Pazinotus hamulifer ( Boettger, 1906 )

Harzhauser, Mathias, Landau, Bernard M. & Merle, Didier, 2025, The Muricidae (Gastropoda, Muricoidea) of the Miocene Paratethys Sea (Aspellinae, Ergalataxinae, Coralliophilinae, Rapaninae), Zootaxa 5611 (1), pp. 1-106 : 36-38

Pazinotus hamulifer ( Boettger, 1906)

Figs 14 , 20A–G

* Murex (Muricidea) hamulifer n. sp. — Boettger 1906: 41.

Muricidea hamulifer (Boettger) — Zilch 1934: 251, pl. 15, figs. 77a–b.

[ Hexaplex ] hamulifer Boettger, 1906 — Vokes 1971: 56.

Hexaplex hamulifer ( Boettger, 1906) — Stojaspal 1978: 334.

Paziella hamulifer ( Boettger, 1906) — Merle et al. 2011: 164, 508, pl. 132, fig. 6.

Pazinotus martonszaboi n. sp. — Kovács & Vicián 2024: 13, figs. 29–32.

Type material. Lectotype (designated by Zilch 1934: 251) SFM 373038 (=12-2275a), SL: 13.5 mm, MD: 9.6 mm, CoŞteiu de Sus ( Romania). illustrated in Zilch (1945: pl. 15, fig. 77), figs. 20A 1 –A 2.

Illustrated material. NHMW 1863/0015/0665, SL: 9.9 mm, MD: 6.5 mm, Niederleis ( Austria), figs. 20B 1 –B 3. NHMW 1862/0001/0226a, SL: 9.8 mm, MD: 7.0 mm, Baden ( Austria), figs. 20C 1 –C 3. NHMW 1862/0001/0226b, SL: 10.5 mm, MD: 7.8 mm, Baden ( Austria), figs. 20D 1 –D 3. NHMW 1866/0001/0843a, SL: 10.2 mm, MD: 6.5 mm, Niederleis ( Austria), figs. 20E 1 –E 3. NHMW 1866/0001/0843b, SL: 9.4 mm, MD: 5.9 mm, Niederleis ( Austria), Fig. 20F . NHMW 1866/0001/0843c, SL: 9.3 mm, MD: 6.1 mm, Niederleis ( Austria), Fig. 20G .

Revised description. Small, broadly fusiform shell with conical spire and deeply incised suture; apical angle ~56°. Protoconch conical of three smooth, weakly convex whorls. Early teleoconch whorls shouldered with P1 and P2 appearing simultaneously at protoconch/teleoconch junction. Subsutural ramp rapidly widening from first to second whorl. Axial sculpture of eight foliose axial ribs forming moderate length recurved spines over P1, very short spines over P2. Weak s1 appearing on third teleoconch whorl and weak s2 on penultimate whorl. Periphery at P2 on first two whorls, at P1 on later whorls. Subsutural ramp convex with weak IP appearing on third whorl; even weaker adis followed by abis on penultimate whorl. P3 appears at abapical suture on penultimate whorl. Last whorl attaining 70% of total height, strongly shouldered with conical base. Axial sculpture of six prominent, weakly foliose varices. Subsutural ramp convex with delicate IP, even weaker adis, abis; P1 forming very long adapically curved spines on varices; P2–P5 and s1–s5 well developed; P6 weaker; ADP, MP and ABP forming short spines; primary and secondary cords terminating in narrow spines at aperture. Aperture pyriform. Outer lip with weakly crenulate edge; ID weak, D1–D5 prominent. Anal canal indistinct. Siphonal canal moderately long, open, narrow, dorsally recurved. Columella moderately excavated in adapical half, slightly angled at transition to siphonal canal. Columellar callus forming broad rim, adherent in parietal area, sharply delimited from base.

Paratethyan synonyms. Pazinotus martonszaboi Kovács & Vicián, 2024 , holotype: HNHM PAL 2024.6.1, SL: 12.1 mm, MD: 7.6 mm, Tekeres ( Hungary), illustrated in Kovács & Vicián (2024: figs. 29–30); paratype: HNHM PAL 2024.7.1, SL: 10.5 mm, MD: 6.9 mm, Letkés ( Hungary), illustrated in Kovács & Vicián (2024: figs. 31–32). The specimens fall within the range of morphological variability of Pazinotus hamulifer ( Boettger, 1906) as understood herein.

Discussion. The lectotype of Pazinotus hamulifer ( Boettger, 1906) is a specimen with strongly curved spine on the terminal varix and narrow, spinose secondary cords. Additional material from the Vienna Basin documents some variability in these features. The terminal spine may be less prominent, straight, or largely reduced, and the cords may be broader ( Fig. 20B ). This species was placed by Merle et al. (2011) in Paziella Jousseaume, 1880 [type species Murex pazi Crosse, 1869 ; present-day, Western Atlantic] but the new material suggests placement in Pazinotus E.H. Vokes, 1970 .

Paleoenvironment. The occurrence at the locality Letkés ( Hungary) suggests inner neritic environments with corals ( Kovács & Vicián 2014).

Distribution in Central Paratethys. Badenian (Middle Miocene): Vienna Basin: Baden, Niederleis ( Austria) (hoc opus); Mecsek Mountains: Tekeres ( Hungary), Pannonian Basin: Letkés ( Hungary) Kovács & Vicián (2024); Făget Basin: CoŞteiu de Sus ( Romania) ( Boettger 1906).

Figures

FIGURE 14. Cord diagrams and apertural features of Aspellinae (Muricopsis, Murexsul, Pazinotus).

FIGURE 20. Pazinotus hamulifer (Boettger, 1906). A1–A2. lectotype, SFM 373038, CoŞteiu de Sus (Romania). B1–B3. NHMW 1863/0015/0665, Niederleis (Austria). C1–C3. NHMW 1862/0001/0226a, Baden (Austria). D1–D3. NHMW 1862/0001/0226b, Baden (Austria). E1–E3. NHMW 1866/0001/0843a, Niederleis (Austria). F. NHMW 1866/0001/0843b, Niederleis (Austria). G. NHMW 1866/0001/0843c, Niederleis (Austria).