Karlidia Lelej, 1999

Williams, Kevin A., 2025, Color syndrome terminology for velvet ant females (Hymenoptera: Mutillidae and Myrmosidae), Zootaxa 5684 (1), pp. 1-238 : 135-139

Karlidia Lelej, 1999 —This is a monotypic genus.

Fig. 1848— Karlidia peterseni Lelej in Lelej & Krombein, 1999: Code 2/1, length 7.5 mm, Thailand (OR), Chiang Mai, Doi Suthep (18.82 98.89), BPBM. Determined by KAW. Photo by KAW at DPI .

Odontomutilla Ashmead, 1899 —There are some regional keys to species in this genus: China (Chen 1957), Pacific Islands (Mickel 1935), India (Terine & Girish Kumar 2024), and mainland Africa (Bischoff 1920).

Fig. 1849— Odontomutilla aegrota (Cameron, 1898) : Code 4/1, length 14 mm, Zimbabwe (AT), Salisbury (-17.82 31.05), FSCA. Determined by D.J. Brothers. Photo by KAW at DPI.

Fig. 1850— Odontomutilla australica André, 1903: Code 4/1, length 10.9 mm, Australia (AU), Queensland, Cairns (-16.92 145.78), UMSP. Compared with type by C.E. Mickel. Photo by KAW at CDFA .

Fig. 1851— Odontomutilla calida André, 1908: Code 2/1, length 18 mm, Democratic Republic of the Congo ** (AT), Shabu Reg., Kapanda (-8.99 26.75), EMUS. Identifier unknown. Photo by KAW at CDFA.

Fig. 1852— Odontomutilla cordigera (Sichel & Radoszkowski, 1869) : Code 1/1, length 10.8 mm, Thailand (OR), Chiang Mai; Mueang Chiang Mai; Doi Suthep (18.81 98.92), NHMD. Identifier unknown. Published in Boutin & Vilhelmsen (2024) .

Fig. 1853— Odontomutilla familiaris familiaris (Smith, 1857) : Code 2/1, length 15 mm, Philippines (OR), Misamis Or., Balason (8.70 125.13), BPBM. Determined by KAW. Photo by KAW at DPI.

Fig. 1854— Odontomutilla fracta fracta (Saussure, 1891) : Code 2/1, length 8.8 mm, Tanzania ** (AT), Selous Game reserve (-9.00 37.50), NHMUK. Determined by KAW. Photo by KAW at CDFA.

Fig. 1855— Odontomutilla fracta ocularis Bischoff, 1920 : Code 2/1, length 8.8 mm, Uganda ** (AT), West Mengo, Entebbe (0.05 32.46), RMNH. Determined by KAW. Photo by KAW at CDFA.

Fig. 1856— Odontomutilla fracta unifasciata Bischoff, 1920 : Code 2/1, length 8.5 mm, Tanzania (AT), Kigonsera (-10.80 35.07), RMNH. Determined by KAW. Photo by KAW at DPI.

Fig. 1857— Odontomutilla grossa Mickel, 1935 : Code 1/1, length 16.7 mm, Indonesia ** (OR), Borneo, Mowong (0.76 109.43), UMSP. Holotype. Photo by KAW at CDFA.

Fig. 1858— Odontomutilla horida (Smith, 1879) : Code 2/1, length 10.1 mm, South Africa ** (AT), Transvall, Shilouvane (-24.04 30.28), BMNHB. Determined by KAW. Photo by KAW at CDFA.

Fig. 1859— Odontomutilla inanis Mickel, 1935 : Code 2/1, length 11 mm, Papua New Guinea (AU), Northeast Karimui (-6.50 144.82), BPBM. Determined by KAW. Photo by KAW at DPI.

Fig. 1860— Odontomutilla jaferpaloti Terine & Girish Kumar, 2024 : Code 3/1, length 14.3 mm, India (OR), No further data, DEI. Determined by KAW. Photo by KAW at CDFA.

Fig. 1861— Odontomutilla listopadovi Lelej, 2023 : Code 2/1, length 12.1 mm, Myanmar (OR), Mandalay, 2 km South Popa (21.80 96.10), IBSS. Holotype. Published in Lelej (2023).

Fig. 1862— Odontomutilla manifesta (Smith, 1859) : Code 2/4, length 14 mm, Papua New Guinea (AU), Gulf, KakoRondonia (-7.85 143.52), UCDC. Determined by KAW. Photo by KAW at CDFA .

Fig. 1863— Odontomutilla mickeli Lelej, 2005 : Code 2/1, length 15 mm, Indonesia (OR), Java, Djampang Tengah (-7.06 106.79), RMNH. Determined by KAW. Photo by KAW at CDFA .

Fig. 1864— Odontomutilla mocquerysi (André, 1895): Code 4/1, length 9.2 mm, Democratic Republic of the Congo (AT), Katanga, Kafakumba (-9.68 23.76), RBINS. Determined by KAW. Photo by KAW at CDFA .

Fig. 1865— Odontomutilla notata chione (Péringuey, 1898) : Code 2/1, length 9.4 mm, South Africa (AT), Rustenburg S (-25.68 27.25), UMMZ. Determined by KAW. Photo by KAW at CDFA.

Fig. 1866— Odontomutilla notata nigrita Bischoff, 1920 : Code 4/1, length 9.5 mm, Kenya ** (AT), Crescent Island, Lake Naivasha (-0.77 36.40), CSCA. Determined by KAW. Photo by KAW at DPI.

Fig. 1867— Odontomutilla notata notata (Lepeletier, 1845) : Code 2/1, length 11 mm, South Africa (AT), Basutoland, Mokhorburg? (-29.29 29.06), MZLU. Determined by KAW. Photo by KAW at CDFA.

Fig. 1868— Odontomutilla notata umtalina Bischoff, 1920 : Code 2/1, length 15 mm, Zimbabwe (AT), Lower Gwelo (-19.30 29.20), CSCA. Determined by KAW. Photo by KAW at DPI.

Fig. 1869— Odontomutilla ovata (Sichel & Radoszkowski, 1869) : Code 1/1, length 9.3 mm, South Africa (AT), S. Cape, 30 km S Steytlerville (-33.56 24.34), EMUS. Determined by KAW. Photo by KAW at CDFA.

Fig. 1870— Odontomutilla ovataeformis Bischoff, 1920 : Code 1/1, length 13.5 mm, Namibia (AT), Gross Spitzkoppe (-21.94 15.38), CSCA. Determined by KAW. Photo by KAW at CDFA.

Fig. 1871— Odontomutilla paderua Das & Girish Kumar, 2017 : Code 1/1, length 11 mm, India (OR), Jaiphur (26.91 75.78), UMMZ. Determined by KAW. Photo by KAW at CDFA.

Fig. 1872— Odontomutilla pulchrina (Smith, 1855) : Code 1/1, length 15.8 mm, India (OR), Karnataka; Shivamogga; Agumbe Ghat. (13.51 75.10), NHMD. Identifier unknown. Published in Boutin & Vilhelmsen (2024) .

Fig. 1873— Odontomutilla rhodesiaca Bischoff, 1920 : Code 2/1, length 12.7 mm, Zimbabwe (AT), Shiloh (-17.80 30.99), SAMC. Holotype. Photo by L. Huntly at SAMC (van Noort 2025).

Fig. 1874— Odontomutilla saussurei (Sichel & Radoszkowski, 1869) : Code 2/1, length 16.4 mm, Liberia ** (AT), Marshall Territory (6.15 -10.39), CMNH. Determined by KAW. Photo by KAW at CDFA.

Fig. 1875— Odontomutilla semifasciata (André, 1896): Code 2/1, length 11.5 mm, Indonesia (OR), Southeast Sulawesi; Kolaka; Mengkoka (-3.63 121.25), NHMD. Identifier unknown. Published in Boutin & Vilhelmsen (2024) .

Fig. 1876— Odontomutilla sikkimensis André, 1903: Code 2/1, length 9.7 mm, India (OR), Uttarakhand; Almora ; Almora (29.59 79.65), NHMD. Identifier unknown. Published in Boutin & Vilhelmsen (2024) .

Fig. 1877— Odontomutilla sinensis (Smith, 1855) : Code 2/1, length 15.2 mm, China (OR), W.Hupeh, Lichuan, Hsiao-Ho (30.30 108.90), CASC. Determined by KAW. Photo by KAW at CDFA.

Fig. 1878— Odontomutilla spectra (Bingham, 1908) : Code 1/1, length 12.2 mm, India (OR), Madhya Pradesh; Mandla; Mandla (22.60 80.37), NHMD. Identifier unknown. Published in Boutin & Vilhelmsen (2024) .

Fig. 1879— Odontomutilla tamensis (Cameron, 1906) : Code 2/1, length 11 mm, Indonesia ** (AU), Papua, Waenda near Jayapura, Kampus Walker (-2.60 140.70), RMNH. Determined by KAW. Photo by KAW at CDFA. Fig. 1880— Odontomutilla thymele Mickel, 1935 : Code 1/1, length 13 mm, Indonesia ** (OR), Sumatra, Aceh, Ketambe (3.69 97.72), RMNH. Determined by KAW. Photo by KAW at CDFA.

Fig. 1881— Odontomutilla tomyris (Péringuey, 1899) : Code 2/1, length 15.6 mm, Botswana ** (AT), Tlokweng (-24.67 25.98), CMNH. Determined by KAW. Photo by KAW at CDFA.

Fig. 1882— Odontomutilla trichocondyla (André, 1894): Code 1/1, length 11.2 mm, India (OR), Karnataka; Bangalore Urban; Bangalore (12.97 77.58), NHMD. Identifier unknown. Published in Boutin & Vilhelmsen (2024). Fig. 1883— Odontomutilla urania (Smith, 1857) : Code 1/1, length 11 mm, Indonesia ** (OR), Sumatra, Gn. Betoeng (-2.83 104.23), RMNH. Determined by KAW. Photo by KAW at CDFA.

Fig. 1884— Odontomutilla uranioides Mickel, 1933 : Code 1/1, length 15 mm, Vietnam (OR), Fyan (11.88 108.20), BPBM. Determined by KAW. Photo by KAW at DPI.

Fig. 1885— Odontomutilla zimrada kenyana Bischoff, 1920 : Code 2/1, length 14 mm, Tanzania ** (AT), Kia (- 3.41 37.06), CSCA. Determined by KAW. Photo by KAW at DPI.

Fig. 1886— Odontomutilla zimrada lukulediana Strand, 1917 : Code 4/1, length 16.7 mm, Tanzania (AT), Kigonsera (-10.79 35.07), BMNHB. Determined by KAW. Photo by KAW at CDFA.

Fig. 1887— Odontomutilla aff. grossa Mickel, 1935 : Code 1/1, length 10 mm, Laos (OR), Xieng Khouang, Ban Thaviang (18.98 103.44), BMNHB. Determined by KAW. Photo by KAW at CDFA.

Fig. 1888— Odontomutilla cf. maxima Bischoff, 1920 : Code 2/1, length 13.5 mm, Zimbabwe (AT), Inyangombe Falls (-8.17 32.41), FSCA. Determined by KAW. Photo by KAW at CDFA.

Fig. 1889— Odontomutilla aff. mickeli Lelej, 2005 : Code 2/1, length 13 mm, Indonesia (OR), Sumba (-9.57 119.39), RMNH. Determined by KAW. Photo by KAW at CDFA.

Fig. 1890— Odontomutilla aff. microcephala (André, 1896): Code 2/1, length 13.5 mm, South Africa (AT), Transvaal, 15 km E Karimo Station (-25.52 31.24), CMNH. Determined by KAW. Photo by KAW at CDFA.

Fig. 1891— Odontomutilla aff. mocqsuerysi (André, 1895): Code 2/1, length 12 mm, Kenya (AT), Gedi (-3.31 40.01), RMNH. Determined by KAW. Photo by KAW at DPI.

Fig. 1892— Odontomutilla sp. nov. 1: Code 2/1, length 10.4 mm, Kenya (AT), Further data not available, Unknown. Determined by A.S. Lelej. Photo by A.S. Lelej.

Fig. 1893— Odontomutilla sp. nov. 2: Code 4/3, length 9.7 mm, Tanzania (AT), Usambara (-4.90 38.30), Unknown. Determined by A.S. Lelej. Photo by A.S. Lelej.

Fig. 1894— Odontomutilla sp. nov. 3: Code 1/1, length 7.8 mm, Thailand (OR), Phitsanulok (16.84 100.87), EMUS. Determined by KAW. Photo by KAW at CDFA.

Fig. 1895— Odontomutilla sp. nov. 4: Code 1/1, length 10.9 mm, Thailand (OR), Chaiyaphum, Tat Tone NP (15.98 102.04), EMUS. Determined by KAW. Photo by KAW at CDFA.

Fig. 1896— Odontomutilla sp. nov. 5: Code 3/1, length 9.3 mm, India (OR), Tamil Nadu; Coimbatore; Coimbatore (11.00 76.96), NHMD. Identifier unknown. Published in Boutin & Vilhelmsen (2024) .

Fig. 1897— Odontomutilla sp. nov. 6 (male): Code 2/1, length 11.5 mm, Tanzania (AT), Mikumi (-7.40 36.99), RMNH. Determined by KAW. Photo by KAW at DPI.

Fig. 1898— Odontomutilla sp. nov. 7: Code 4/1, length 9 mm, Ivory Coast (AT), Guiglo, 12 km E Pauleoula (5.82 -7.29), RMNH. Determined by KAW. Photo by KAW at CDFA.

Fig. 1899— Odontomutilla sp. nov. 8: Code 1/1, length 8.4 mm, Iran (PA), Further data not available, Unknown. Determined by A.S. Lelej. Photo by A.S. Lelej.

Yamanetilla Lelej, 1996 —No published keys include all of the species, but Chen (1957) and Mickel (1935) are the best starting points.

Fig. 1900— Yamanetilla andromeda (Mickel, 1934) : Code 2/1, length 5.5 mm, Philippines (OR), Mindanao, S.Luis Calapan (7.77 122.81), EMUS. Determined by B. Petersen. Photo by KAW at DPI.

Fig. 1901— Yamanetilla cassiope (Smith, 1857) : Code 2/1, length 6 mm, Malaysia (OR), Sungei Linam [?Kampang Sungei Lui] (3.17 101.88), BPBM. Determined by KAW. Photo by KAW at DPI.

Fig. 1902— Yamanetilla nipponica (Tsuneki, 1972) : Code 2/1, length 5.8 mm, Japan (PA), Urayama aguechi, Saitama (35.91 139.07), CASC. Identifier unknown. Photo by KAW at CDFA .

Fig. 1903— Yamanetilla pedaria (Mickel, 1934) : Code 2/1, length 5 mm, Philippines (OR), Palawan, Puerto Princesa (9.76 118.75), EMUS. Determined by B. Petersen. Photo by KAW at DPI .

Fig. 1904— Yamanetilla yamanei Lelej, 2024 : Code 2/1, length 6.2 mm, Indonesia (OR), Sulawesi, Gorontalo, Mts. Tilogkabilu (0.62 123.16), SKYC. Holotype. Published in Lelej (2024a).

Fig. 1905— Yamanetilla sp. nov. 1: Code 2/1, length 6 mm, Bhutan (OR), Thimhu, Lungtenphu (27.46 89.66), RMNH. Determined by KAW. Photo by KAW at CDFA.

Fig. 1906— Yamanetilla sp. nov. 2: Code 2/1, length 7 mm, Nepal (OR), Janakpur, Dolakha (27.61 86.08), BMNHB. Determined by KAW. Photo by KAW at CDFA.

Fig. 1907— Yamanetilla sp. nov. 3 (KhmerF1): Code 2/1, length 5.2 mm, Thailand (OR), Loei (16.84 101.69), EMUS. Determined by KAW. Photo by KAW at CDFA.

Fig. 1908— Yamanetilla sp. nov. 4 (KhmerF2): Code 2/1, length 5.8 mm, Thailand (OR), Loei (19.50 101.34), EMUS. Determined by KAW. Photo by KAW at CDFA.

SUBFAMILY MYRMILLINAE Bischoff, 1920 —Keys to genera are available for the Oriental (Lelej 2005) and Palaearctic (Lelej 2002, Turrisi et al. 2015) realms. For Africa, Bischoff (1920) is a starting point, but subsequent papers should be consulted (Nonveiller 1975, 1996a, 1996b).