Pseudomops septentrionalis Hebard, 1917

Estrada-Álvarez, Julio Cesar & Sormani H., Carlo G., 2024, Revision of the genus Pseudomops Serville, 1831 (Blattodea: Blattellidae) in Mexico, Revista Chilena de Entomología (Rev. Chil. Entomol.) 50 (3), pp. 549-580 : 569-571

Pseudomops septentrionalis Hebard, 1917

( Figs. 4 a-4d; 8a-8J; 12a-12v; 16a-16d; 21)

Thyrsocera Sallei Saussure 1862: 168 (D?) [sp. n.] [ Mexico calida (LT)] (new synonymy). Saussure 1864: 123 (male and female) [n. rec. Les parties chaudes du Mexique...de la côte du golfe] [Trouvée par M. Sallé]. Scudder 1868: 81. Scudder 1901: 325.

Thyrsocera cincta not P. cinctus (Burmeister) . Brunner von Wattenwy 1865: 122, 412 (male and female) [part sin. Thyrsocera sallei ] [n. rec. Mexique, Oaxaca]. (var. c) Saussure 1870: 50. Saussure and Zehntner 1893: 32 [part. Var. sallei ]. Finot 1897: 186 [part synonymy sallei ].

Pseudomops Sallei Var. (Saussure) In : Walker 1868: 77 (female) [comb. n.] [n. rec. Orizaba]. Scudder 1901: 276. Kirby 1904: 74 [ Mexico].

Pseudomops oblongata [ID err.] not P. oblongatus (Linnaeus) . Rehn 1904 513 (male) [Tuxpan, Jalisco].

Pseudomops cincta [ID err.] not P. cinctus (Burmeister) . Shelford 1908 4 [part Central America, Texas]. Shelford 1908 4 [part Texas]. Princis 1969: 950 [part sinonymy sallei ]. Estrada-Álvarez 2013: 276 [part].

Pseudomops intercepta not P. cinctus (Burmeister) . Hebard 1932: 208 [part] [Jalapa, Vera Cruz; San José del Cabo, BaJa California; Cuernavaca, Morelos; Cuautla, Morelos] [ID err.].

Pseudomops septentrionalis Hebard1917 156; Lam.VI, Figs. 5-8 (D male, female) [Brownsville, Cameron County, Texas (LT); Saltillo, Coahuila, Mexico; San José, Tamaulipas, Mexico]. Hebard 1921 209 [n. rec. Saltillo, Coahuila, San José, Tamaulipas, Venvidio (sic)[=Venadillo, Sinaloa]. Hebard 1922 [1923]: 170 [Venvidio (sic)[=Venadillo]. Hebard 1932: 208 [n. rec. Matamoros, Tamaulipas]. Hebard 1942 [1943]: 268 […Northern Mexico it is foundd from the Golf of Mexico to the Pacific]. McKittrick 1964: 11, 17, 31, 61, 70, 80, 94, 95, 96, 100; Figs. 45E, 61, 62, 116, 204A, B. Princis 1969: 945 [ USA; Mexico; Panama]. Reddell and Mitchell 1971: 191 (det. A.B. Gurney) [n. rec. SIERRA DE GUATEMALA. TAMAULIPAS, Sótano de la Joya de Salas]. Reddell 1981: 206 [caves in San Luis Potosí and Tamaulipas]. Atkinson et al. 1991: 65 [ Mexico, Coahuila, Sinaloa, Tamaulipas; U.S….]. Ruiz-Cancino and Coronado-Blanco 2002: 95 [n. rec. NL]. Hoffmann et al. 2004: 309 [Sótano de la Joya de Salas, TAMAULIPAS]. Estrada-Álvarez 2013: 277 [part]. Palacios-Vargas et al. 2015: 58 [Tamaulipas]. Gaspar et al. 2015: 124; Figs. 2a, 2b. [n. rec. Arkansas, EE. UU.]. Halbert 2015: 6 [n. rec. Florida, EE. UU.]. Estrada-Álvarez et al. 2021: 748; Figs. 1 A-1J [n. rec. N.L, México] [Chihuahua, Sinaloa, Nuevo León, Tamaulipas, Zacatecas, San Luis Potosí, Queretaro, State of Mexico and Veracruz].

Note. Thyrsocera sallei Saussure, 1862 and Pseudomops septentrionalis Hebard, 1917 , they are synonyms by priority principle T. sallei would be senior synonymy of P. sepentrionalis . Based on Art. 81.2.2. ( ICZN 1999), we propose to Thyrsocera sallei as Junior synonym of P. septentrionalis , for the widespread use of P. septentrionalis before the T. sallei .

Brief diagnosis. Males, form of paraprocts ( Figs. 8c; 12g, 12h) and styles ( Figs. 8d, 8e; 12i, 12J). Females form of the paraprocts ( Figs. 8J; 1 tv) and complex valvar ( Fig. 11v); separate this species from P. interceptus .

Variations. It is the species with less chromatic variation. We found only melanic variation in this species.

Form monocolor ( Fig. 16 bc): It is a clearform, with pronotum and tegmina of the same color.

Form bicolor ( Fig. 16c): The pronotum is clearer than the tegmina.

Form melanica ( Fig. 16d): It is a dark form, with a pronotum and tegmina of the same color.

Type material. Pseudomops septentrionalis Hebard, 1917 . Holotype male from Brownsville , Cameron Co., Texas, H; July 31-aug. 5, 1912. (ANSP), photos by Heidi Hopkins revised. Male syntype Thyrsocera sallei ; Cordova, Mexique; coll. H. de Saussure [ MHNG], revised.

Examined material (n= 33; 19 males, 10 females, 4 not sexed): USA, Dallas. 1 female, Dallas, Texas [ID Saussure Pseudomops cincta [ = sallei ]] [ MHNG]. Ooteca Dallas, Texas [ID Saussure Pseudomops cincta [ = sallei ] [ MHNG]. USA, Texas: Somerville, Co., 5 mi W Glen Rose Dinosaut Valley State Parkmay 6, 1986; col. L. German [ AMNH]. MÉXICO, Nuevo León. 1 female, Monterrey, Nuevo León; 8-VIII-76; col. H. Brailovsky [ CNIN].?[sin abdomen] Chipinque, N.L.; 23-VII-77; col. Anonymous [ CNIN]. 1 male, Chipinque, N.L.; 23-VII-77; col. Anonymous [ CNIN]. 10 males, 2 female, Nuevo León, Cerro de la Silla; 27- 04-2021; col. Jorge J. Rodríguez-RoJas [ CER]. 5 males, 1 female, Nuevo León, Cerro de la Silla; 27-05-2021 [ CER]. Linares, Mex., Nuevo Leon; V-22-1952; col. M. Cazier, W. Gertsch, R. Schrammel [ AMNH]. Querétaro. 1 female, México, Queretaro, Misión de Bucareli, 21°16.83’N, 99°36.46’W, 800 msnm; 28-V-1998; col. E. Barrera [ CNIN]. Tamaulipas. 1 female, México, Tamaulipas, Gómez Farías, Alta Cima, Casa de piedra, 1448 msnm, 23°00’32’’N, 9908’09’’W, Bosque de Pino; 11-V-2007; col. E. Barrera, H. Brailovski, L Cervantes [ CNIN]. 1 female México, Tamaulipas, Gómez Farias-Rio Frio; 4-X-85; col. F. Arias, L. Cervantes, M. Garcia [ CNIN]. Sinaloa. Venvidio, Sinaloa, Mexico, VIII-2-6 1918; col. J. A. Kusche [ AMNH]. Veracruz. 1 male, Atoyac, Veracruz; May; col. H. H. Smith [ID Saussure Pseudomops cincta [ = sallei ]] [ MHNG]. 1 male, México, Veracruz, Mpio. Buena Vista, Ixtaczoquitlán, 1000 msnm; 28-VII-2000; col. E. Barrera, A. Ibarra [ CNIN].

Distribution. USA: Alabama, Arkansas, Louisiana, Mississippi, Oklahoma and Texas. MÉXICO: Baja California Sur, Mpty. Los Cabos (San José del Cabo). Chihuahua, Mptys. Chínipas, Chihuahua, Meoqui, Julimes. Coahuila, Mpty. Saltillo (Saltillo). Jalisco, Mpty. Tuxpan (Tuxpan). Nuevo León: Mpty. Guadalupe ( Guadalupe, Cascadas de Guadalupe in Cerro de la Silla); Mpty. San Pedro Garza (Chipinque), and Mptys. Allende, Aramberri, García, General Zaragoza, Hualahuises, Iturbide, Juárez, Linares, Montemorelos, Monterrey, Santiago, Zuazua. Querétaro, Mpty. Pinal de Amoles (Misión de Bucareli). Sinaloa, Mpty. Mazatlán (El Venadillo). San Luis Potosí, caves (without more data). Tamaulipas, Mpty. Gómez Farías (San José, Alta Cima, Gómez Farías-Puente Rio Frio); Mpty. Matamoros (Heroica Matamoros); Mpty. Jaumave (Sotano de la Joya de Salas en la Sierra Guatemala). Caves (without more data). Veracruz, Mpty. Atoyac (Atoyac); Mpty. Ixtaczoquitlán (Buena Vista, Moyoapan); Mpty. Veracruz (Veracruz), Zacatecas, Mpty. Tlaltenango de Sánchez Román ( Fig. 21).

Biogeography. Neartic, mostly in Tamaulipas province (tam), with reminiscences in the provinces Sierra Madre Oriental (sme), Trans-Mexican Volcanic Belt (tvb), Mexican plate (mp), Mexican Pacific coasts (mpc), Mexican Gulf (mg), and Baja California (baj); at an altitude of 6-1,299 m ( Fig. 21).

Bionomics. This species has been observed perching on a plant leaf during the day and presented activity at night.

Other examined material. Pseudomops neglectus Shelford, 1906 . 3 females, Lagos, Bresil [ Brasil]; col. Mr. Brunner d. W. [MHNG]. 2 males, 5 females Bresil; col. Dr. Thering [MHNG]. 1 female mis?[intelligible], Argentina; XII/20/19; col. P. Jorgensen [determinated by Hebard in 1926]. [MHNG]. Pseudomops annulicornis ( Burmeister, 1838) ? [sex not determinated] Sta. Para, Brazil [determinated by Hebard 1926]. [MHNG]. Pseudomops flavipes ( Burmeister, 1838) . Holotype male [ T. dubia ]. Rio, Bresil; col. Guérin [MHNG]. Pseudomops crinicornis ( Burmeister, 1838) . 6 females, Cayenne; col. M. H de Saussure [MHNG].

Calayotlus discicollis ( Burmeister, 1838) holotipo (Jürgen Deckert, MFN). 1 male, Valle de Bravo, Edo de México, Loc. 1600; 12/VIII/69; Col. Anonymus [ CNIN]. 1 male, México, Guerrero, 10 Km. Al oeste de Tololaopan; 23/VI/1990 /; col. H. Braylovsky y E. Barrera [ CNIN.] 1 male, México, Michoacán, Pedernales , Arr. Frío; 10/XI/1987; col. D. Barrera [ CNIN]. 3 males, México, Morelos, Loc. 2, 2.5 Km. N 4 km. O Huautla, Estación CEAMISH, 940 m de Alt., Colecta diurna; 06-10/VIII/1996; col. S. Zaragoza [ CNIN]. 1male, 3 females. México, Estado de México, Amanalco, El Salto; 19/V/2009; col. J. C. Estrada-Álvarez y Guadarrama C. Cols. , en casa habitación. 1 male, Estado de México, Malinalco, Malinalco ; 15/III/2007; col. Julio C. Estrada-Álvarez.