Epithemateae

M., Taram, D., Borah, H., Tag, M., MÖller & Weber, A., 2022, Whytockia (Gesneriaceae), a new generic record for India, based on a new species, W. arunachalensis, from Arunachal Pradesh, India, Rheedea 32 (1), pp. 18-29 : 19

Key to the genera of Tribe Epithemateae in India

1. Leaves mostly opposite, those of a pair ± of same shape and size, symmetrical or slightly asymmetrical at base, first cauline leaf solitary and distinctly larger than the paired leaves; inflorescences in terminal and axillary positions, consisting of a stalk, a single denseflowered head (much condensed pair-flowered, ebracteolate cyme, flowers in four rows), and a small or conspicuous cucullate subtending bract; fruit a pyxis (capsule with circumscissile dehiscence, bowl-shaped when open) ................................................. Epithema

1. Leaves alternate, or if opposite then those of a pair strongly unequal (anisophyllous), always strongly asymmetrical at base; inflorescences either unilateral racemes in terminal position or pair-flowered cymes in axillary positions; fruit capsular, opening with two or four valves, or breaking irregularly in pieces .................. 2

2. Leaves (in the Indian species) always alternate (by complete reduction of smaller leaf); inflorescence a terminal, unilateral raceme, flowers several to many, placed in two rows (superficially similar to a scorpioid cyme), all or some associated with a small subtending bract or bracts lacking; corolla strongly bilabiate, with enlarged, undivided plate-like or 3-parted lower lip, and upper lip reduced to a small erect or reflexed bilobed rim, mouth closed (personate) .................. Rhynchoglossum

2. Leaves in pairs, those of a pair strongly unequal in size; inflorescences cymose, either pair-flowered or conventional cymes, ebracteolate; corolla almost actinomorphic or with bilabiate limb, lower lip not much enlarged, mouth open ............................................................... 3

3. Small leaves ± of same shape as the large leaves; inflorescences pair-flowered cymes, emerging only from the axils of the large leaves, ebracteolate; calyx imbricate (margins overlapping); corolla zygomorphic, bilabiate, short-tubed-infundibuliform; ovary bilocular, with axile placentae (four ovule-bearing arms arising from the middle of the complete septum) ........................................... Whytockia

3. Small leaves reduced to scale- or stipule-like appendages; inflorescence a terminal thyrse, in vigorous plants comprising 2–3 pair-flowered or conventional cymes; calyx valvate-plicate; corolla either actinomorphic and ecalcarate, or bilabiate and with distinct spur at base; ovary unilocular, placentae parietal ...... Stauranthera