Orania agapus ( De Gregorio, 1885 )

Harzhauser, Mathias, Landau, Bernard M. & Merle, Didier, 2025, The Muricidae (Gastropoda, Muricoidea) of the Miocene Paratethys Sea (Aspellinae, Ergalataxinae, Coralliophilinae, Rapaninae), Zootaxa 5611 (1), pp. 1-106 : 57-59

Orania agapus ( De Gregorio, 1885)

Figs 33 , 35A–H

Fusus variabilis Jan. —Pusch 1837: 141, pl. 12, fig. 4 [non Fusus variabilis Lamarck, 1803 = Siphonaliopsis variabilis (Lamarck,

1803)]. [ Murex ] Barrandei Hörnes — Hörnes 1848: 18 [nomen nudum]. Raphitoma cheilotoma Partsch — Hörnes 1848: 21 [nomen nudum]. Fus [us]. variabilis Jan. — Eichwald 1853: 179. Murex intercisus Micht. —Hörnes 1853: 241, pl. 25, fig. 2 [non Murex intercisus ( Michelotti, 1847) ]. Murex flexicauda Bronn —Hörnes 1853: 241, pl. 25, figs. 8a–c [non Murex flexicauda Bronn, 1831 = Orania turrita (Borson,

1821)]. * [ Murex (Aplus) plicatus (L.) Brocc.] agapus De Greg. — De Gregorio 1885: 284 [nov. nom. pro Murex flexicauda sensu Hörnes

1853: pl. 25, fig. 8]. Pollia cheilotoma Partsch sp.— Hoernes & Auinger 1890: 234: pl. 30, figs. 1–2. Pollia Barrandei M. Hoern. sp.— Hoernes & Auinger 1890: 235, pl. 30, figs. 3a–c. Pollia chilotoma [sic] (Ptsch.)— Boettger 1902: 33. Pollia barrandei (M. Hö.) — Boettger 1902: 33. Pollia chilotoma [sic] (Ptsch.)— Boettger 1906: 30. Pollia barrandei (M. Hö.) — Boettger 1906: 30. Pollia cheilotoma Partsch — Friedberg 1912: 182, pl. 11, figs. 20–21. Tritonidea cheilotoma Partsch — Friedberg 1938: 119. C [antharus]. (P [ollia]). cheilotomus (Partsch) — Sieber 1958: 151. C [antharus]. (P [ollia]). barrandei Hörn. — Sieber 1958: 151. Cantharus (Pollia) cheilotoma (Partsch in Hoernes und Auinger 1890)—Kojumdgieva in Kojumdgieva & Strachimirov 1960:

173, pl. 43, figs. 18a–b. Coralliophila (Orania) barrandei Hörnes , sp. 1848— Glibert 1963: 39. Cantharus (Pollia) cheilotomus Partsch (in Hörnes), (1848) 1856— Strausz 1966: 308, pl. 35, figs. 2–3. Cantharus cheilotomus (Partsch in R. Hoernes und Auinger, 1879)— Zelinskaya et al. 1968: 191, pl. 45, figs. 18–19. [? Cantharus ] barrandei Hörnes, 1848 — Vokes 1971: 22. Cantharus (Pollia) cheilotomus Partsch — Csepreghy-Meznerics 1972: 28, pl. 11, figs. 14–16. Cantharus (Pollia) cheilotomus (Hörnes) —A.W. Janssen 1984: 231, pl. 61, figs. 7–8. Cantharus (Pollia) cheilotomus (Partsch in Hoernes, 1848)— Atanacković 1985: 149, pl. 33, figs. 7–10. Coralliophila (Orania) cheilotoma (Hörnes & Auinger) — Bałuk 1995: 232, pl. 37, figs. 5–6. Orania cheilotoma (Hörnes & Auinger, 1890) — Harzhauser 2002: 98, pl. 6, fig. 15. Orania cheilotoma (Hoernes & Auinger) — Landau et al. 2013: 156, pl. 23, figs. 5–6. Orania cheilotoma ( Hoernes et Auinger, 1890) — Kovács et al. 2018: 126, figs. 8K–N. Orania cheilotoma ( Hoernes et Auinger, 1890) — Kovács 2019: 133. Orania fusulus ( Brocchi, 1814) — Kovács 2019: 133, figs. 65–66 [non Orania fusulus ( Brocchi, 1814) ].

Type material. Holotype, NHMW 1949/0005/0017, SL: 15.2 mm, MD: 7.3 mm, Steinebrunn ( Austria), illustrated in Hörnes (1853: pl. 25, fig. 8) and Hoernes & Auinger (1890: pl. 30, fig. 1), figs. 35A 1 –A 2.

Illustrated material. NHMW 1949/0005/0018, SL: 14.8 mm, MD: 8.6 mm, Niederleis ( Austria), illustrated in Hoernes & Auinger (1890: pl. 30, fig. 2), figs. 35B 1 –B 2. NHMW 1857/0014/0016, SL: 16.3 mm, MD: 8.4 mm, Steinebrunn ( Austria), figs. 35C 1 –C 2. NHMW 1871/0011/0017, SL: 16.0 mm, MD: 7.7 mm, Steinebrunn ( Austria), figs.35D 1 –D 2. Lectotype of Pollia barrandei Hoernes &Auinger, 1890 (designated herein): NHMW 1846/0037/0227, SL: 15.4 mm, MD: 8.0 mm, Steinebrunn ( Austria), illustrated in Hörnes (1853: pl. 25, fig. 2) and Hoernes & Auinger (1890: pl. 30, fig. 3), figs. 35E 1 –E 2. Paralectotypes of Pollia barrandei Hoernes & Auinger, 1890 : NHMW 1868/0001/0443, SL: 14.8, MD: 7.1 mm, Lăpugiu de Sus ( Romania), figs. 35F 1 –F 2. NHMW 1859/0037/0025, SL: 15.2, MD: 7.5 mm, Lăpugiu de Sus ( Romania), figs. 35G 1 –G 2. NHMW 1858/0043/0049, SL: 16.7, MD: 7.1 mm, Lăpugiu de Sus ( Romania), Fig. 35H .

Additional material. 17 spec., 1855/0043/0036, Lăpugiu de Sus ( Romania) ; 92 spec. NHMW 1846/0037/0348, Steinebrunn ( Austria) ; 93 spec., NHMW 1871/0010/0343, Steinebrunn, Gainfarn, Bad Vöslau, Niederleis and Vienna /Pötzleinsdorf ( Austria) .

Revised description. Small, slender to stout fusiform shell; apical angle 52–66°. Protoconch broad conical of three convex whorls. Teleoconch of up to five whorls. Suture deeply incised, shallowly undulating. Early teleoconch whorls with broad, weakly concave subsutural ramp, shoulder placed slightly below mid-whorl, weakly convex below. Axial sculpture of eight to ten broad ribs, fading over subsutural ramp. Spiral sculpture of broad, close-set P1 and P2 and weaker SP; s1 intercalated on second whorl, rapidly becoming subequal in strength. Cords swollen over axial ribs. P3, s2 and s3 appearing on last spire whorls. Last whorl attaining 61–65% of total height, with moderately steep subsutural ramp displaying SP, adis and IP of equal strength), weak shoulder angulation, convex periphery, moderately constricted at base. Axial sculpture of up to nine broad, low axial ribs, prominent along periphery, fading over subsutural ramp and base. P1–P6 of subequal strength, forming low, broad cords with slightly weaker s1–s6; P1 weakly tuberculate; P5 marking transition to base; P6 on siphonal canal. Crowded growth lines give surface finely scabrous appearance, especially over subsutural ramp. Fasciole weakly swollen delimiting narrow pseudoumbilicus.Aperture moderately wide, ovate. Outer lip thickened with slightly elongate ID, D1–D5; D2 bifid. Anal canal well defined, accentuated by weak anal denticle. Siphonal canal short, moderately wide, open, slightly bent to the left, dorsally recurved, shallowly notched. Columellar callus forming broad, adherent rim. Columella moderately excavated in adapical part, weakly twisted, with faint columellar fold at transition to siphonal canal.

Paratethyan synonyms. Pollia cheilotoma Hoernes & Auinger, 1890 . The name Murex (Aplus) plicatus agapus De Gregorio, 1885 was overlooked by Hoernes & Auinger (1890) when establishing Pollia cheilotoma . By selecting specimen NHMW 1949/0005/0017 from Steinebrunn ( Austria), illustrated in Hoernes & Auinger (1890: pl. 30, fig. 1) as lectotype, Pollia cheilotoma becomes an objective synonym of Murex (Aplus) plicatus agapus De Gregorio, 1885 . Pollia barrandei Hoernes & Auinger, 1890 differs from Orania agapus in its more prominent spiral sculpture. Both morphologies co-occur at many localities and intermediate specimens are present. Therefore, we consider them to be conspecific and treat Pollia barrandei as subjective junior synonym of Orania agapus ( De Gregorio, 1885) as already supposed by Landau et al. (2007: 51) and Landau et al. (2013: 156).

Discussion. See Landau et al. (2013: 156).

Paleoenvironment. Shallow marine, inner neritic with sea grass (own data, M.H.).

Distribution in Central Paratethys. Karpatian (Early Miocene): Korneuburg Basin: Stetten ( Austria) ( Harzhauser 2002). Badenian (Middle Miocene): Polish-Ukrainian Fore-Carpathian Basin: Jasinów, Zborów (Zboriv), Żukowce (Zhukivtsi) ( Ukraine) ( Friedberg 1912); Voronyaky Hills: Hołubica (Holubytsia), Podhorce (Pidhirtsi) ( Ukraine) ( Friedberg 1912); Korytnica Basin: Korytnica ( Bałuk 1995); North Alpine-Carpathian Foreland Basin: Grund ( Austria) ( Hoernes & Auinger 1890); Vienna Basin: Bad Vöslau, Gainfarn, Niederleis, Pfaffstätten, Steinebrunn, Vienna /Pötzleinsdorf ( Austria), Mikulov ( Czech Republic) ( Hoernes & Auinger 1890); Pannonian Basin: Letkés ( Hungary) ( Kovács et al. 2018); Southern Pannonian Basin: Hrvaćani ( Bosnia and Herzegovina) ( Atanacković 1985); Bükk Mountains: Borsodbóta ( Hungary) ( Csepreghy-Meznerics 1972); Bakony Mountains: Bánd ( Hungary) ( Kovács 2020); Făget Basin: CoŞteiu de Sus, Lăpugiu de Sus ( Romania) ( Kovács 2019); Dacian Basin: Urovene ( Bulgaria) (Kojumdgieva in Kojumdgieva & Strachimirov 1960).

Figures

FIGURE 33. Cord diagrams and apertural features of Microdrupa?, Orania, Lindapterys and Daphnellopsis.

FIGURE 35. Orania agapus (De Gregorio, 1885). A1–A2. holotype, NHMW 1949/0005/0017, Steinebrunn (Austria). B1– B2. NHMW 1949/0005/0018, Niederleis (Austria). C1–C2. NHMW 1857/0014/0016, Steinebrunn (Austria). D1–D2. NHMW 1871/0011/0017,Steinebrunn(Austria).E1–E2. NHMW1846/0037/0227,Steinebrunn(Austria).F1–F2.NHMW1868/0001/0443, Lăpugiu de Sus (Romania). G1–G2. NHMW 1859/0037/0025, Lăpugiu de Sus (Romania). H. NHMW 1858/0043/0049, Lăpugiu de Sus (Romania).