Cathymorula exilis (Hörnes, 1852)

Harzhauser, Mathias, Landau, Bernard M. & Merle, Didier, 2025, The Muricidae (Gastropoda, Muricoidea) of the Miocene Paratethys Sea (Aspellinae, Ergalataxinae, Coralliophilinae, Rapaninae), Zootaxa 5611 (1), pp. 1-106 : 45-47

Cathymorula exilis (Hörnes, 1852)

Figs 26 , 27A–C

Purpura exilis Partsch — Hauer 1837: 417 [nomen nudum].

Purpura exilis Partsch — Hörnes 1848: 17 [nomen nudum].

* Purpura exilis Partsch —Hörnes 1852: 169 (pars), pl. 13, figs. 20–23.

Purpura exilis Partsch —Roemer 1870: 380, pl. 47, fig. 7.

Purpura (Stramonita) exilis Partsch —Hoernes & Auinger 1882: 152 (pars).

Purpura subumbilicata Bell. — Bellardi 1882: 175, pl. 11, fig. 2.

Purpura exilis — Quenstedt 1884: 653, pl. 211, fig. 33.

Purpura (Stramonita) exilis Ptsch. — Boettger 1906: 34.

Purpura exilis Partsch — Friedberg 1912: 134, pl. 7, fig. 9.

Purpura exilis Partsch — Friedberg 1938: 118.

Thais (Stramonita) exilis Partsch — Csepreghy-Meznerics 1954: 37, pl. 4, figs. 12–13.

Thais (Stramonita) exilis Partsch — Korobkov 1955: plate captions, pl. 73, figs. 13a–b.

Th [ais]. (S [tramonita].) exilis (Partsch) — Sieber 1958: 146.

Thais (Stramonita) exilis (Partsch in Hoernes 1856)—Kojumdgieva in Kojumdgieva & Strachimirov 1960: 153, pl. 41, fig. 2.

Thais (Stramonita) exilis Partsch (in Hörnes), 1856— Strausz 1966: 283, text-fig. 129b, pl. 64, figs. 2, 5.

Thais (Stramonita) exilis Partsch — Csepreghy-Meznerics 1972: 27, pl. 9, fig. 22.

Thais exilis (Partsch in Hoernes)— Krach 1981: 69 (pars), pl. 17, figs. 8, 10–11, pl. 22, fig. 13 [non pl. 18, figs. 7–8].

Thais (Stramonita) exilis (Partsch in Hörnes, 1856)— Bałuk 1995: 222, pl. 27, figs. 1–4.

Thais (Stramonita) exilis (Partsch in Hörnes, 1856)— Harzhauser 2002: 94, pl. 6, fig. 6.

Thais (Stramonita) exilis (Partsch in Hörnes)— Bałuk 2006: 208, pl. 12, figs. 4–7 [non figs. 3,5 = Cathymorula exilissima sp. nov.].

Cathymorula exilis (Hörnes) — Landau et al. 2013: 158, pl. 23, fig. 9.

Thais (Stramonita) exilis (Partsch in Hörnes)— Popa et al. 2014: 14, pl. 4, fig. 5.

Cathymorula exilis (Partsch in Hörnes, 1852)— Kovács et al. 2018: 128, figs. 8T–V.

Cathymorula exilis (Hörnes, 1852) — Kovács 2019: 134.

Cathymorula exilis (Hörnes) — Kovács 2020: 464, pl. 4, figs. 13–14.

Cathymorula exilis (Hörnes) — Kovács & Vicián 2024: 24, fig. 64.

Type material. Lectotype (designated herein): NHMW 1846/0037/0165, SL: 30.6 mm, MD: 19.3 mm, Steinebrunn ( Austria), illustrated in Hörnes (1853, pl. 13, fig. 21), figs. 27A 1 –A 3 . Paralectotypes: NHMW 1860/0001/0167a, SL: 33.1 mm, MD: 19.2 mm, Mikulov ( Czech Republic), figs. 27B 1 –B 3. NHMW 1860/0001/0167b, SL: 27.3 mm, MD: 17.4 mm, Mikulov ( Czech Republic), figs. 27C 1 –C 3 .

Additional material. 25 spec., NHMW 1855/0045/0411, Gainfarn ( Austria) and Mikulov ( Czech Republic); 3 spec., NHMW A1520, Enzesfeld ( Austria) ; 2 spec., NHMW 1864/0042/0007, Niederkreuzstetten ( Austria) ; 5 spec., NHMW 1866/0040/0187, Baden-Sooss ( Austria) ; 1 spec., NHMW 1849/0001/0250, Möllersdorf ( Austria) ; 12 spec., NHMW 1866/0001/1050, Forchtenau ( Austria) ; 5 spec., NHMW 1853/0010/0014, Enzesfeld ( Austria) ; 7 spec., NHMW 1869/0001/0470, Grund ( Austria) ; 2 spec., NHMW 1847/0046/0018, Szob ( Hungary) ; 7 spec., NHMW 1870/0033/0066, Lăpugiu de Sus ( Romania) .

Revised description. Medium-sized, solid, ovate shell; apical angle ~80°. Protoconch conical, of 2.5 whorls. Teleoconch of up to 4.5 whorls. Suture weakly impressed, shallowly undulating. Early teleoconch whorls with weakly concave subsutural ramp and shoulder (marked by P1) placed short distance above suture. Sculpture of narrow adis, IP, abis and subequal secondary cords and nine rounded nodes along shoulder (P1). Last whorl attaining ~82–85% of total height; ovate with broad, weakly concave subsutural ramp, rounded shoulder, convex periphery, moderately constricted below. Numerous weak spiral cords on subsutural ramp (including SP, adis, IP and abis) about nine to ten weak nodes along shoulder. P1–P4 on convex part of whorl, P5–P6 on siphonal canal. Primary cords broad, flattish, bifid cords occasionally with low, indistinct nodes. Secondary cords of subequal strength (s1–s4), separated by narrow grooves. Fasciole swollen, relatively short, delimiting broad, deep pseudoumbilicus. Aperture moderately narrowly ovate; outer lip thin with weak ID and prominent D1–D4, passing into four lirae persisting far into aperture. Anal canal deeply incised, U-shaped, accentuated by blunt parietal denticle. Siphonal canal, moderately short, wide, open, straight. Columella deeply excavated in adapical half, almost straight below, with faint denticle and weak columellar fold at transition to siphonal canal. Columellar callus forming broad rim, poorly delimited from base, weakly erect at siphonal canal, adherent above.

Discussion. Purpura exilis was first used as name for a Miocene species by Paul Maria Partsch (1791–1856) on collection labels and was made available by Hörnes (1852). Both had overlooked that the name was already used by Link (1807: 121), who provided a German description for an extant species. We are not aware of this name being used as a valid name after 1899 and therefore we refer to Article 23.9.1.1. of the ICZN (1999) and assume that Purpura exilis Link, 1807 is a nomen oblitum.

Cathymorula cathyae Landau, Houart & C. M. Silva, 2007 , from the Pliocene of the Mediterranean Sea, differs in its broader spiral cords, the weaker shoulder, which lacks nodes, and develops the presence of five denticles (ID simple, D1 bifid, D2 to D4 simple) in the outer lip (see Landau et al. 2007: pl. 14, figs. 1–4). Similarly, Cathymorula stazzanensis ( Bellardi, 1882) , from the Tortonian of Italy and Portugal ( Bellardi 1882: pl. 11, fig. 23; Ferrero Mortara et al. 1981: pl. 33, fig. 3; Landau et al. 2007: pl. 14, fig. 5) lacks an angled shoulder and has more numerous denticles (ID simple, D1 bifid, D2 probably bifid, D3–D4 simple).

Paleoenvironment. Shallow marine, inner neritic.

Distribution in Central Paratethys. Karpatian (Early Miocene): Korneuburg Basin: Stetten ( Austria) ( Harzhauser 2002); Vienna Basin: Niederkreuzstetten ( Austria) (Hoernes & Auinger 1882); Badenian (Middle Miocene): Korytnica Basin: Korytnica ( Bałuk 1995); Polish-Ukrainian Fore-Carpathian Basin: Rybnica ( Poland) ( Friedberg 1938); Vienna Basin: Bad Vöslau, Baden, Enzesfeld, Gainfarn, Pfaffstätten, Steinebrunn, Vienna / Pötzleinsdorf, Vienna /Grinzing ( Austria), Mikulov ( Czech Republic) (Hoernes & Auinger 1882); Eisenstadt-Sopron Basin: Forchtenau ( Austria) (Hoernes & Auinger 1882); Pannonian Basin: Letkés, Szob ( Hungary), Mecsek Mountains: Hidas, Hosszúhetény, (Hoernes &Auinger 1882; Strausz 1966; Kovács 2020; Kovács et al. 2018; Kovács & Vicián 2024); Bakony Mountains: Bánd ( Hungary) ( Kovács 2020); Bükk Mountains: Borsodbóta ( Hungary) ( Csepreghy-Meznerics 1972); Cserhát Hills: Kelet-Cserhát, Márkháza, Sámsonháza, ( Hungary) ( Csepreghy-Meznerics 1972); Făget Basin: CoŞteiu de Sus, Lăpugiu de Sus ( Romania) ( Kovács 2019; hoc opus); Dacian Basin: Târnene, Staropatica ( Bulgaria) (Kojumdgieva in Kojumdgieva & Strachimirov 1960).

Proto-Mediterranean Sea. Serravallian (Middle Miocene): Karaman Basin: Akpınar, Seyithasan ( Turkey) ( Landau et al. 2013).

Figures

FIGURE 26. Cord diagrams and apertural features of Ergalataxinae.

FIGURE 27. Cathymorula exilis (Hörnes, 1852). A1–A3. lectotype, NHMW 1846/0037/0165, Steinebrunn (Austria). B1–B3. paralectotype, NHMW 1860/0001/0167a, Mikulov (Czech Republic). C1–C3. paralectotype, NHMW 1860/0001/0167b, Mikulov (Czech Republic).