Elongeuma speophilum Golovatch, 1982

Golovatch, S. I. & Spelda, J., 2022, Review of the Central Asian millipede genus Elongeuma Golovatch, 1982, with descriptions of two new species (Diplopoda: Chordeumatida: Kirkayakidae), Arthropoda Selecta 31 (1), pp. 1-9 : 6-7

Elongeuma speophilum Golovatch, 1982

Figs 24–31 , Map.

Elongeuma speophilum Golovatch, 1982: 944 (D).

Elongeuma speophilum — Read, Golovatch, 1994: 64 (M); Turbanov et al., 2016: 1291 (M).

HOLOTYPE ♂ ( ZMUM ρ1530), revised, Tien-Shang (= Tian Shan) Mts, Kyrgyzstan (= Kirghizia), Fergana Mt. Range , Kirgyzindan River valley , Kendyn-kyry (= Kyndyn-kyry, or Zindan) Cave, 100–120 m inside (off entrance), X.1978, V. Ogudin leg. (https:// speleo.kg/expeditions/2021/alternativnyj-karstovyj-rajon-v-dolinereki-kuraves/).

DIAGNOSIS. Differs from congeners by a pallid coloration with only five light brown ocelli on each side of the head. ♂ legs 7 are with a pair of strong hooks on the coxae and small glandular parabasal knobs on the prefemora ( Fig. 26 ). Coxal pouches (cs) of the posterior gonopods are membranous (see also key below).

BRIEF REDESCRIPTON. Length ca 14 mm, width 1.1 mm on mandibular genae, 1.0 mm on midbody segments (♂).

All characters as in E. reductum sp.n., except as follows.

Coloration pallid, only 5 ocelli on each side of head light brown. Body with 28 segments. Antennae longer, reaching segment 6 when stretched laterally, antennomere 3 being the longest. Metatergal macrochaetae mostly bacilliform and blunt ( Fig. 24 ). ♂ leg-pairs 3–6 only slightly enlarged, with tarsal papillae distally; each ♂ leg 7 ( Fig. 26 ) with a laterally micropapillate coxa carrying a strong ventral hook directed distad, while prefemur with a small, glandular, parabasal knob ventrally; ♂ legs 10 and 11 with coxal sacs/glands ( Figs 27, 28 ), usual, devoid of tarsal papillae like following legs.

Anterior gonopods ( Figs 29, 30 ) much like in E. reductum sp.n., but differing in the absence of a lobule (j) on each solenomere (sl), thus being very similar to those of E. chichkan sp.n.. Posterior gonopods ( Fig. 31 ) also very similar to those in E. chichkan sp.n., both coxites being quite high, sac-shaped and densely setose, while coxal pouches or sacs (cs) with membranous projections.

REMARK. This species was observed living on ice inside the cave at an elevation of> 2000 m a.s.l. (https:// speleo.kg/expeditions/2021/alternativnyj-karstovyj-rajon-vdoline-reki-kuraves/).

Figures

Fig. 24–31. Elongeuma speophilum Golovatch, 1982, # holotype: 24 — left half of midbody ring, dorsal view; 25 — gnathochilarium, ventral view; 26 — legs 7, posterior view; 27 — leg 10, posterior view; 28 — coxa 11, posterior view; 29, 30 — anterior gonopods, lateral and posterior views, respectively; 31 — posterior gonopods, anterior view. Abbreviations: cs — coxal sac vestige; hl — hyaline lobe; pfl — pseudoflagellum; sg — seminal groove; sl — solenomere; st — sternum. Scale bars: 0.5 mm (24, 29–31) and 0.2 mm (25–28). After Golovatch [1982]. Рис. 24–31. Elongeuma speophilum Golovatch, 1982, голотип #: 24 — леваЯ половина среднетуловиЩного сегмента, сверху; 25 — гнатохилЯрий, сниЗу; 26 — ноги 7, сЗади; 27 — нога 10, сЗади; 28 — таЗик 11, сЗади; 29, 30 — передние гоноподы, соответственно сбоку и сЗади; 31 — Задние гоноподы, спереди. ОбоЗначениЯ: cs — остаток коксального кармана; hl — гиалиноваЯ пластина; pfl — ложный жгутик; sg — семенной канал; sl — соленомер; st — стернит. МасШтаб: 0,5 мм (24, 29–31) и 0,2 мм (25–28). По: Головач [1982].