Galinhapodus, Santos & Moratalla & Royo-Torres & Belvedere & Castanera & Ceríaco & Mocho & Merino & Meyer & Sciscio, 2024

Santos, Vanda F., Moratalla, José J., Royo-Torres, Rafael, Belvedere, Matteo, Castanera, Diego, Ceríaco, Luis M. P., Mocho, Pedro, Merino, Marc, Meyer, Christian A. & Sciscio, Lara, 2024, A revised name and new insights into the Middle Jurassic sauropod trackways from Portugal. A correction of Santos et al. 2009, Acta Palaeontologica Polonica 69 (4), pp. 587-590 : 588

Ichnogenus Galinhapodus nom. nov.

pro Polyonyx Santos et al., 2009 nec Stimpson, 1858

Zoobank LSID: urn:lsid:zoobank.org:act:.

Type ichnospecies: Polyonyx gomesi Santos et al., 2009 .

Etymology: Combination of Galinha, to honour the name of the tracksite where this morphotype was identified. Furthermore, it is also to acknowledge the former quarry owner, Rui Galinha; and Greek podus, foot.

Diagnosis.—As for the type and only known ichnospecies; see Fig. 1 and SOM 1 (Supplementary Online Material available at http://app.pan.pl/SOM/app69-Santos_etal_SOM.pdf) and Morphosource (https://doi.org/10.17602/M2/M634167, https://doi. org/10.17602/M2/M623555, and https://doi.org/10.17602/M2/ M623561), Santos et al. 2009: figs. 3–5.

Galinhapodus gomesi ( Santos et al., 2009) comb. nov. Holotype: An in situ sauropod trackway (trackway G5), 142 m long with 94 consecutive manus-pes print sets.

Type horizon: Serra de Aire Formation, close to the Bajocian–Bathonian boundary, Middle Jurassic ( Azerêdo 1993, 2007; Azerêdo et al. 1995).

Type locality: Galinha tracksite, Municipal area of Bairro, Serra de Aire , Maciço Calcário Estremenho, west-central Portugal. This locality is the first geological site to be declared a natural monument: Monumento Natural das Pegadas de Dinossáurio da Serra de Aire ( Serra de Aire Dinosaur Tracks Natural Monument) by law in Portugal (Law Decree no. 12/96 of 22nd October) and it is under the management of the Instituto da Conservação da Natureza e das Florestas .

Diagnosis.— Wide gauge sauropod trackway revealing low heteropody (manus−pes area ratio 1:2) and two autapomor- phies: (1) asymmetric manus prints with large digit I marks oriented in a medial direction with a large, posteriorly oriented, triangular claw mark, and impressions of digits II– V; (2) pes prints with four claw marks: claws I–II with an anterior orientation, and III–IV laterally oriented. Manus digits II– V show a slightly bent arrangement ( Santos et al. 2009).

Description.— The same as described in Santos et al. (2009: 411–413).

Remarks.— Trackway G1 from the same locality was assigned to Polyonyx isp. Both the similarities and differences between the holotype trackway G5 and G1 have already been noted (see Santos et al. 2009; Castanera et al. 2016), especially in the manus impressions. G1 shows the characteristic speech-bubble-shaped manus of Galinhapodus , but they are more symmetrical, slightly longer, and no individual digits can be clearly identified with the exception of digit I, which projects posteriomedially (instead of medially). Santos et al. (2009) concluded that “it is still uncertain which features are diagnostic at the ichnospecies level”. Thus, trackway G1 is now classified as Galinhapodus isp.

Figures

Fig. 1. Manus-pes pair from the Galinhapodus igen. nov. trackway G5. Photograph (A), contour-line outline (5 mm spacing) (B), and false-colour depth map (C) of a right manus-pes set from trackway 5 (G5) at Galinha tracksite (Bairro, Serra de Aire, west-central Portugal, Bajocian–Bathonian boundary, Middle Jurassic).