Lathyrus L.
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publication ID |
https://doi.org/10.5281/zenodo.293200 |
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persistent identifier |
https://treatment.plazi.org/id/03B0402C-FF5F-E3F4-F777-FE6CDD6DFB87 |
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treatment provided by |
Plazi |
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scientific name |
Lathyrus L. |
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51. Lathyrus L. View in CoL 1
(Inch Orobus L. )
Annual or perennial herbs, often climbing by means of tendrils. Leaves usually paripinnate and terminated by a tendril, rarely reduced to a tendrilor a grass-like phyllode; leaflets usually distinctly parallel-veined; stipules usually herbaceous. Flowers in axillary racemes, or solitary, axillary. Calyx actinomorphic to bilabiate; keel usually obtuse; stamens diadelphous; style pubes cent on the upper side, rarely glabrous. Legume usually oblong, compressed, dehiscent. Seeds 2 or more.
No clear distinction exists between this genus and Vicia . The majority of the species can readily be separated from Vicia by the often winged stem, the parallel-veined, and often fewer leaflets and the style-pubescence. On the other hand species 1-10 can be separated only by style-pubescence, and it must be doubtful whether this character alone can justify the assignment of these species and, for example Vicia orobus or V. oroboides , to separate genera.
In a number of species, particularly 12-14 and 39-45, thecolour of the corolla changes markedly on drying, becoming pale purple or blueinstead of a bright red or reddish-purple. Because of this some of the descriptions of corolla-colour may not be accurate.
Most species occur in dry grassland or scrub or as ruderals.
Literature: M. Bässler, Feddes Repert. 72: 69-97 (1966). K. Brunsberg, Bot. Not. 118: 377-402 (1965). Z. Czefranova, Nov. Syst. Pl. Vase. (Leningrad) 1965: 152-167 (1965).
1 Leaves without leaflets
2 Rhachisforminga grass-likephyllode; stipulesminute;corolla crimson 53. nissolia 2 Rhachis forming a tendril; stipules 6-30(-50) mm, ovate, hastate; corolla yellow 54. apbaca 1 At least the upper leaves with 1 or more pairs of leaflets
3 Stem winged, at least in the upper part
4 Lower leaves without leaflets,the rhachisbroadlywingedand resembling a leaf
5 Upper leaves with 1-2 pairs of leaflets; corolla yellow; dorsal suture of legume with 2 wings 52. ochrus 5 Upper leaves with 2-4 pairs of leaflets; corolla purple with violet, lilac, white or pink wings; dorsal suture of legume
not winged
6 Wings violet or lilac; standard emarginate; dorsal suture of legume channelled 50. clymenum 6 Wings white or pink; standard mucronate; dorsal suture
of legume not channelled 51. articulatus 4 All leaves with 1 or more pairs of leaflets 7 At least some leaves with 2 or more pairs of leaflets; racemes 2- to many-flowered
8 Leaveswithout a tendril; rhachis mucronate
9 Corolla cream, sometimes with a pink or purple tinge;
hilum c. I of the circumference of the seed
15. pannonicus 9 Corolla red-purple or crimson; hilum c. | of the circum
ference of the seed
10 Calyx pubescent 18. alpestris 10 Calyx glabrous, sometimes with ciliate teeth 19. montanus 8 Leaves with atendril
11 Rhachis at least 4 mm wide, broadly winged
34. heterophyllus 11 Rhachis not more than 3 mm wide
12 Stipules usually at least 25 mm, about as large as
leaflets 9. pisiformis 12 Stipules less than 25 mm, distinctly smaller than leaflets
13 Lowest tooth of the calyx about as long as tube
23. palustris
13 Lowest tooth of the calyx distinctly shorter than tube
14 Leaflets (3-)4-5 pairs; legume 30-35 x 5-5-5 mm; seeds smooth 11. incurvus 14 Leaflets 2-3 pairs; legume 40-70x 7-11 mm; seeds tuberculate 26. cirrhosus 7 Allleaves withonly1 pairofleaflets; rarelysomewith
2 pairs and then the flowers solitary
15 Racemes (3-)5- to many-flowered
16 Stipules less than I as wide as stem 32. sylvestris 16 Stipules at least | as wide as stem
17 Margin of leaflets undulate; calyx-teeth only slightly unequal, the lowest tooth not more than 1 | times as long as the upper 2 31. undulatus 17 Margin of leaflets not undulate; calyx-teeth very unequal, the lowest tooth at least twice as long as the upper 2
18 Leaflets not more than twice as long as wide, the lateral veins not extending more than about 1-way to the apex 30. rotundifolius 18 Leaflets more than twice as long as wide, with 3 or more parallel veins ± reaching the apex 33. latifolius 15 Racemes 1 - to 3(-4)-flowered
19 Corolla yellow, sometimes with a pinkish tinge
20 Corolla 7-12 mm; legume 5-7 mm wide; seeds rugulose 47. hierosolymitanus 20 Corolla 12 mm or more; legume 7-12 mm wide
21 Calyx-teeth equalling or slightly longer than tube; corolla 12-18 mm; seeds tuberculate or papillose
46. annuus 21 Calyx-teeth 2-3 times as long as tube; corolla 18-25
mm; seeds smooth 48. gorgoni 19 Corolla variously coloured, but never yellow
22 Corolla 20 mm or more
23 Peduncles not more than 60 mm; legume with 2 wings on the dorsal suture 44. sativus 23 Peduncles more than 70 mm; legume not winged
24 Pubescent, at leaston the calyx and legume 37. odoratus 24 Glabrous
25 Leaflets 1-4 mm wide, linear-lanceolate; calyxteeth longer than tube 35. tremolsianus 25 Leaflets 4-18 mm wide, lanceolate to ovate; calyxteeth shorter than tube 36. tingitanus 22 Corolla less than 20 mm
26 Calyx-teeth not or only slightly longer than tube
27 Leaflets 5-12 mm; corolla blue; seeds smooth
25. neurolobus 27 Leaflets 15-90 mm; corolla crimson with blue wings,
or orange-red; seeds tuberculate or reticulaterugose
28 Sparsely pubescent; corolla crimson with pale blue wings; legume tuberculate and densely pubescent
49. hirsutus 28 Glabrous; corolla orange-red; legume glabrescent but persistently pubescent on the suture
42. setifolius 26 Calyx-teeth l|-3 times as long as tube
29 Corolla red; legume with 2 wings on the dorsal and ventral sutures 45. amphicarpos 29 Corolla white, pink or purple
30 Legume with 2 keels on the dorsal suture; peduncles 10-30 mm 43. cicera 30 Legume with 2 wings on the dorsal suture; peduncles 30-60 mm 44. sativus 3 Stem not winged
31 Atleasttheupperleaveswitha tendril
32 Corolla 25-30 mm; legume 60-90 mm 29. grandiflorus 32 Corolla less than 25 mm; legume less than 60 mm
33 Racemes 2- to many-flowered; calyx-teeth ±distinctly unequal
34 Corolla yellow (20-22). pratensis group 34 Corolla purple or bluish
35 Leaflets 1 pair 27. tuberosus 35 Leaflets 2-5 pairs 36 Leaflets linear, oblong or lanceolate, distinctly parallel-veined, the lateral veins extending ± to the apex 23. palustris 36 Leaflets lanceolate-elliptical to orbicular-elliptical, pinnately veined
37 Stipules 10-20 mm wide, triangular-hastate; legume 6-10 mm wide 4. japonicus 37 Stipules 2-10 mm wide, linear-lanceolate to orbicular-ovate, semi-sagittate; legume 4-5-5-5 mm wide 10. humilis 33 Flowers solitary, very rarely 2 together; calyx-teeth usually equal
38 Peduncles 2-5 mm; seeds smooth, with hilum less than of the circumference 41. inconspicuus 38 Peduncles 5-70 mm; seeds with hilum more than of the circumference
39 Legume 7-11 mm wide, pubescent when immature; seeds 2-3, coarsely reticulate-rugose; peduncles not aristate 42. setifolius 39 Legume 3-7 mm wide, glabrous; seeds 8-15; peduncles usually aristate
40 Peduncles 5-20 mm; legume 4-7 mm wide, with prominent longitudinal veins; seeds smooth or slightly rugose 39. sphaericus 40 Peduncles 20-70 mm; legume 3-4 mm wide, with indistinct reticulate venation; seeds tuberculate
40. angulatus 31 All leaves without a tendril
41 Flowers solitary, with peduncles2-10 mm; calyx-teeth
± equal
42 Pubescent; leafletsofupperleaves7 -20mm; legume
5-7 mm wide, glabrous 38. saxatilis 42 Glabrous; leaflets of upper leaves 25-40 mm; legume 2-5
mm wide, densely pubescent when young 41. inconspicuus 41 Flowers in racemes, rarely solitary and then with peduncles more than 10 mm and calyx-teeth unequal
43 Corolla pale cream, yellow or orange-yellow, sometimes with a red or purple tinge
44 Leaflets pinnately veined; stipules 2-12 mm wide, lanceolate to ovate (5-8). laevigatus group 44 Leaflets distinctly parallel-veined; stipules 0-5-2(-4) mm wide, linear or linear-lanceolate, semi-sagittate, semihastate or sagittate
45 Corolla yellow; leaflets 1 pair, 8-20 mm; legume black (20-22). pratensis group 45 Corolla cream or pale yellow sometimes with a red or purple tinge; leaflets 1 -5 pairs, usually more than
20 mm; legume pale brown
46 Calyx glabrous or cibate on the margin; style filiform
15. pannonicus 46 Calyx pubescent or villous; style dilated at apex
47 Pubescent; leaflets 1 -3 pairs; legume glabrous
16. pallescens 47 Villous; leaflets 3-5 pairs; legume villous 17. pancicii 43 Corolla red-purple, purple or bluish
48 Stipules 8-20 mm wide, usually hastate or sagittate, almost as large as the leaflets
49 Leaflets 2-5 pairs; calyx-teeth less than twice as long as tube; legume 6-10 mm wide 4. japonicus 49 Leaflets 1 pair; calyx-teeth 2-3 times as long as tube; legume 3-5 mm wide 24. laxiflorus 48 Stipules 0-5-8 mm wide
50 Leaflets pinnately veined or very feebly parallelveined, the lateral veins much weaker than midrib
51 Legume covered with brown glands; racemes 6- to 30- flowered 2. venetus 51 Legume eglandular; racemes 1 - to 10-flowered
52 Leaflets acuminate; stipules 10-25 mm 1. vernus 52 Leaflets obtuse or subacute; stipules 4-10 mm
53 Leaflets 3—6(— 11) pairs; legume black 3. niger 53 Leaflets 1 pair; legume pale reddish-brown 28. roseus 50 Leaflets parallel-veined, the lateral veins reaching
± to the apex of the leaflets
54 Roots fusiform, fleshy; leaflets 1-2 pairs; calyx-teeth only slightly unequal; legume 5-9 mm wide
14. digitatus
54 Roots slender; leaflets 2-4 pairs; calyx-teeth usually very unequal; legume 4-6 mm wide
55 Leaflets 5-11 mm wide, usually less than 10 times as long as wide, obtuse or subacute, mucronate
18. alpestris
55 Leaflets 2-6 mm wide, always at least 10 times as long as wide, acuminate
56 Keel more or less winged at apex; style dilated at apex 12. flliformis
56 Keel acute, not winged at apex; style ±filiform,
not dilated at apex 13. bauhinii
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