Lonadinae Oberpr. & Töpfer, 2022

Oberprieler, Christoph, Töpfer, Alisha, Dorfner, Marco, Stock, Miriam & Vogt, Robert, 2022, An updated subtribal classification of Compositae tribe Anthemideae based on extended phylogenetic reconstructions, Willdenowia 52 (1), pp. 117-149 : 132

15. Lonadinae Oberpr. & Töpfer , subtrib. nov.

Type: Lonas Adans. ( Lonas annua ( L.) Vines & Druce).

Description — Glabrous annual herbs. Leaves alternate, pinnatisect. Capitula in a dense corymb, discoid. Involucre hemispheric to cylindric. Phyllaries in 3 or 4 rows, with narrow, pale scarious margins. Receptacle narrowly conic, paleate; paleae narrowly obovate, flat to slightly canaliculate, with a central resin canal. Florets hermaphrodite, fertile; corolla 5-lobed, yellow. Achenes narrowly obovoid, round in cross-section or slightly dorsiventrally flattened, with 1 adaxial and 2 lateral ribs; apex with a scarious corona; pericarp with myxogenic cells abaxially and on ribs and with a resin sac apically in adaxial rib. Embryo sac development monosporic. Base chromosome number x = 9.

Distribution — NW Africa, Sicily.

Members — Lonas Adans. (1).

Notes — The unispecific genus Lonas was assigned to the extremely polyphyletic subtribe Matricariinae sensu Bremer & Humphries (1993) , who acknowledged that the position of the genus “is difficult to assess” (l.c.: 155). The species L. annua was described by Linnaeus (1753) as a member of the genus Santolina and a close relationship with Santolina and other members of the Santolininae is reasonable both on morphological (receptacular paleae with a central resin duct) and molecular phylogenetic grounds. However, none of the gene and species trees presented here revealed a strongly supported position within the Santolininae ; conversely, in the concatenated sequence analysis ( Fig. 3), it is strongly supported as a sister-group to a monophyletic Santolininae + Glebionidinae lineage.