Beilschmiedia dictyoneura Kosterm.

de kok, R. P. J., 2016, A revision of Beilschmiedia (Lauraceae) of Peninsular Malaysia, Blumea 61 (2), pp. 147-164 : 151

2. Beilschmiedia dictyoneura Kosterm. — Map 1

Beilschmiedia dictyoneura Kosterm.(1965) 24, f. 4; Kochummen (1989) 119; I.M. Turner (1995) 276; Sach. Nishida (2008) 354. — Type: Kostermans 4464 (holotype BO [sheet number 0118371]; isotype K [K000768691]), Indonesia, north of Balikpapan, Sungai Wain region.

Trees 10–35 m tall, dbh 10–30 cm; bark grey, reddish or pale brown, smooth to finely fissured or pitted; inner bark reddish to dark brown, aromatic; sapwood white to yellowish. Twigs slender, 1.5–3 mm diam, round in cross-section, glabrous to sparsely hairy, more densely hairy at nodes; terminal leaf buds ovoid, 1.8–2.6 mm long, velutinous; hairs short, appressed, dark brown. Leaves (sub)opposite, blades lanceolate to elliptic, (5–)8–16 by 2.3–4.5 cm, thickly leathery; apex acute with a clear point; base cuneate; margins flat; secondary veins 6 –10 pairs, curving and joining near margin; tertiary veins laxly reticulate; upper surface glabrous, slightly glossy, midrib sunken to flattened, secondary veins raised, tertiary veins distinct; lower surface glabrous, midrib raised, secondary veins raised, tertiary veins distinct. Petiole 14–22 mm long, glabrous, channelled. Inflorescence 14–70 mm long, sparsely hairy to velutinous, not enclosed at base by bracts; bracteoles lanceolate, 1–1.8 mm long, apex acute, persistent. Flowers greenish white to white; perianth tube absent; perianth lobes broadly ovate, almost equal or inner three lobes slightly smaller, 0.5–1 by 1–1.7 mm, sparsely hairy, apices broadly acute. Stamens 6, representing first and second whorls, c. 0.7 mm long, glabrous or pubescent only around base, filaments shorter than anthers; anthers glabrous, acute at apex. Staminodes 3–6 (3 in third whorl, 0–3 in fourth whorl). Ovary c. 0.6 mm long, glabrous; style c. 0.5 mm long. Fruit (dried) ellipsoid, 12–17 by 9.5–13.3 mm, apex rounded, base rounded, surface pitted when young to warty when mature. Stalk slightly swollen, 2–2.6 mm diam, 25 mm long, not constricted at apex.

Distribution — Peninsular Malaysia, Borneo and Sumatra.

Ecology — Lowland to mountain forest, sometimes on sandy, loamy soil, at 20–1100 m altitude.

IUCN Conservation Assessment — Least Concern.

Phenology — Flowering and fruiting all year round.

Note — This is one of the few Beilschmiedia species that only has six stamens, a character that it shares with some collections of the Peninsular Malaysian species B. glauca and the two Borneo endemics B. crassa Sach.Nishida and B. kinabaluensis Kosterm. ( Nishida 2008) .

Figures

Map 1 Distribution of Beilschmiedia atra (●), B. dictyoneura (▲) and B. glabra (▼). Map based on a map provided by d-maps.com.