Dryopsophus Fitzinger 1843: 30

Donnellan, Stephen C., Mahony, Michael J., Esquerré, Damien, Brennan, Ian G., Price, Luke C., Lemmon, Alan, Lemmon, Emily Moriarty, Günther, Rainer, Monis, Paul, Bertozzi, Terry, Keogh, J. Scott, Shea, Glenn M. & Richards, Stephen J., 2025, Phylogenomics informs a generic revision of the Australo-Papuan treefrogs (Anura: Pelodryadidae), Zoological Journal of the Linnean Society 204 : -

Dryopsophus Fitzinger 1843: 30

( Fig. 15)

Type species: ‘ Hyla citropa Péron’ (= Hyla citripoda Péron, 1807 = Hyla citropa Duméril & Bibron 1841 ), by original designation.

Content: 10 species— Dryopsophus barringtonensis * ( Copland, 1957) , Dryopsophus citropa * ( Péron, 1807) , Dryopsophus daviesae * (Mahony, Knowles, Foster & Donnellan, 2001) , Dryopsophus kroombitensis * (Hoskin, Hines, Meyer, Clarke & Cunningham, 2013) comb. nov., Dryopsophus nudidigitus * (Copland, 1963) , Dryopsophus pearsoniana * ( Copland, 1961) , Dryopsophus phyllochrous * ( Günther, 1863) , Dryopsophus piperata ( Tyler & Davies, 1985) , Dryopsophus spenceri * ( Dubois, 1984) , Dryopsophus subglandulosus * ( Tyler & Anstis, 1983) .

Diagnosis: Dryopsophus can be diagnosed from the other members of the Cyclorana – Eremnoculus Clade as follows: from Chlorohyla by the absence of the AMES vs. presence, by overall tadpole morphology Type 5 or 6 vs. 4; from Cyclorana by presence of the intercalary structure vs. absent except for C. alboguttata , small vs. large inner metatarsal tubercle, expanded vs. and unexpanded finger and toe discs, overall tadpole morphology Type 5 or 6 vs. Type 3; from Eremnoculus by medium vs. large ED/SVL, absent vs. well-developed palpebral reticulum, tadpole oral disc Type 1, 4, or 5 vs. Type 6, overall tadpole morphology Type 5 or 6 vs. Type 7; from Leptobatrachus by overall tadpole morphology Type 5 or 6 vs. Type 4; from Megatestis by medium vs. small ED/ SVL, minimally or reduced vs. fully webbed toes, developed vs. undeveloped finger discs, absence of the AMES vs. presence, a multi-divided vs. reduced m. palmaris longus, a high vs. low call dominant frequency; from Melvillihyla by a multiple note vs. single note call, by a complex call in seven of the 10 species vs. a simple call; from Mosleyia by granular vs. spinous nuptial pads, small or medium pigmented eggs vs. large unpigmented eggs, tadpole oral disc Type 1, 4, or 5 vs. Type 6, overall tadpole morphology Type 5 or 6 vs. Type 7; from Pelodryas by overall tadpole morphology Type 5 or 6 vs. Type 1 or 1A, absence vs. presence of the m. extensor brevis medius digiti IV, by a complex call in seven of the 10 species vs. a simple call; Ranoidea by the production of short higher pitched calls with fully amplitude modulated notes compared to relatively long low pitched calls of densely pulsatile notes, overall tadpole morphology Type 5 or 6 vs. Type 1, note repetition vs. densely pulsatile call, a high vs. low call dominant frequency; from Rhyaconastes by presence vs. absence of a vocal sac, absence of the AMES vs. presence, tadpole oral disc Type 1, 4, or 5 vs. Type 3; from Spicicalyx by unornamented hindlimb vs. crenulations on hindlimb, absence vs. presence of a heel spike, overall tadpole morphology Type 5 or 6 vs. Type 4, by a complex call in seven of the 10 species vs. a simple call. Refer to Tables 1 and 2.

Distribution and ecology: Forests in eastern Australia. Arboreal frogs found in lower riparian vegetation along flowing streams where they breed.

Etymology: Not stated by Fitzinger (1843), but presumably from the Greek ΔΡῦς (drys, oak tree) and ψόφος (psophos, a sound not produced by the human voice), so a sound calling from the trees. Both the original Greek noun and the Latinized version psophus are masculine (Article 30.1.3).

Remarks: Dryopsophus is the equivalent of the Litoria citropa and the Litoria maculata ( D. spenceri ) Groups of Tyler and Davies (1978).