Prostoma graecense, Bohmig, 1893

Kajihara, Hiroshi, 2025, A taxonomic revision of the freshwater monostiliferous hoplonemertean genus Prostoma Dugès, 1828 (Nemertea: Eumonostilifera): a radical solution or an over-lumping?, Zootaxa 5646 (4), pp. 451-500 : 467-468

9. graecense Böhmig, 1893

Source. In the binomen Tetrastemma graecensis [sic] by Böhmig in Mojsisovics (1893: LXXXII [82]).

Type locality. A waterbody in a botanical garden in Graz , Styria, Austria ( Fig. 9 ) .

Etymology. Latin adjective, graecensis , - is, - e, meaning “from Graz”, while in many other cases it also refers to “from Greece ”.

Remarks. The species name “ Tetrastemma graecensis ” was proposed during a presentation by Ludwig Böhmig at a meeting of the zoology section of the Naturwissenschaftlicher Verein für Steiermark [Natural-Science Association of Styria] on 17 December 1892. The contents of Böhmig’s talk were included in a report by Mojsisovics (1893: LXXXII–LXXXIX [82–89]). The author of the name graecensis ( graecense ) is therefore Böhmig, rather than Mojsisovics, according to Article 50.2. Strict adherence to Recommendation 51E would be that the author of graecense be cited as ‘Böhmig in Mojsisovics, 1893 ’. In the present paper, however, it is simplified as ‘Böhmig, 1893’. Mojsisovics’ article appeared in Volume 29 of the Mittheilungen des naturwissenschaftlichen Vereines für Steiermark, which contained various reports of the Verein ’s activity in 1892 (Jahrgang 1892, ‘business year 1892’). While “ Jahrgang 1892 ” is printed on the title page of the volume, the actual publication year, 1893, is also printed on the same title page. The publication date of the name graecensis is thus 1893, although it has long been erroneously ascribed to 1892 in all subsequent literature to the best of my knowledge (e.g., Bürger 1895: 590; Bürger 1904: 69; Gibson & Moore 1976: 200; Gibson 1995: 516; Chernyshev et al. 1998: 61; Kajihara 2007: 311).

Prostoma graecense was fully described by Böhmig (1898) and has been regarded as valid by many researchers (e.g., Stiasny-Wijnhoff 1938: 227; Gibson & Moore 1976: 203; Chernyshev et al. 1998: 61; Sundberg & Gibson 2008: 62). The invalidation of P. graecense as a junior synonym of P. clepsinoides will likely cause confusion, but the Principle of Priority is strictly adhered in this study (see Discussion below). Gibson (1995: 495) erroneously indicated Prostoma graecense as the type species for Prostoma .

Figures

FIGURE 9. World map showing distribution of 25 nominal species of freshwater monostiliferous hoplonemerteans indicated by boxes and respective species-group names (three subterranean species are indicated with ‘S’ in parentheses); circles, diamond, and triangles indicate localities of DNA-barcoded individuals; circles, Haplogroup 1 from the UK, Sweden (SE), Italy (IT), Macedonia (MK), Japan (JP), New Zealand (NZ), Mexico (MX), Indiana (IN), and Massachusetts (MA); diamond, Haplogroup 2 from Norway (NO); triangles, Haplogroup 3 from Sweden (SE) and California (CA).