Favartia suboblonga (d’Orbigny, 1852)

Harzhauser, Mathias, Landau, Bernard M. & Merle, Didier, 2025, The Muricidae (Gastropoda, Muricoidea) of the Miocene Paratethys Sea (Aspellinae, Ergalataxinae, Coralliophilinae, Rapaninae), Zootaxa 5611 (1), pp. 1-106 : 25-27

Favartia suboblonga (d’Orbigny, 1852)

Figs 7 , 13A–D

Murex oblongus — Grateloup 1833: 100 [non Murex oblongus Brocchi, 1814 = Pleurotomoides gracilis ( Montagu, 1803) ]. Morex oblongus — Grateloup 1845: pl. 3, fig. 13 [non Murex oblongus Brocchi, 1814 = Pleurotomoides gracilis ( Montagu,

1803)]. * Murex suboblongus d’Orb. —d’Orbigny 1852: 73, no. 1334 [nom. nov. pro Murex oblongus Grateloup, 1833 , non Brocchi,

1814]. Murex incisus Broderip —Hörnes 1853: 223, pl. 23, figs. 7a–c [non Favartia incisa (Broderip, 1833) ]. Murex incisus Brod. — D’Ancona 1871: 23, pl. 4, fig. 6. Murex (Muricidea) incisus Brod. —Hoernes & Auinger 1885: 206 [non Favartia incisa (Broderip, 1833) ]. Murex (Muricidea) incisus Brod. — Boettger 1902: 28 [non Favartia incisa (Broderip, 1833) ]. Murex (Favartia) incisus Brod. — Sacco 1904: 20, pl. 5, figs. 19–22 [non Favartia incisa (Broderip, 1833) ]. Murex (Muricidea) incisus Brod. — Boettger 1906: 44 [non Favartia incisa (Broderip, 1833) ]. Murex (Favartia) excisus var. Peyroti n.— Montanaro 1935: 27, pl. 2, fig. 11. AspelIa (Favartia) incisa excisa Grateloup, 1833 — Glibert 1952: 298, pl. 6, fig. 10 [non Favartia excisa ( Grateloup, 1833) ]. A [spella]. (F [avartia].) incisa excisa Grat. — Sieber 1958: 146. [ Favartia ] suboblongus d’Orbigny, 1852 — Vokes 1971: 103. Pterynotus excisus ( Grateloup, 1833) — Stojaspal 1978: 334 [non Favartia excisa ( Grateloup, 1833) ]. Favartia incisa ( Broderip, 1832) — Inzani & Bertarelli 1985: 27, pl. 1, fig. 4 [non Favartia incisa (Broderip, 1833) ]. Favartia incisa ( Broderip, 1832) — Andreoli & Marsigli 1992: 7, pl. 2, fig. 4 [non Favartia incisa (Broderip, 1833) ]. Dermomurex scalaroides (Blainville, 1829) — Cavallo & Repetto 1992: 86, fig. 185 [non Dermomurex scalaroides (de Blainville,

1829)]. Favartia cf. incisa ( Broderip, 1832) — Cavallo & Repetto 1992: 90, fig. 194 [non Favartia incisa (Broderip, 1833) ]. Favartia cf. incisa ( Broderip, 1832) —Muñiz-Solis & Guerra-Merchán 1994: 28, figs. 6C–D [non Favartia incisa (Broderip,

1833)]. Favartia (Favartia) suboblonga (d’Orbigny) — Vokes 1994: 122, pl. 19, fig. 3. AspelIa (Favartia) incisa ( Broderip, 1832) — Bałuk 1995: 221, pl. 26, figs. 7–9, pl. 38, fig. 5 [non Favartia incisa (Broderip,

1833)]. Favartia suboblonga (d’Orbigny, 1852) — Merle 1999: 300, text-figs 39, 40, pl. 5, figs. 7–9, pl. 6, figs. 1–4. Favartia excisa ( Grateloup, 1833) — Chirli 2000: 23, pl. 11, figs. 9–12 [non Favartia excisa ( Grateloup, 1833) ]. Favartia incisa ( Broderip, 1832) — Chirli 2000: 24, pl. 11, figs. 13–16 [non Favartia incisa (Broderip, 1833) ]. Favartia suboblonga (d’Orbigny) — Landau et al. 2007: 46, text-fig. 12, pl. 12, figs. 5–8. Favartia suboblonga (d’Orbigny, 1852) — Landau et al. 2013: 161, pl. 24, fig. 5. Favartia suboblonga (d’Orbigny, 1852) — Goret et al. 2013: 17, text-figs 13a–b, pl. 4, fig. 6. Favartia suboblonga (d’Orbigny) — Ceulemans et al. 2016: 40, pl. 1, figs. 8–9. Favartia (s.s.) suboblonga (d’Orbigny, 1852) — Kovács et al. 2018: 121, figs. 5Q–T. Favartia (Favartia) suboblonga (d’Orbigny, 1852) — Kovács 2019: 124, figs. 34–35. Favartia suboblonga (d’Orbigny, 1852) — Landau et al. 2019: 21, pl. 18, figs. 1–3. Favartia (Favartia) suboblonga (d’Orbigny, 1852) — Kovács 2020: 455, pl. 2, figs. 18–19. Favartia suboblonga (d’Orbigny, 1852) — Merle et al. 2022: 280, pl. 56, figs. 8–14.

non Murex (Favartia) suboblongus d’Orb. — Cossmann & Peyrot 1924: 130, no. 752, pl. 12, figs. 21–22 [= unnamed species according to Merle 1999].

non Murex (Favartia) suboblongus d’Orb. — Cossmann & Peyrot 1924: 130, no. 752, pl. 12, fig, 23 [= Pygmaepterys subdecussatus (d’Orbigny, 1852) ].

non Murex (Favartia) suboblongus var. quadrangulatu s nov. var.— Cossmann & Peyrot 1924: 131, no. 753, pl. 12, figs. 31, 32, 43 [= Favartia excisa ( Grateloup, 1833) ].

Type material. One syntype (Grateloup‘s coll. n°441, see Merle (1999: pl. 5, fig. 7) is stored in the University of Bordeaux ( France).

Illustrated material. NHMW 1855/0045/0081a, SL: 15.8 mm, MD: 10.0 mm, Steinebrunn ( Austria), illustrated in Hörnes (1853: pl. 23, fig. 7), figs. 13A 1 –A 3. NHMW 1871/0011/0016, SL: 13.3 mm, MD: 8.6 mm, Steinebrunn ( Austria), figs. 13B 1 –B 3. NHMW 1865/0001/0197, SL: 20.1 mm, MD: 13.6 mm, Lăpugiu de Sus ( Romania), figs. 13C 2 –C 3. NHMW 1853/0043/0056, SL: 16.7 mm, MD: 11.3 mm, Lăpugiu de Sus ( Romania), figs. 13D 1 –D 3.

Additional material. 10 spec., NHMW 1855/045/0081b, Steinebrunn ( Austria) .

Description. See Landau et al. (2007: 47).

Discussion. See Landau et al. (2007; 47).

Paleoenvironment. Shallow marine, inner neritic.

Distribution in Central Paratethys. Badenian (Middle Miocene): Korytnica Basin: Korytnica ( Bałuk 1995); Vienna Basin: Steinebrunn ( Austria), Porzteich ( Czech Republic) (Hoernes & Auinger 1885); Pannonian Basin: Letkés, Mecsekpölöske ( Hungary) ( Kovács 2019, 2020; Kovács et al. 2018).

Făget Basin: CoŞteiu de Sus, Lăpugiu de Sus ( Romania) (Hoernes & Auinger 1885; Boettger 1902).

(Proto-)Mediterranean Sea. Tortonian (Late Miocene): Po Basin: Montegibbio ( Italy) ( Montanaro 1935). Pliocene: Spain ( Landau et al. 2007), Italy ( Chirli 2000, Cavallo & Repetto 1992; Landau et al. 2007).

Northeastern Atlantic. Aquitanian (Early Miocene): Aquitaine Basin: Saint-Paul-lès-Dax (Maïnot) (Grateloup 1847), Meilhan ( Merle 1999). Langhian (Middle Miocene): Loire Basin: Bossée, Ferrière-Larçon, La Chapelle- Blanche (La Houssaye), Louans, Manthelan, Paulmy, Pontlevoy, Sainte-Maure, Sainte-Catherine-de-Fierbois, Sepmes (Grande Barangerie) ( France) ( Glibert 1952).

Genus Muricopsis Bucquoy & Dautzenberg, 1882

Type species. Murex blainvillii Payraudeau, 1826 [= Muricopsis cristata ( Brocchi, 1814) ]; original designation by Bucquoy & Dautzenberg (1882: 16). Present-day, Mediterranean Sea.

Remarks. Middle Miocene specimens of Muricopsis from the Paratethys and the Proto-Mediterranean Sea have often been confused with the extant Muricopsis cristata ( Brocchi, 1814) , which originated during the Pliocene. In the last years it has become clear that the species concept applied since Hörnes (1853) was too broad and that additional species occurred in the European Neogene. Recently Merle et al. (2022) recognized that specimens from the Serravallian of the Karaman Basin represent a distinct species, which he described as Muricopsis landaui Merle in Merle et al. 2022. Specimens from the Badenian of the Paratethys were compared with Muricopsis dujardini Peyrot, 1938 , originally described from the Langhian of the Loire Basin, by Kovács et al. (2018) or have been listed in open nomenclature as Muricopsis cf. cristata by Merle et al. (2022). This Paratethyan species is discussed herein as Muricopsis emus ( De Gregorio, 1885) . In addition, a larger, spinose species occurred in the southern Paratethyan Basins, which is described herein as Muricopsis hystrix sp. nov. Murex holubicensis Friedberg, 1912 , placed in Muricopsis by Korobkov (1955) and Vokes (1971), was removed from this genus and was placed in Ocenebra by Harzhauser et al. (2024). Similarly, Murex (Muricidea) moravicus Hoernes & Auinger, 1885 , which was placed in Muricopsis by Kovács (2019), is excluded from Muricopsis and placed in Murexsul herein.

Figures

FIGURE 7. Cord diagrams and apertural features of Aspellinae (Favartia).

FIGURE 13. Favartia suboblonga (d’Orbigny, 1852). A1–A3. NHMW 1855/0045/0081a, Steinebrunn (Austria). B1–B3. NHMW 1871/0011/0016, Steinebrunn (Austria). C1–C3. NHMW 1865/0001/0197, Lăpugiu de Sus (Romania). D1–D3. NHMW 1853/0043/0056, Lăpugiu de Sus (Romania).