. 2326— Smicromyrme atrithorax (André, 1902): Code 3/1, length 8.5 mm, Azerbaijan (PA), Lerik, Talysh Mts. (38.65 48.38), FSCA. Determined by KAW. Photo by KAW at DPI .

Fig. 2327— Smicromyrme ausonia Invrea, 1950 : Code 2/1, length 4.6 mm, Italy (PA), Villanova, Albenga (44.05 8.14), CASC. Determined by G. Pagliano. Photo by KAW at CDFA.

Fig. 2328— Smicromyrme autonoe Mickel, 1934 : Code 2/1, length 4.1 mm, Philippines (OR), Palawan, Puerto Princesa (9.76 118.75), UMSP. Paratype. Photo by KAW at CDFA .

Fig. 2329— Smicromyrme basalis annularis Mickel, 1934 : Code 2/1, length 5.3 mm, Philippines (OR), Los Banos (14.16 121.25), UMSP. Determined by C.E. Mickel. Photo by KAW at CDFA.

Fig. 2330— Smicromyrme basalis basalis (Smith, 1879) : Code 2/1, length 5.8 mm, Philippines (OR), Alcate Victoria Madero (13.37 121.20), EMUS. Determined by B. Petersen. Photo by KAW at DPI.

Fig. 2331— Smicromyrme benefactrix (Turner, 1916) : Code 2/1, length 4.6 mm, Malawi (AT), Lake Nyasa, Monkey Bay (-14.08 34.91), UMSP. Determined by C.E. Mickel. Photo by KAW at CDFA.

Fig. 2332— Smicromyrme bicolor Hammer, 1962 : Code 1/1, length 4.5 mm, India (OR), Karnataka; Bangalore Urban; Bangalore , Allalsandra (12.97 77.58), NHMD. Identifier unknown. Published in Boutin & Vilhelmsen (2024) .

Fig. 2333— Smicromyrme brothersi Lelej, 2024 : Code 2/1, length 5.8 mm, Indonesia (OR), Lombok, near Senggigi (-8.45 116.05), IBSS. Holotype. Published in Lelej (2024a).

Fig. 2334— Smicromyrme burgeri Lelej, 2020 : Code 3/2, length 6 mm, India (OR), Karnataka, 15 km SW Bangalore (12.85 77.36), MZUF. Holotype. Published in Lelej (2020a).

Fig. 2335— Smicromyrme carosellii Invrea, 1936 : Code 1/2, length 7 mm, Kenya ** (AT), Mwingi, Kangoride vadi (-0.94 38.06), FSCA. Determined by M. Romano. Photo by KAW at DPI.

Fig. 2336— Smicromyrme coromandelica (Motschulsky, 1863) : Code 1/1, length 5.3 mm, India (OR), Coimbatore (11.02 76.96), CSCA. Determined by KAW. Photo by KAW at CDFA.

Fig. 2337— Smicromyrme deidamia (Smith, 1857) : Code 2/1, length 8.9 mm, Malaysia (OR), Sabah, Sandakan (5.84 118.12), UMSP. Determined by C.E. Mickel. Photo by KAW at CDFA .

Fig. 2338— Smicromyrme degiovannii Pagliano & Matteini Palmerini, 2014 : Code 3/1, length 5.2 mm, Tunisia (PA), 10 km SE Mat-mata (33.50 10.02), CASC. Determined by G. Pagliano. Photo by KAW at CDFA.

Fig. 2339— Smicromyrme difficilis Bischoff, 1921 : Code 2/1, length 4 mm, Democratic Republic of the Congo ** (AT), Shaba, Kisanga Station (-7.84 26.36), CSCA. Determined by KAW. Photo by KAW at DPI.

Fig. 2340— Smicromyrme dolichothoracicus Bischoff, 1920 : Code 2/1, length 4 mm, Nigeria ** (AT), Uyo (5.03 7.91), PURC. Determined by KAW. Photo by KAW at DPI.

Fig. 2341— Smicromyrme durga (Bingham, 1897) : Code 3/2, length 7.8 mm, India (OR), West Bengal; Galalea (?) (22.98 87.75), NHMD. Identifier unknown. Published in Boutin & Vilhelmsen (2024) .

Fig. 2342— Smicromyrme enippe (Péringuey, 1899) : Code 1/1, length 5.5 mm, Democratic Republic of the Congo ** (AT), Shaba, Kisanga Station (-7.84 26.36), CSCA. Determined by KAW. Photo by KAW at DPI.

Fig. 2343— Smicromyrme fasciaticollis (Spinola, 1843) : Code 3/2, length 8 mm, Italy (PA), Lampedusa, Guitgia (35.50 12.60), FSCA. Determined by M. Romano. Photo by KAW at DPI.

Fig. 2344— Smicromyrme ferruginata Bischoff, 1921 : Code 1/2, length 7.9 mm, South Africa (AT), Pretoria, Transvaal Boekenhouts Kloof (-25.53 28.47), CMNH. Determined by KAW. Photo by KAW at CDFA.

Fig. 2345— Smicromyrme frankburgeri Schmid-Eger, 2022 : Code 2/1, length 4.5 mm, Germany (PA), Hessen, Darmstadt (49.86 8.64), CSCA. Determined by KAW. Photo by KAW at CDFA .

Fig. 2346— Smicromyrme frederici (André, 1898): Code 1/1, length 3.6 mm, India (OR), Utar Pradesh, New Dehli, 28 37.8’N 77 9.8’E (28.63 77.17), UCRC. Determined by KAW. Photo by KAW at CDFA .

Fig. 2347— Smicromyrme fura anthraceipes Mickel, 1934 : Code 2/1, length 8.4 mm, Philippines (OR), Luzon , Benguet, Baguio (16.40 120.60), USNM. Holotype. Photo by KAW at CDFA.

Fig. 2348— Smicromyrme fura fura Mickel, 1934 : Code 2/1, length 6.5 mm, Philippines (OR), Lagos, Los Banos (14.17 121.24), EMUS. Determined by B. Petersen. Photo by KAW at DPI.

Fig. 2349— Smicromyrme gineri Invrea, 1953 : Code 2/1, length 4.4 mm, Morocco (PA), Sidi Bienzane, foce Oued Massa Agadir (30.40 -9.58), CASC. Determined by G. Pagliano. Photo by KAW at CDFA.

Fig. 2350— Smicromyrme gridellii (Invrea, 1960) : Code 1/1, length 3.5 mm, Egypt (PA), El Kharga, Kharga Oasis (25.40 30.60), USNM. Identifier unknown. Photo by KAW at DPI.

Fig. 2351— Smicromyrme gudenzii Pagliano & Matteini Palmerini, 2014 : Code 3/2, length 6.7 mm, Tunisia (PA), Douz (33.47 9.02), GPC. Holotype. Photo by A. Degiovanni in Pagliano & Matteini (2014).

Fig. 2352— Smicromyrme helarctos Williams in Williams et al., 2019: Code 2/1, length 3.5 mm, Thailand (OR), Kanchanaburi, Phalad Waterfall (14.24 99.34), CSCA. Determined by KAW. Photo by KAW at CDFA .

Fig. 2353— Smicromyrme hyale (Mickel, 1934) : Code 2/1, length 5.3 mm, Philippines (OR), Mindanao, Surigao (8.54 126.11), UMSP. Paratype. Photo by KAW at CDFA .

Fig. 2354— Smicromyrme innesi (André, 1893): Code 3/1, length 6.4 mm, Egypt (PA), Barrage (27.08 32.00), USNM. Determined by Alfieri. Photo by KAW at CDFA.

Fig. 2355— Smicromyrme kuanfuana Tsuneki, 1972 : Code 2/1, length 4.3 mm, Taiwan (OR), Hualien Pref., Kuangfu (23.64 121.42), USNM. Holotype. Photo by KAW at CDFA.

Fig. 2356— Smicromyrme lewisi Mickel, 1935 : Code 2/1, length 5.9 mm, Japan (PA), Yorii , Saitama (36.12 139.19), CASC. Identifier unknown. Photo by KAW at CDFA .

Fig. 2357— Smicromyrme lybica Invrea, 1932 : Code 2/1, length 3.8 mm, Tunisia ** (PA), Tozeur (33.92 8.12), CASC. Determined by G. Pagliano. Photo by KAW at CDFA .

Fig. 2358— Smicromyrme lochia Mickel, 1937 : Code 2/1, length 5.7 mm, Malaysia (OR), Penang (5.43 100.30), UMSP. Paratype. Photo by KAW at CDFA .

Fig. 2359— Smicromyrme longigena Bischoff, 1921 : Code 1/1, length 6.1 mm, South Africa ** (AT), KwaZulu-Natal, Tembe Elephant Park (-27.03 32.38), FSCA. Determined by G. Pagliano. Photo by KAW at DPI .

Fig. 2360— Smicromyrme mareotica Invrea, 1951 : Code 3/1, length 9.5 mm, Egypt ** (PA), Burj el Arab (30.90 29.50), USNM. Identifier unknown. Photo by KAW at DPI.

Fig. 2361— Smicromyrme margheritae Pagliano & Matteini Palmerini, 2014 : Code 2/1, length 3.8 mm, Tunisia (PA), Douz (33.47 9.02), GPC. Holotype. Photo by A. Degiovanni in Pagliano & Matteini (2014).

Fig. 2362— Smicromyrme mauromoustakisi Invrea, 1940 : Code 1/1, length 7.5 mm, Cyprus (PA), Limassol (34.71 33.02), USNM. Identifier unknown. Photo by KAW at DPI .

Fig. 2363— Smicromyrme melpomene hecuba (Péringuey, 1898) : Code 1/1, length 7.1 mm, South Africa (AT), Cape Province, Wellington, Rooshoek (-33.57 19.07), EMUS. Determined by F.J. Suárez. Photo by KAW at CDFA.

Fig. 2364— Smicromyrme neglecta Hammer, 1962 : Code 1/1, length 5 mm, India (OR), Pondichery, Karaikal (10.93 79.84), CASC. Determined by G. Pagliano. Photo by KAW at CDFA.

Fig. 2365— Smicromyrme nigriceps Nonveiller, 1959 : Code 2/1, length 4 mm, Greece (PA), Creta, Koutsounari, Terapetra (35.01 25.74), FSCA. Determined by M. Romano. Photo by KAW at DPI .

Fig. 2366— Smicromyrme nigrigena (André, 1894): Code 1/1, length 4.4 mm, India (OR), Karnataka; Bangalore Rural; Bangalore (12.97 77.58), NHMD. Identifier unknown. Published in Boutin & Vilhelmsen (2024) .

Fig. 2367— Smicromyrme nonveilleri 1 Invrea, 1954: Code 2/1, length 6.2 mm, Greece (PA), Assiros (40.82 23.03), CASC. Determined by G. Pagliano. Photo by KAW at CDFA.

Fig. 2368— Smicromyrme nonveilleri 2 Invrea, 1954: Code 1/1, length 5.1 mm, Turkey ** (PA), Manisa (38.61 27.43), NHMUK. Determined by KAW. Photo by KAW at CDFA .

Fig. 2369— Smicromyrme ocellata (Saussure, 1867) : Code 1/1, length 8 mm, Sri Lanka (OR), Wilpattu NP, 0.5 mi NE Kokmotte (8.45 80.05), USNM. Determined by B. Petersen. Photo by KAW at CDFA.

Fig. 2370— Smicromyrme parthenia (Cameron, 1900) : Code 2/1, length 6 mm, India (OR), Karnataka, Gersoppa (14.24 74.65), EMUS. Determined by B. Petersen. Photo by KAW at DPI.

Fig. 2371— Smicromyrme partita obscurithorax (André, 1903): Code 3/1, length 6.3 mm, Morocco (PA), 15 km North Ifrane (33.64 -5.11), CASC. Determined by KAW. Photo by KAW at CDFA.

Fig. 2372— Smicromyrme partita partita (Klug, 1835) : Code 3/2, length 6.5 mm, Italy (PA), Liguria, Ortovero (44.05 8.09), FSCA. Identifier unknown. Photo by KAW at DPI .

Fig. 2373— Smicromyrme peregrina (Cameron, 1897) : Code 1/1, length 4 mm, India (OR), Pondichery (11.94 79.81), FSCA. Determined by KAW. Photo by KAW at DPI.

Fig. 2374— Smicromyrme perisii (Sichel & Radoszkowski, 1869) : Code 2/1, length 5 mm, Italy (PA), Sardinia, Magomadas (40.26 8.53), FSCA. Determined by M. Romano. Photo by KAW at DPI .

Fig. 2375— Smicromyrme posthuma (Cameron, 1898) : Code 1/1, length 6.1 mm, India ** (OR), Karnataka; Chikkamagaluru; Mudigere area (13.14 75.64), NHMD. Identifier unknown. Published in Boutin & Vilhelmsen (2024) .

Fig. 2376— Smicromyrme pruinosa Bischoff, 1921 : Code 4/1, length 6.9 mm, Zimbabwe (AT), Pakasa (-16.00 30.00), UMSP. Identifier unknown. Photo by KAW at CDFA.

Fig. 2377— Smicromyrme raptrix Bischoff, 1921 : Code 3/2, length 12 mm, South Africa (AT), Further data not available, Collection unknown. Identifier unknown. Photo by M. Romano in Pagliano et al. (2020).

Fig. 2378— Smicromyrme riccardoi Pagliano & Matteini Palmerini, 2014 : Code 3/2, length 4.8 mm, Tunisia (PA), Elhamma (33.89 9.88), GPC. Holotype. Photo by A. Degiovanni in Pagliano & Matteini (2014).

Fig. 2379— Smicromyrme rufescens (André, 1901): Code 1/2, length 4.7 mm, South Africa (AT), Free State, Bothaville (-27.38 26.62), MNHN. Holotype. Photo by D.J. Brothers.

Fig. 2380— Smicromyrme ruficollis ceresae Invrea, 1952 : Code 3/2, length 5.5 mm, France (PA), Corsica, Ghisonaccia (42.02 9.40), CSCA. Determined by G. Pagliano. Photo by KAW at DPI.

Fig. 2381— Smicromyrme ruficollis ruficollis (Fabricius, 1793) : Code 2/1, length 4 mm, Spain (PA), Pto.Sta.Ha (Valdelagrana) (36.58 -6.21), CSCA. Determined by G. Pagliano. Photo by KAW at DPI.

Fig. 2382— Smicromyrme rufipes (Fabricius, 1787) : Code 2/1, length 3.5 mm, Estonia ** (PA), Tartu (58.39 26.72), FSCA. Determined by KAW. Photo by KAW at DPI .

Fig. 2383— Smicromyrme sexmaculata Hammer, 1962 : Code 2/1, length 4.3 mm, India (OR), Nedungadu PO, Tanjore Dt. (10.97 79.78), CSCA. Determined by KAW. Photo by KAW at CDFA.

Fig. 2384— Smicromyrme sicana (De Stefani, 1887) : Code 1/1, length 4.5 mm, Italy (PA), Liguria, Casanova Lerrone (44.03 8.05), FSCA. Identifier unknown. Photo by KAW at DPI .

Fig. 2385— Smicromyrme signata (Klug, 1829) : Code 1/1, length 4.8 mm, Egypt (PA), Phorette [Rasheed] (31.39 30.41), USNM. Determined by Alfieri. Photo by KAW at CDFA.

Fig. 2386— Smicromyrme songkhwae Sittichaya & Williams in Sittichaya et al., 2022: Code 2/1, length 2.9

mm, Thailand (OR), Phitsanulok, Chat Trakan, Phu Soi Dao National Park (17.70 100.95), THNHM. Holotype. Published in Thaochan et al. (2022).

Fig. 2387— Smicromyrme suarezi Matias, 2024 : Code 3/2, length 5.9 mm, Portugal (PA), Castelo Branco, Alfrivida (39.72 -7.51). Holotype. Published in Matias (2024).

Fig. 2388— Smicromyrme suberrata* Invrea, 1957 : Code 2/1, length 4.5 mm, Italy (PA), Sicily, Portella di Femmina Morta (37.92 14.65), CSCA. Determined by G. Pagliano. Photo by KAW at DPI. The female is not currently known in the literature. This female was identified as S. suberratus by G. Pagliano, who studied velvet ants extensively in the Mediterranean region, and was found in a location where males of S. suberratus occur. Fig. 2389— Smicromyrme subfasciata (André, 1901): Code 2/1, length 6.5 mm, South Africa (AT), Eastern Cape, Gqeberha (-33.96 25.60), USNM. Determined by C.E. Mickel. Photo by KAW at CDFA.

Fig. 2390— Smicromyrme substita (André, 1896): Code 2/1, length 7.5 mm, India (OR), Sikkim (27.34 88.61), NHMD. Identifier unknown. Published in Boutin & Vilhelmsen (2024) .

Fig. 2391— Smicromyrme sulcisia Invrea, 1955 : Code 3/2, length 6 mm, Italy (PA), Liguria, Ortovero (44.05 8.10), CSCA. Determined by G. Pagliano. Photo by KAW at DPI .

Fig. 2392— Smicromyrme tekensis Skorikov, 1935 : Code 3/2, length 6.3 mm, Kazhakstan (PA), Taldykorgan (45.02 78.38), USNM. Determined by A.S. Lelej. Photo by KAW at CDFA.

Fig. 2393— Smicromyrme tenasserimica (Magretti, 1892) : Code 2/1, length 4.3 mm, India ** (OR), Karnataka; Kanara; Kanara (14.88 74.58), NHMD. Identifier unknown. Published in Boutin & Vilhelmsen (2024) .

Fig. 2394— Smicromyrme tettensiella Bischoff, 1921 : Code 2/1, length 11 mm, Democratic Republic of the Congo ** (AT), Katanga, Kafakumba (-9.68 23.76), RBINS. Determined by KAW. Photo by KAW at CDFA .

Fig. 2395— Smicromyrme thaochani Williams in Williams et al., 2019: Code 2/1, length 8.8 mm, Myanmar (OR), Sagaig (22.32 94.48), CASC. Determined by KAW. Photo by KAW at CDFA.

Fig. 2396— Smicromyrme triguttata Mickel, 1933 : Code 2/1, length 4.2 mm, Thailand (OR), Kanchanaburi, Vimarndin farm stay (14.66 98.52), CSCA. Determined by KAW. Photo by KAW at CDFA .

Fig. 2397— Smicromyrme turanica (Morawitz, 1893) : Code 2/1, length 7 mm, Greece (PA), Creta, Lefka, Ori Mts. , Omalos Plateau (35.07 25.46), FSCA. Determined by M. Romano. Photo by KAW at DPI .

Fig. 2398— Smicromyrme vladani Nonveiller, 1972 : Code 2/1, length 6 mm, Greece (PA), Creta, Koutsomatodos, Topolia (35.43 23.69), FSCA. Determined by M. Romano. Photo by KAW at DPI .

Fig. 2399— Smicromyrme williamsi Terine & Girish Kumar, 2024 : Code 1/1, length 6.9 mm, India (OR), Anamalai Hills, Cinchona (10.30 77.00), CASC. Determined by KAW. Photo by KAW at CDFA.

Fig. 2400— Smicromyrme cf. constanceae (Cameron, 1892) : Code 3/2, length 4.5 mm, India (OR), Narmada R. (22.50 77.05), NHMUK. Determined by KAW. Photo by KAW at DPI.

Fig. 2401— Smicromyrme cf. maidli Bischoff, 1921 : Code 2/1, length 4.5 mm, South Africa (AT), Cape Town (- 33.95 18.43), USNM. Determined by KAW. Photo by KAW at DPI .

Fig. 2402— Smicromyrme cf. omphale (Péringuey, 1898) : Code 2/1, length 5 mm, South Africa (AT), Cape Town (-33.95 18.43), USNM. Determined by KAW. Photo by KAW at DPI .

Fig. 2403— Smicromyrme cf. rufescens (André, 1901): Code 1/4, length 4 mm, South Africa (AT), Transvaal, Nylsvley Nature Reserve, Aylstroom & Naboomspruit (-24.65 28.68), FSCA. Determined by KAW. Photo by KAW at DPI.

Fig. 2404— Smicromyrme sp. nov. 1: Code 2/1, length 6.6 mm, Nepal (OR), near Pokhara (28.23 83.98), RMNH. Determined by KAW. Photo by KAW at CDFA.

Fig. 2405— Smicromyrme sp. nov. 2: Code 3/1, length 5.9 mm, Sri Lanka (OR), Northern Province, Tharakunda, 4 mi NW Mannar (9.02 79.87), MZLU. Determined by KAW. Photo by KAW at CDFA.

Fig. 2406— Smicromyrme sp. nov. 3: Code 1/1, length 5 mm, Egypt (PA), Kom Awshim (29.38 31.80), USNM. Determined by KAW. Photo by KAW at DPI.

Fig. 2407— Smicromyrme sp. nov. 4: Code 3/2, length 5.1 mm, India (OR), Tamil Nadu, 41 km E Kodaikanal (10.25 77.85), CASC. Determined by KAW. Photo by KAW at CDFA .