Janssenia spinosa (Kojumdgieva, 1960)

Harzhauser, Mathias, Landau, Bernard M. & Merle, Didier, 2025, The Muricidae (Gastropoda, Muricoidea) of the Miocene Paratethys Sea (Aspellinae, Ergalataxinae, Coralliophilinae, Rapaninae), Zootaxa 5611 (1), pp. 1-106 : 87-89

Janssenia spinosa (Kojumdgieva in Kojumdgieva & Strachimirov, 1960)

Figs 48 , 49D–E

Purpura (Sistrum) inconstans Michti. —Hoernes & Auinger 1882: 152, pl. 16, figs. 10–13 [non Purpura inconstans Michelotti, 1847 ].

Ricinula (Sistrum) inconstans Michti. — Boettger 1906: 34 [non Purpura inconstans Michelotti, 1847 ].

* Thais (Stramonita) spinosa n. sp. — Kojumdgieva in Kojumdgieva & Strachimirov 1960: 153, pl. 41, fig. 3.

Thais (Stramonita) austriaca Hoernes & Auinger — Krach 1981: 69, pl. 22, fig. 10 [non Claremontiella pauli ( Hilber, 1882) ].

Janssenia spinosa ( Kojumdgieva, 1960) — Kovács et al. 2018: 126, figs. 8R–S.

Janssenia spinosa ( Kojumdgieva, 1960) — Kovács 2019: 134.

Thais (Stramonita) spinulosa [sic] Kojumdgieva & Strachimirov, 1960 — Landau et al. 2013: 157.

Janssenia spinosa (Kojumdgieva) — Kovács & Vicián 2024: 24, fig. 63.

Type material. We are not aware of the whereabouts of the type specimens.

Illustrated material. NHMW 1867/0019/0065, SL: 26.8 mm, MD: 16.3 mm, CoŞteiu de Sus ( Romania), illustrated in Hoernes & Auinger (1882: pl. 16, fig. 13), figs. 49D 1 –D 3. NHMW 2024/0186/0001 (A 1519), SL: 23.3 mm, MD: 15.7 mm, Lăpugiu de Sus ( Romania), illustrated in Hoernes & Auinger (1882: pl. 16, fig. 12), figs. 49E 1 –E 3.

Revised description. Medium-sized, moderately broad ovate species; apical angle ~66°. Protoconch unknown. Teleoconch consisting of up to five whorls. Suture superficial, shallowly undulating. Early teleoconch whorls with deeply concave subsutural ramp and shoulder at abapical suture with about 8–10 rounded nodes. Spiral sculpture of numerous delicate threads including indistinct SP, adis, IP, abis. Later whorls with broad subsutural collar and prominent, subtrigonal tubercles corresponding to P1 placed just above suture. Last whorl attaining ~75% of total height; with subsutural collar, concave subsutural ramp, angled shoulder, subcylindrical periphery, moderately constricted below. Spiral sculpture of prominent P1 and P2 with eight to nine prominent, trigonal tubercles; P3 atrophied; P4 prominent delimiting base. P5, P6 weak. Secondary cords s1 to s5 broad, flattish separated by narrow grooves, overrun by delicate tertiary threads. Fasciole strongly swollen, twisted, delimiting deep, broad pseudoumbilicus. Aperture moderately narrow, ovate; outer lip thin, with weak D1–D4 (see Kovács et al. 2018: 126, fig. 8R). Anal canal deeply incised, narrowly V-shaped. Siphonal canal moderately short, wide, open, slightly bent to the left. Columella straight, smooth, faintly angled at siphonal canal. Columellar callus poorly delimited, adherent.

Discussion. Janssenia spinosa (Kojumdgieva in Kojumdgieva & Strachimirov, 1960) differs from Janssenia echinulata (Pusch, 1837) in its smaller size, spiny tubercles and atrophied P3. Janssenia connectens ( Bellardi, 1884) , from the Early or Middle Miocene of the Colli Torinesi ( Italy) and Janssenia tuberculata ( Bellardi, 1884) , from the Burdigalian of Valle Ceppi ( Italy) are both comparable in sculpture and apertural features but differ in their more inflated outline.

Paleoenvironment. Shallow marine, inner neritic.

Distribution in Central Paratethys. Badenian (Middle Miocene): Bakony Mountains: Bánd ( Hungary) ( Kovács 2020); Pannonian Basin: Letkés, Márkháza ( Hungary) ( Kovács et al. 2018; Kovács & Vicián 2024); Făget Basin: CoŞteiu de Sus, Lăpugiu de Sus ( Romania) (Hoernes & Auinger 1882; Kovács 2019); Dacian Basin: Târnene, Staropatica ( Bulgaria) (Kojumdgieva in Kojumdgieva & Strachimirov 1960).

Genus Zoltania nov. gen.

Type species. Zoltania styriaca ( Hilber, 1879) ; Badenian (Langhian), Middle Miocene; Austria, Hungary, Romania, Central Paratethys Sea.

Diagnosis. Medium-sized, solid, stocky biconic shell with low conical spire; early teleoconch whorls low, with very broad, shallow, weakly concave subsutural ramp, shoulder placed at or just above suture. weak axial ribs overrun by prominent spiral cords; last whorl with broad, strongly scabrose subsutural collar, concave subsutural ramp, strongly angled shoulder, slowly contracting below; broad axial ribs with strongly developed primary (P1– P4) and secondary cords (s1–s4); primary cords P1 to P4 wide, duplicated in appearance and split by a small spiral groove; outer lip thickened, with paired crenulations at edge, with ID, D1–D4 placed short distance behind peristome; columella with prominent, broad central columellar fold.

Description. As for type species.

Etymology. In honor of Zoltán Kovács (Liszt Ferenc Academy of Music, Budapest) and Zoltán Vicián (Budapest), in recognition for their contributions on Paratethyan paleontology.

Included species. Zoltania styriaca ( Hilber, 1879) , Badenian (Middle Miocene), Central Paratethys Sea. Cymia (s. l.) cf. styriaca of Lozouet (2023), Burdigalian (Early Miocene), Léognan ( France), northeastern Atlantic (this species will need a new name).

Stratigraphic and geographic range. Early Miocene to Middle Miocene; northeastern Atlantic and Central Paratethys Sea.

Discussion. Zoltania is reminiscent of the extant Semiricinula von Martens, 1879 [type species Purpura muricina de Blainville, 1832 ; present-day, Indo-West Pacific Region], which has a comparable mode of sculpture formation and may develop up to two columellar folds ( Raven 2016) (e.g., S. muricina (de Blainville, 1832) , S. hadrolineae ( Houart, 1996) . However, the columellar folds are weaker, and most Semiricinula species have higher spires, a thinner peristome, and they develop lirae, which start some distance behind the peristome (e.g., Semiricinula turbinoides (de Blainville, 1832) , S. tongasoa Bozzetti, 2018 , S. bozzettii Houart & Héros, 2013 ), whereas only subadult specimens of Zoltania have lirae.

Janssenia Landau, Harzhauser, İslamoğlu & C. M. Silva, 2013 differs in its higher spire, the thinner peristome and the four to five large denticles placed some distance behind the peristome

Paleoenvironment. Shallow marine, inner neritic lagoonal environments.

Figures

FIGURE 48. Cord diagrams and apertural features of Rapaninae.

FIGURE 49. A–C. Janssenia echinulata (Pusch, 1837). A1–A3. NHMW 1851/0010/0027, Mikulov (Czech Republic). B1–B3. NHMW 1861/0033/0030, Buituri (Romania). C1–C3. NHMW 1855/0045/0409, Gainfarn (Austria). D–E. Janssenia spinosa (Kojumdgieva in Kojumdgieva & Strachimirov, 1960). D1–D3. NHMW 1867/0019/0065, CoŞteiu de Sus (Romania). E1–E3. NHMW 2024/0186/0001 Lăpugiu de Sus (Romania).