Coralliophila badensis, Harzhauser & Landau & Merle, 2025
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publication ID |
https://doi.org/10.11646/zootaxa.5611.1.1 |
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publication LSID |
lsid:zoobank.org:pub:78FDE0BC-8C7A-4E67-B387-71A58ADD333D |
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persistent identifier |
https://treatment.plazi.org/id/039A879F-FF8D-FFC0-2785-D02DFC30FE7E |
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treatment provided by |
Plazi |
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scientific name |
Coralliophila badensis |
| status |
sp. nov. |
Coralliophila badensis sp. nov.
Figs 37 View FIGURE 37 , 39A–D View FIGURE 39
Pyrula granifera Michti View in CoL —Hörnes 1853: 273, pl. 28, fig. 10 [non Coralliophila granifera ( Michelotti, 1847) View in CoL ].
Pyrula (Rapana) granifera Michti. View in CoL var.— Hoernes & Auinger 1890: 247 [non Coralliophila granifera ( Michelotti, 1847) View in CoL ]. C [oralliophila]. (C [oralliophila].) granifera (Micht.) View in CoL — Sieber 1958: 147 [non Coralliophila granifera ( Michelotti, 1847) View in CoL ]. Coralliophila granifera ( Michelotti, 1847) View in CoL — Kovács 2019: 138, figs. 77–78 [non Coralliophila granifera ( Michelotti, 1847) View in CoL ].
Type material. Holotype: NHMW 2024/0163/0001 (A1442), SL: 20.9 mm, MD: 15.3 mm, Baden-Sooss ( Austria), illustrated in Hörnes 1853: 273, pl. 28, fig. 10, figs. 39A 1 –A 3. Paratypes: NHMW 2024/0163/0002, SL: 19.0 mm, MD: 14.8 mm, Baden-Sooss ( Austria), pl. 28, fig. 10, figs. 39B 1 –B 3. NHMW 2013/0078/0265a, SL: 26.4 mm, MD: 17.6 mm, Baden ( Austria), figs. 39C 1 –C 3. NHMW 2013/0078/0265b, SL: 21.3 mm, MD: 14.6 mm, Baden ( Austria), figs. 39D 1 –D 3. NHMW 2013/0078/0265c, SL: 20.9 mm, MD: 14.4 mm, Baden ( Austria)
Additional material. 2 spec., NHMW 2013/0078/0266, Baden ( Austria) ; 1 spec., 2010/0040/0766, Bad Vöslau ( Austria) ; 3 spec., 2010/0040/0765, Bad Vöslau ( Austria) ; 3 spec., 2010/0040/0767, Bad Vöslau ( Austria) ; 1 spec., 1971/1477/0005, Gainfarn ( Austria) ;
Type locality. Baden ( Austria), Vienna Basin .
Type stratum. Baden Formation.
Age. Middle Miocene, middle Badenian (Langhian).
Etymology. Referring to the city of Baden (Lower Austria).
Diagnosis. Medium-sized, pyriform shell with low conical spire and globose, strongly constricted last whorl with delicate sculpture of subequal primary and secondary cords with delicate tertiary thread in interspaces; P1 placed short distance above anterior suture; aperture wide with long, wide and open siphonal canal.
Description. Medium-sized, pyriform shell with low conical spire; apical angle ~80–86°. Protoconch worn, high conical, with moderately convex whorls. Teleoconch of up to 3.5 whorls. Suture weakly impressed, very shallowly undulating. Early teleoconch whorls weakly convex with broad, straight-sided subsutural ramp and rounded shoulder marked by P1 placed short distance above anterior suture. Axial sculpture of 11 broad, low ribs, weakening over subsutural ramp. Axial ribs overrun by prominent, narrow scabrose adis, IP and abis with slightly weaker tertiary cords. P1 and P2 prominent, close set. Last whorl attaining ~81–85% of total height; with weakly convex subsutural ramp, convex periphery, strongly constricted below. Axial sculpture of about 11 weak, broad axial ribs being moderately prominent on subsutural ramp, fading on periphery, subobsolete on base. Spiral sculpture delicate of subequal adis, IP, abis, P1–P6, s1–s6, ADP, MP, ABP, EAB1, EAB2 and delicate tertiary thread in interspaces; all cords faintly scabrose. Fasciole narrow, moderately swollen, delimiting moderately narrow pseudoumbilicus. Aperture very wide, pyriform; outer lip thin, smooth. Anal canal wide, indistinct; siphonal canal long, wide, open, slightly bent to the left. Columella broadly excavated, smooth, weakly twisted at siphonal canal. Columellar callus indistinct, poorly delimited from base, slightly erect at siphonal canal, adherent at parietal area.
Discussion. Coralliophila badensis sp. nov. is characterized by its very low spire and inflated non-shouldered last whorl with narrow base, and much finer sculpture than usual for the genus, with primary and secondary cords of subequal strength. Coralliophila badensis is reminiscent of some morphotypes of the Pliocene to extant Mediterranean Coralliophila sofiae (Aradas & Benoit, 1876) (e.g., Brunetti 2004: fig. 20), which differs in its much stronger axial ribs, coarser primary cords and higher spire. Coralliophila badensis was identified by Hörnes (1853) as Coralliophila granifera ( Michelotti, 1847) from the Early or Middle Miocene of the Colli Torinesi ( Italy). This view was followed by subsequent authors. However, C. granifera [= Coralliophila varicolonga Sacco, 1904 ] differs in its much higher and slenderer spire and the broader, shallower subsutural ramp (see Michelotti 1847: 266, pl. 17, fig. 6; Bellardi 1882: 196, pl. 12, figs. 7–8) [ Sacco (1904) introduced Coralliophila granifera varicolonga as name for the specimen illustrated by Bellardi (1882) as var. A. The drawing, which shows a deformation on the last whorl, suggests that this specimen is identical to the specimen illustrated in Michelotti (1847: pl. 17, fig. 6). Therefore, Coralliophila granifera varicolonga is an objective synonym of Coralliophila granifera ]. Coralliophila inflatissima Sacco, 1904 , from the Early or Middle Miocene of the Colli Torinesi ( Italy) and introduced as name for the specimen illustrated by Bellardi (1882) as var. B. differs in its coarser spiral sculpture, broader, more prominent axial ribs and the adapically widening aperture ( Bellardi 1882: 196, pl. 12, fig. 6). Coralliophila costata Bellardi, 1882 , from the Early and Middle Miocene of Termô-Fôurà and Sciolze ( Italy), is similar to C. badensis in general shape, but differs in its prominent, fold-like axial ribs and the deep pseudoumbilicus (see Bellardi, 1882 pl. 12, fig. 11). Coralliophila angusta Bellardi, 1882 , from the Middle Miocene of the Villa Forzano ( Italy), differs in its higher spire and less globose last whorl (see Ferrero Mortara et al. 1981: pl. 33, fig. 6). Coralliophila brevispira Bellardi, 1882 , from the Early and Middle Miocene of Termô-Fôurà and Rio della Batteria ( Italy), develops a similar spire but has a narrowly conical base (see Bellardi, 1882: pl. 12, fig. 9).
Specimens from the Early/Middle Miocene of the North Sea Basin, identified as Coralliophila granifera by van Voorthuysen (1944: pl. 6, figs. 12–13, 18) and Stein et al. (2016: pl. 40, fig. 4) differ from the Paratethyan species in their slightly angled periphery with prominent spiral cord and the less constricted base.
Paleoenvironment. Occurrences in the Baden Formation of the Vienna Basin suggests middle to outer neritic environments in up to 250 m water depth (Kranner et al. 2021).
Distribution in Central Paratethys. Badenian (Middle Miocene): Vienna Basin: Baden, Bad Vöslau, Gainfarn ( Austria) (hoc opus); Făget Basin: CoŞteiu de Sus ( Romania) ( Kovács 2019).
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