Neodubrovnikella turonica ( Said & Kenawy, 1957 )

Simmons, Michael, Bidgood, Michael, Consorti, Lorenzo & Schlagintweit, Felix, 2025, A Review Of The Identity And Biostratigraphy Of Cenomanian “ Larger ” Benthic Foraminifera: Part 2 - The Order Loftusiida (Excluding The Suborder Orbitolinina), Acta Palaeontologica Romaniae 21 (1), pp. 103-192 : 134-136

Neodubrovnikella turonica ( Said & Kenawy, 1957)

Figure 12

T 1957 Peneroplis turonicus n. sp. Said & Kenawy, p. 82, pl. 13, figs. 14a–c, 16a–c; Turonian (now considered Cenomanian fide Schlagintweit & Yazdi-Moghadam, 2022 ), Egypt.

Non 1973 Peneroplis turonicus – Kerdany et al., pl. 1, fig. 16-18; Cenomanian, Egypt [= Biconcava bentori Hamaoui fide Hamaoui in Schroeder & Neumann, 1985].

1974 Peneroplis cf. turonicus – Saint-Marc, p. 235, pl. 8, figs. 5–9; middle-late Cenomanian, Lebanon (Turonian age discounted fide Schlagintweit & Yazdi-Moghadam, 2022 ).

1974 Foraminifer F-50 – Radoičić, pl. 6, figs. 5-6; early Turonian (now considered Cenomanian fide Schlagintweit & Yazdi-Moghadam, 2022 ), Serbia.

1976 Peneroplis cf. turonicus – Charvet et al., pl. 7, figs. 2-3; late early – late Cenomanian, Greece.

1981 Peneroplis cf. turonicus – Saint-Marc, pl. 4, figs. 11–12; middle-late Cenomanian, Lebanon (Turonian age discounted).

1989 Merlingia cf. cretacea (sic) Hamaoui & Saint-Marc – Kuss & Malchus, text-figs. 48-49; Cenomanian, Egypt.

1990 Pseudolituonella reicheli Marie – Šribar & Pleničar , pl. 2, fig. 8; middle – late Cenomanian, Slovenia.

1994 Peneroplis cf. turonicus -Velić & Vlahović, pl. 5, figs. 1-4; middle-late Cenomanian, Croatia.

2004 Peneroplis cf. turonicus – Chiocchini et al., pl. 6, fig. 6; late Cenomanian, southern Italy.

2006 Peneroplis cf. turonicus – Ettachfini, pl. 15, figs. 1- 2; late Cenomanian, Morocco.

2006 Pseudolituonella reicheli – Taslı et al., Fig. 6 D ; middle – late Cenomanian, Turkish Taurides.

2012? Peneroplis cf. turonicus – Chiocchini et al., pl. 109, fig. 1 (pars), 2–8; late Cenomanian, southern Italy.

2013 Pseudolituonella reicheli – Al-Dulaimi et al., fig. 9/8; late Cenomanian, Southern Iraq.

? 2013 Peneroplis parvus De Castro – Ghanem & Kuss , fig. 14(10); late Cenomanian, Syria.

Non 2013 Peneroblis cf. turonicus (sic) – Shahin & Elbaz, p. 278, pl. 3, fig. 3; Cenomanian, Egypt [= Peneroplis parvus ].

2016 Peneroplis turonicus – Assadi et al., fig. 6 a5; middle – late Cenomanian, Iranian Zagros.

2018 Peneroplis parvus – Omidi et al., pl. 4, fig. 8; middle – late Cenomanian, Iranian Zagros.

2020 Peneroplis turonicus – Solak et al.,?fig. 6M, fig. 10S–T,?fig. 13I; late Cenomanian, Western Taurides, Türkiye.

2021 Pseudolituonella reicheli – Dousti-Mohajer et al., pl. 1i; middle – late Cenomanian, Iranian Zagros.

2021 Peneroplis planatus (Fichtel and Moll) – Brčić et al., fig. 9i; late Cenomanian, Croatia.

2021 “ Peneroplis ” turonicus – Consorti & Schlagintweit, fig. 2H; late Cenomanian, Albania-Kosova.

? 2021 Peneroplis sp. – Solak, pl. 2E; middle – late Cenomanian, Turkish Taurides.

? 2021 Peneroplis turonicus – Dousti-Mohajer et al., pl. 3a; middle – late Cenomanian, Iranian Zagros.

2022 Neodubrovnikella turonica – Schlagintweit & Yazdi-Moghadam, p. 4, figs. 3A; 4A-F, 5-6; upper early – late Cenomanian, southern Neotethys.

Non 2022 Peneroplis turonicus – Esfandyari et al., fig. 25j; Cenomanian – early Turonian, Iranian Zagros [= Pseudolituonella reicheli ].

2023 Neodubrovnikella turonica – Simmons & Bidgood, p. 70, fig. 19; (upper early?) middle-late Cenomanian, global review.

2023 Neodubrovnikella turonica – Xu et al., fig. 3c, Cenomanian, Iranian Zagros.

Non 2023 Peneroplis turonicus – Moghaddam et al., fig. 5v; Cenomanian – early Turonian, Iranian Zagros [= Pseudolituonella reicheli ].

2024 Peneroplis turonicus – Moghaddam et al., fig. 2l; Cenomanian, Iranian Zagros.

2024 Neodubrovnikella turonica – Křížová et al., fig. 4rs; late Cenomanian, north-east Italy.

Reference Images: Schlagintweit & YazdiMoghadam (2022) p. 4, figs. 3A; 4A-F, 5-6. Taxonomy/Identity: The taxonomic status of this species and its identity has recently been reviewed by Schlagintweit & Yazdi-Moghadam (2022) who recognised that this is in fact an agglutinated taxon with a pseudokeriothecal wall, not a species of the porcellaneous genus Peneroplis (indeed also not Turonian). It was assigned to the genus Neodubrovkinella , the only other species of which, Neodubrovnikella maastrichtiana Schlagintweit & Rashidi , is Maastrichtian ( Schlagintweit & Rashidi, 2018; Schlagintweit & Yazdi-Moghadam, 2022). See also Simmons & Bidgood (2023).

It is similar to Peneroplis parvus De Castro and appears to be an agglutinated isomorph of this miliolid genus (see also Peneroplis cairensis Chiocchini, 2008 ). N. turonica is smaller (c. 0.25-0.8 mm) and with fewer chambers in the last whorl (7-8) than P. parvus and P. cairensis (Chiocchini, 2008) . Peneroplis is also always planispiral whereas N. turonica is frequently not perfectly planispiral.

It is also often confused in the literature with Pseudolituonella reicheli Marie which has a muchreduced coiled part, a cylindrical rather than flattened uncoiled portion, and apertures that are cribrate, rather than in a row as in N. turonica .

Confident Stratigraphic Range: middle-late Cenomanian.

Uncertain Stratigraphic Range: upper early Cenomanian.

Originally regarded as being restricted to the Turonian (hence the name), a re-assessment of the age of the type specimens from Egypt by Schlagintweit & Yazdi-Moghadam (2022), together with a review of other ostensibly Turonian records, indicates that this species is in fact restricted to the (middle-late) Cenomanian (see also Velić, 2007). Records in the upper part of the early Cenomanian are based on illustrated but uncertainly agecalibrated records (Charvet et al., 1976), or unillustrated records ( Decrouez, 1975), from Greece (and see also Schlagintweit & Yazdi-Moghadam, 2022, for records from the Iranian Zagros).

Geographic Distribution: Throughout Neotethys, from Morocco to Iran (see Schlagintweit & Yazdi-Moghadam, 2022).

Figures

Fig. 12. Representative illustrations of Neodubrovnikella turonica: a. Equatorial section, Schlagintweit & Yazdi- Moghadam (2022, fig. 4(E), Iran); b. Re-illustration of holotype, Schlagintweit & Yazdi-Moghadam (2022, fig. 4 (A), Egypt); c. Equatorial section, Schlagintweit & Yazdi-Moghadam (2022, fig. 6(B), Croatia); d. Axial section, Schlagintweit & Yazdi-Moghadam (2022, fig. 6(C), Kosovo); e. Apertural face, Said & Kenawy (1957, holotype, pl. 13, fig. 14c, Egypt) (Reproduced after Simmons & Bidgood, 2023).

Fig. 6. Representative illustrations of Reissella ramonensis: a. Exterior view, (Hamaoui 1963, pl. 1, fig. 3, holotype, Israel); b. Exterior view (schematic), (Hamaoui 1963, pl. 1, fig. 11, Israel); c. Enlarged cutaway exterior view showing internal features (Hamaoui 1963, text fig. 1, Israel) (Reproduced after Simmons & Bidgood, 2023).