Attiliosa evae ( Kovács & Vicián, 2024 )

Harzhauser, Mathias, Landau, Bernard M. & Merle, Didier, 2025, The Muricidae (Gastropoda, Muricoidea) of the Miocene Paratethys Sea (Aspellinae, Ergalataxinae, Coralliophilinae, Rapaninae), Zootaxa 5611 (1), pp. 1-106 : 10

Attiliosa evae ( Kovács & Vicián, 2024)

Figs 1 , 3A

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* Gracilipurpura? evae n. sp. — Kovács & Vicián 2024: 16, figs. 35–36.

Type material. Holotype: HNHM PAL 2024.8.1., SL: 19.0 mm, MD: 10.0 mm, Letkés ( Hungary), illustrated in Kovács & Vicián (2024: figs. 35–36), figs. 1, 3A 1 –A 2.

Description. See Kovács & Vicián (2024: 16). Antepenultimate whorl: P1 and P2 more prominent than s1; penultimate whorl: P1, s1, P2 prominent, IP weak; last whorl: delicate adis, IP, abis and tertiary cord on sutural ramp, P1–P6 and s1–s6 on convex part of whorl, s4–s5 weaker, short P1 spine; ADP–EAB1 and abs on siphonal canal.

Discussion. Based on its general morphological similarity (slender shape, numerous spiral cords, aperture devoid of internal denticles) to the Miocene—Recent Gracilipurpura craticulata ( Bucquoy & Dautzenberg, 1882) , this species was provisionally assigned to Gracilipurpura Jousseaume, 1880 with question mark on advice of one author of the present paper (DM). The problem of this generic attribution is that species of Gracilipurpura display a sealed canal when the aperture is completely built and have a longer siphonal canal than Attiliosa evae ( Kovács & Vicián, 2024) . A deeper analysis of the first teleoconch whorl shows that Gracilipurpura craticulata bears IP, P1, s1, P2, whereas that Attiliosa evae bears only P1, the series IP, P1, s1, P2 appearing on second or third whorl. The short P1 spine on the first teleoconch whorl and the late appearance of P1, IP, s1 and P2 are reminiscent of those of Attiliosa (e.g., Attiliosa houarti Vokes, 1999 , present-day Thailand). Members of Attiliosa usually have denticles within the outer lip, but the aperture of the holotype of Attiliosa evae is not thickened and is probably not completely built. The shape with an elongate spire and a short siphonal canal is reminiscent of Attiliosa juhaszi Kovács & Vicián, 2024 , but this species differs by having a larger shape and in lacking P1 cord spine.

Paleoenvironment. The occurrence at the locality Letkés ( Hungary) suggests inner neritic environments with corals ( Kovács & Vicián 2014).

Distribution in Central Paratethys. Badenian (Middle Miocene): Pannonian Basin: Letkés ( Hungary) Kovács & Vicián (2024).

Figures

FIGURE 1. Cord diagrams and apertural features of Aspellinae (Aspella, Attiliosa, Dermomurex).

FIGURE 2. A–B. Aspella emmae (Boettger, 1902). A1–A2. lectotype, SFM 366519, CoŞteiu de Sus (Romania). B1–B3. NHMW 1871/0001/0182, Steinebrunn (Austria). C–E. Aspella teter sp. nov. C1–C3. paratype, NHMW 1846/0037/0235, Steinebrunn (Austria). D1–D3. holotype, NHMW 1865/0001/0560a, Steinebrunn (Austria). E1–E3. paratype, NHMW 1865/0001/0560b, Steinebrunn (Austria).

FIGURE 3. A1–A2. Attiliosa evae (Kovács & Vicián, 2024). holotype, HNHM PAL 2024.8.1., Letkés (Hungary).B1–B2. Attiliosa juhaszi Kovács & Vicián, 2024. holotype, HNHM PAL 2024.1.1., Letkés (Hungary).