HAPLOPHRAGMIOIDEA Eimer & Fickert 1899

Simmons, Michael, Bidgood, Michael, Consorti, Lorenzo & Schlagintweit, Felix, 2025, A Review Of The Identity And Biostratigraphy Of Cenomanian “ Larger ” Benthic Foraminifera: Part 2 - The Order Loftusiida (Excluding The Suborder Orbitolinina), Acta Palaeontologica Romaniae 21 (1), pp. 103-192 : 107

Superfamily HAPLOPHRAGMIOIDEA Eimer & Fickert 1899 (diagnosis sensu Kaminski, 2014)

Family CYCLAMMINIDAE Marie 1941 (diagnosis sensu Loeblich & Tappan, 1987)

Subfamily BUCCICRENATINAE Loeblich & Tappan 1985 (diagnosis sensu Loeblich & Tappan, 1987)

Genus Buccicrenata Loeblich & Tappan 1949 (see Table 1 for diagnosis)

Buccicrenata is an involute planispiral form with rapidly enlarging chambers that also frequently uncoils rapidly. Debate about the nature of the wall and septa of Buccicrenata - whether they are, or are not, alveolar - is not yet fully resolved, although the alveolar nature of the wall seems reasonably well established (see Loeblich & Tappan, 1985; Simmons & Bidgood, 2023). The alveolar nature of the septa in Buccicrenata (if and when unequivocally proven) separates it from Everticyclammina Redmond. Buccicrenata is also very similar to Pseudocyclammina Yabe & Hanzawa , although the former genus has a single, sinuous slit-like aperture compared with cribrate in the latter and typically somewhat thicker walls. Nevertheless, the two apertural types are hard to discern when observed in oblique/random thin-sections. In addition, and marking another distinction from Pseudocyclammina ), “… the septal base thickened against the previous whorl, and may form a continuous imperforate basal layer with solid triangular chomata-like mounds at the position of the septa ” (Loeblich & Tappan, 1985, p. 98). The two genera can also be distinguished by predominantly reniform chambers and a distinctive lobate periphery in Buccicrenata (see Simmons & Bidgood, 2023 and species key chart therein). Nonetheless, in much illustrated material in the literature it is often hard to judge the nature of aperture and the alveolar nature of the wall and septa, thus making it difficult to separate possible Buccicrenata from Everticyclammina , Pseudocyclammina , Ammobaculites and Lituola species, to name but a few.

Buccicrenata ex gr. subgoodlandensis ( Vanderpool,