Onthophagus anthracinus, Harold, 1873

Arriaga-Jiménez, Alfonsina, Escobar-Hernández, Fernando, Rös, Matthias & Kohlmann, Bert, 2020, The establishment of the Onthophagus anthracinus (Coleoptera: Scarabaeidae) species complex and the description of a new species, The Canadian Entomologist 152 (1), pp. 1-17 : 13

The Onthophagus anthracinus species complex

We establish the O. anthracinus species complex as having the following characteristics: anterior edge of pronotal punctures lacking tubercles; pygidial punctures very shallow, at least in basal half, sometimes deep and distinct if apical half of pygidium is shiny; male with pronotal protuberance; female with protuberance usually feebly indicated; pronotal punctures not crowded, separated by at least one diameter, anterior edge of pronotal punctures lacking any small shiny tubercle; surface dull, alutaceous, brown or black; length less than 7.5 mm.