Dehaasia incrassata (Jack) Kosterm., 1952

Fijridiyanto, I. A., Smets, E. & Arifiani, D., 2020, Taxonomic revision of Dehaasia (Lauraceae) in Sumatra, Blumea 65 (2), pp. 167-175 : 171

4. Dehaasia incrassata (Jack) Kosterm. — Map 1

Dehaasia incrassata (Jack) Kosterm.(1952a) 91; Merr.(1952) 230;Backer & Bakh.f. (1963) 131; Kosterm. (1964) 468;(1973) 441; Kochummen (1989) 139. — Laurus incrassata Jack (1822) 33;Kosterm.(1964) 637. — Machilus incrassatus (Jack) Nees (1831) 70; Kosterm.(1964) 910. — Haasia incrassata (Jack) Nees (1836) 376, nom. inval., genus name rejected. — Persea incrassata (Jack) Nees (1836) 127, 376; Kosterm. (1964) 1230. — Type: Jack s.n. † (see note), [ Indonesia], Sumatra,Natal; neotype (selected here): Teysmann s.n. (neo L.0036318), [ Indonesia], Sumatra.

Haasia microcarpa Blume in Nees (1836) 373. — Dehaasia microcarpa (Blume) Blume (1837) 162; Kosterm. (1964) 469. — Lectotype (selected here): Hasskarl s.n. (lecto L.0036319; isolecto U.0002728), [ Indonesia,] Java, Bantam [Banten].

Dehaasia squarrosa Zoll. & Moritzi in Zoll. (1854) 115; Hassk. (1855) 8, as nom. nov. — Haasia squarrosa (Zoll. & Moritzi) Miq. (1858) 929; Kosterm. (1964) 537. — Type: Zollinger 1376 (holo BO not found; iso P.01753085*, U0002732 ), [ Indonesia,] Java.

Dehaasia media Blume (1837) 163; Kosterm. (1964) 469. — Lectotype (selected here): sine coll. (lecto L.0036322; isolecto L.0036323, L.0036324), [ Indonesia], Ambonia (Ambon).

Beilschmiedia purpurea Elmer (1910) 703;Kosterm.(1964) 145. — Syntypes: Elmer 11288 A.00041494*,BM.000950898*,E.00386537*,HBG.509735*, G.00368769*, L.0036325, LE.00012682*, MO247193 *, NY.00022716*, US.00099411*), [ Philippines,] Island of Mindanao, District of Davao, Todaya , Mt Apo .

Trees, up to 20 m tall, stem up to 40 cm diam; bark smooth; twigs sympodial, slightly striate, slender to thick, 3–7 mm diam, glabrous; terminal leaf buds glabrous to glabrescent, cataphylls ciliate. Leaves spirally arranged at the end of twigs; lamina chartaceous to subcoriaceous, elliptic to broadly elliptic or subobovate, (6.5–)12–26(–32) by (2.5–)3.5–12(–15) cm, glabrous on both sides, apex acute to acuminate, base cuneate, midrib flat to slightly raised above, midrib raised below, secondary veins (6–)8–10(–13) pairs, flat to raised above, raised and joining near margin below, tertiary veins scalariform-reticulate, conspicuous above, prominent below. Petioles 7–25 mm long, 1–4 mm diam, glabrous, flat to channelled above. Inflorescences terminal and axillary, paniculate-cymose, (4–)10–13(–18) cm long; rachis pale green to red when fresh, filiform, with minute light brown appressed hairs. Flowers greenish white; tepals unequal, ovate, outer tepals 1.1–1.3 by 1.5–2.5 mm, glabrous or with sparse minute light brown appressed hairs, margin ciliate; inner tepals broadly ovate, 2.5–2.7 by 2–2.7 mm, apex broadly acute, glabrous or with sparse minute light brown appressed hairs, margin ciliate; receptacle shallow; pedicel c. 6.7 mm long. Stamens 1.5–2 mm long, anthers rounded to truncate or slightly emarginate at apex; filaments longer than anthers, pubescent, filaments of third whorl with two sessile glands each. Staminodes absent. Pistil c. 2.5 mm long; ovary subgloboid to ovoid, c. 1 by 1.3 mm, glabrous; style terete, glabrous, slender, longer than ovary, c. 1.5 by 0.1 mm; stigma triangular. Infructescences 2–8.5 cm long, glabrous. Fruit ellipsoid, 2–3.5 by 1–2.5 cm, shiny black when fresh; stalk distinctly swollen, 2.5–3 cm long, fleshy and warty, bright red when fresh.

Distribution — Thailand, Malay Peninsula, Sumatra (Aceh, North Sumatra, South Sumatra, West Sumatra, Bengkulu, Lampung, Simalur (Simeulue) Island, Sipora Island, Siberut Island and Enggano Island), Java, Borneo, Sulawesi, Maluku and New Guinea.

Habitat & Ecology — Primary and secondary forests, at 50–1200 m altitude. Flowering: February to December; fruiting: February to December.

Vernacular names — Kayu madang kuning, kayu si marhalu- wang, masasen kuning, medang kayu mesang, medang keladi, medang kuning.

Note — The type specimen of Jack’s collection from Sumatra was destroyed in 1824 ( Merrill 1952, Balakrishnan & Chakrabarty 2011). Therefore a neotype is designated here using Teijsmann’s collection, also collected from Sumatra. This specimen is a paratype of Dehaasia squarrosa , a synonym of D. incrasssata .