Phaseolus augusti Harms (1921: 503)

Debouck, D. G., 2021, Phaseolus Beans (Leguminosae: Phaseoleae): A Checklist And Notes On Their Taxonomy And Ecology, Journal of the Botanical Research Institute of Texas 15 (1), pp. 73-111 : 80-81

Phaseolus augusti Harms (1921: 503) [clade B; sect. Paniculati Freytag]. TYPE: A Weberbauer 6439 ( HOLOTYPE: B n.v .; ISOTYPES: G, GH-64117 , MOL, US-1496218 ).

The type was probably lost in Berlin during WWII; while the isotype at G consists only of fragments, the other three are complete blooming specimens. If there is any need (see Turland 2013, p. 64 and following), on the basis of the number of flowering racemes, the author wishes to indicate: “ Neotype designated here: A Weberbauer 6439 (GH-64117)”. Using this specimen and the specimen US-1058964 (= the type of Phaseolus bolivianus Piper ), the author fully endorses the synonymy of the latter with P. augusti proposed by James Zarucchi and Alfonso Delgado-Salinas (1993, p. 508). Incidentally, it is quite strange that Charles Vancouver Piper (1926) did not mention P.augusti (thus creating a need for P.bolivianus ) while he mentioned P.pachyrrhizoides (with one “r,” p. 700). The original population of P.bolivianus is probably gone by now due to the expanding city of Cochabamba; an alternative could be the population found NW of Quillacollo DG Debouck 3000 (DAV-167638, MO-5613159, NY) ( Freyre et al. 1996). Along recommendation 60C.2 of the Code ( McNeill et al. 2012, p. 134), the spelling of the current termination of the epithet can be used.

Phaseolus augusti is a species well distributed in the Andes in alder seasonal forest at 1400–3200 m, related to the Andean form of wild P. lunatus ( Delgado-Salinas et al. 2006; Serrano-Serrano et al. 2010), and thus included into the Paniculati where the Lima bean is the reference species ( Fig. 10 ).

Specimens: ECUADOR. Azuay: DG Debouck 2764 (QCA). Loja: L Ellemann 91733 (K). PERU. Piura: DG Debouck 2816 (COL-332611). Cajamarca: N Angulo 1609 (F-1746428). La Libertad: S Leiva 453 (HAO-3037). Huánuco: C Ochoa 1085 (MOL). Junín: DG Debouck 2146 (UC-1571124). Apurimac: C Vargas 8858 (CUZ-7418). Huancavelica: O Tovar 1919 (US-2490509). Cuzco: E Petersen 1535 (LIL-410643). Puno: J Soukup 726 (CUZ-7417). BOLIVIA. La Paz: SG Beck 8659 (BAB). Cochabamba: DG Debouck 2483 (UC-1716208). Norcinti: DG Debouck 2504 (LPB). Chuquisaca: DG Debouck 2490 (LPB). Tarija: JRI Wood 9539 (K). ARGENTINA. Jujuy: A Burkart 11591 (SI). Salta: DG Debouck 1774 (BAA). Tucumán: R Fortunato 3565 (BAB). Catamarca: M Cerana 318 (CORD).

Figures

FIG. 10. Close-ups of pods of: a. P.augusti (DG Debouck 2571), b. P.pachyrrhizoides (DG Debouck 2155), c. P.lignosus (C Watlington 3), d. P.lunatus (DG Debouck 3236), and e. P.viridis (GJuárez 1086).