Astata kashmirensis NURSE , 1909

Hans-J, 2025, Astata LATREILLE, 1796 (Hymenoptera, Astatidae) from the Palearctic region with descriptions of nine new taxa and a key to species, Linzer biologische Beiträge 57 (1), pp. 71-168 : 115-118

Astata kashmirensis NURSE, 1909

Astata kashmirensis NURSE, 1909: 511 , 3. Holotype 3, Kashmir, no specific locality (BMNH).

Astata stecki BEAUMONT, 1942: 407 , ♀, 3. Holotype ♀, Switzerland, Valais, Eusigne (Basel). Synonymized with Astata kashmirensis by PULAWSKI 1975: 320.

Astata stecki melanotica PULAWSKI, 1973: 2, ♀, 3. Holotype ♀, Syria, Damaskus (Torino?).

M a t e r i a l e x a m i n e d: ALBANIA: Valbona , 42°27'N 19°53'E, 08.07.2012, 13, Deneš ( BOL) . ARMENIA: Jerewan, 1000 m, 10.06.2013, 1♀, Schlaefle (WS) . BULGARIA: Beronovo, S. Plan. , 10.08.1997, 533, Zaykov ( BOL) ; Galabovo, Rodopi , 18.06.1996, 13, 15.08.1997, 13, 20.08.1997, 13, Zaykov ( BOL) ; Hrabrino, Rodopi , 20.08.1996, 13, Zaykov ( BOL) ; Ludogorsko Plato , W Vetovo vill., 43°41'57''N 26°14'31''E, 202 m, 14.09.2010, 1♀, Ljubomirov ( SE) ; Nesebar , 08.1970, 13, Kocourek ( BOL) ; Parvenez, Rodopi , 10.07.1996, 13, Zaykov ( BOL) ; Plovdiv, 05.07.1997, 13, Zaykov ( BOL) ; Proslav, Trakia , 15.08.1996, 233, Zaykov ( BOL) ; Slancev Brjag , 05.08.1965, 233, 27.07.1968, 13, 28.06- 14.07.1971, 933, 23♀♀, Padr ( BOL) ; Slancev Brjag , 10.06.1972, 13, Kocourek ( BOL) ; Studenec, Rodopi , 25.07.1997, 13, Zaykov ( BOL) ; Vlahi , 14.08.1993, 13, M. Halada ( BOL) ; Voivodino, Trakia, 05.09.1997, 13, Zaykov ( BOL). CRIMEA: Eupatoria , 10.07.1999, 233, 1♀, Gurko ( BOL) ; Simferopol, Anatra , 07.2000, 1♀, V. Gurko ( BOL) ; Sevastopol , 44°36'N 33°47'E, 08.2009, 13, Kantner ( BOL) . CROATIA: Istrien, Strand 5 km SO Rovinj, 11.08.1994, 13, Schmid-Egger ( SE) ; Pula 10 km N, 27.08.1998, 2♀♀, J. Halada ( BOL) ; Vodnjan env., Istria, 09.07.1999, 13, J. Halada ( BOL) . CYPRUS: Ayia Napa , 20.08.2000, 13 Schlaefle (WS) ; Kouklia , 15 km Se Pafos, 34.72°N 32.55°E, 20.06.2013, 13, Schmid-Egger ( SE) . CZECHIA: Moravia, Znojmo , 48.753°N 16.139°E, 20.07.2020, 1♀, M. Halada ( BOL) ; Pouzdrany , 01.08.1993, 233, M. Halada ( BOL) . FRANCE: Conques s. O., 25.06.2008, 13, Grouet (JA) ; Port la Nouvelle , 26.06.2011, 13, Grouet (JA) ; Alpes maritimes, Tende N, 700-1100 m, 44.09°N 7.60°E, 12.07.2009, 1♀, Schmid-Egger ( SE) . GERMANY: Waren / Müritz , 53.52°N 12.69°E, 05.08.1940, 13, Hainmüller ( MMW) . GREECE: Kreta, Amoudara, St. Marina , 12.06.1993, 13, Tiefenthaler ( BOL) ; Ermioni , 06.1992, 1♀, Schlaefle (WS) ; Peloponnese, 12 km S of Andritsena , 37°24'N 21°50'E, 500 m, 28.08.2019, 3133, J. Halada ( BOL) ; Lesbos, Scala Polichnitos , 09.2014, 13, 1♀, Schlaefle (WS) ; Peloponnese, Marathopoli 20 km N Pilos, 08.07.1996, 433, Ma. Halada ( BOL) ; Epeiros, Preveza , 29.08.1997, 13, J. Halada ( BOL) ; Lebos, Skala Eresou , 19.06.2011, 13, Schlaefle (WS) . HUNGARY: N of Varpalota , 47°12'5''N 18°07'4''E, 05.07.2018, 13, M. Halada ( BOL) . IRAN: Mazdaran prov., Chaloos 10 km S, 380 m, 36°51'N 51°33'E, 15.06.2010, 533, Mi. Halada ( BOL) ; Razawi Ko., Kadkan 2 km S, 1877 m, 35°34'00''N 58°52'41''E, 15.07.2022, 13, W.-H. Liebig ( LI) ; Golestan prov., Minudasht 45 km E, 960 m, 37°36'N 55°93'E, 11.06.2010, 433, Mi. Halada ( BOL) ; Golestan prov., Minudasht NP 40 km E, 750 m, 37°22'N 55°57'E, 01.06.2014, 13, Mi. Halada ( BOL) ; Acer e Shabestar, Sis , 10 km E Shabestar, 38°26'N 45°86'E, 1540 m, 19.06.2010, 733, Mi. Halada ( BOL, JA) . ITALY: Gargano NW, 08.09.2000, 1♀, J. Halada ( BOL) ; Sardinien, Muravera , 02.07.2000, 13, J. Halada ( BOL) ; Toskana, Orbetello Isl. , 06.08.1999, 1533, 5♀♀, J. Halada ( BOL) ; Sizilia, Mt. Madonie S, Petralia Sot env., 600 m, 16.06.2012, 13, J. Halada ( BOL) ; Emilia Romagna, Ravena 60 km N of, 08.07.2000, 1♀, J. Halada ( BOL) ; Schluderns , 1000 m, 06.08.1992, 1♀, Jacobs (JA) . JORDAN: Shuna N env., 29- 30.04.1996, 1♀, Mi. Halada ( BOL) . KAZAKHSTAN: Aksay env. 20 km SE, 19.06.1992, 13, K. Deneš ( BOL) ; Alma Ata , 25.06.1992, 433, 2♀♀, J. Halada, Kocourek, Deneš ( BOL) ; Dzhabagly 3 km W, Talas Mt. R., 42°26'N 69°58'E, 05.08.2000, 13, Makogonova ( BOL) ; Fabrichny , 40 km W of Alma Ata, 09.07.1992, 13, Jirousek ( BOL) ; NO Bajanaul NP, 410 m, 50°49'08''N 75°20'40''E, 18.06.2008, 13, 19.06.2008, 13, W.-H. Liebig ( LI) ; NO Bajanaul NP, 443 m, 50°48'57''N 75°19'20''E, 12.06.2008, 13, W.-H. Liebig ( LI) ; Koktuma, Lake Alakol , 22.06.1995, 733, 3♀♀, M. Mučka, J. Halada ( BOL, JA) ; Kuyuk 40 km SW Djambul, 01.07.1992, 1♀, M. Halada ( BOL) ; Lugovoe 150 km W Djambul, 07.1993, 13, M. Halada ( BOL) ; Malaysari 144 km N Alma-Ata, 21.06.1992, 13, M. Halada ( BOL) . KYRGYZSTAN: Afleatum env., 41°6'N 71°6'E, 01- 03.06.1996, 1♀, J. Halada ( BOL) ; Ala-Buka , 41°5'N 71°2'E, 30.04.1996, 1♀, Mučka ( BOL) ; Alash forest, Alash-Too Mts. , Ferghan. Mt. R., 08.2000, 13, Gurko ( BOL) ; Aravan 25 km N Osh, 06.2000, 13, Gurko ( BOL) ; Burgandi 60 km W Jalal-Abad, 14.06.1995, 333, J. Halada ( BOL) ; Chamadi-Sai , 41°2'N 71°8'E, 30.05.1995, 13, J. Halada ( BOL) ; Frunse (now Bishkek) 75 km S., Cin-Arik , 08.07.1981, 333, Kocourek ( BOL) ; Kashkasu N of Bishkek, Ala-Archa riv., 06.1999, 13, 1♀, Gurko ( BOL) ; Kashka-Suu, Ala-Archa riv., 1650 m, 07.2000, 13, Gurko BOL ); Ooru-Say, Ala-Archa riv., Kirghizsky Mt. R., 1650 m, 07.2000, 13, Gurko ( BOL) ; Toskol-Ata, Ferghanskyi Mts. , 1200 m, 01.08.2001, 233, Gurko ( BOL) ; Toskool-Ata, Ferghan. Mts. , 1500 m, 29.07.2000, 533, Gurko ( BOL) ; Toskool-Ata, Ferghan. Mts., Pistacea forest , 08.2000, 13, Gurko ( BOL) ; Tschui-Tal, Fluss Tschon-Kemin , 1800 m, 42°41'2''N 75°56'42''E, 16.06.2008, 13, Hüttinger ( BOL) ; Uzun-Bulag, Ala-Archa riv., 07.2000, 2♀♀, Gurko ( BOL) . MOROCCO: Agadir 5 km N, 18.05.1997, 2♀♀, J. Halada ( BOL) ; Aghbalou Akourar , 621 m, 33.8627°N 4.6691°W, 17.06.2019, 1♀ L. Hamroud & P. Lhomme ( SE) ; Essaouria , 09.05.1995, 13, Mi. Halada ( BOL) ; Guelmin SE of, 09.05.2003, 1♀, M. Halada ( BOL) ; Guercif 40 km S, 15- 17.05.1995, 1♀, Ma. Halada ( BOL) ; Meknes , 06.1962, 13 Schlaefle (WS) ; Meknes 50 km W, 19.05.1997, 13, M. Halada ( BOL) ; Sidi Ahmed Ben Yussef , 769 m, 33.8349°N 4.7476°W, 16.06.2019, 13, L. Hamroud & P. Lhomme ( SE) ; Sidi Ahmed Ben Yussef , 792 m, 33.8250N 4.7542W, 01- 02.07.2019, 13, L. Hamroud & P. Lhomme ( SE) ; Taroudant, Aoulouz , 17.05.1997, 13, J. Halada ( BOL) ; Tissa env., 08.05.1997, 233, Deneš ( BOL) ; Youssoufia , 11.05.1995, 13, Ma. Halada ( OML) . MONGOLIA: Bayankhongor 86 km NW, 2070 m, 46°50'N 100°04'E, 14.07.2004, 13, Kadlecova ( BOL) ; Ulaanbataar 50 km N, riv. E of Mandal, 1180 m, 13.08.2007, 13, Kadlecova ( BOL) . RUSSIA: Baschkirien, See Aslikul 120 km SW Ufa, Mischwald-Weidesteppe , 09.07.1982, 1♀, Taeger (JA) ; Baikal, Ust-Orda, Steppe , 545 m, 52°50'25''N 104°47'14''E, 26.06.2011, 1♀, Liebig ( LI) ; Primorje, Anisimovka , 43.1830°N 132.7603°E, 06.08.1983, 13, Lelej ( SE) . SLOVAKIA: Chotin , 22.06.1960, 13, Padr ( BOL) ; Chotin , 26.07.1993 233, M. Halada ( BOL) ; Malacky , 05.07.1994, 13, Deneš ( BOL) ; Modra , 14.06.1976, 13, Kocourek ( BOL) ; Pezinok-Baba , 30.06.1952, 13, Kocourek ( BOL) ; Sekule SW, strk. Mlaky , 48.617°N 17.003°E, 22.07.2020, 1♀, M. Halada ( BOL) ; Somotor , 07.1952, 13, Kocourek ( BOL) ; Somotor , 07.1962, 13, Kocourek ( BOL) ; Streda nad Bodrogom , 26.06.1977, 13, Padr ( BOL) ; Sturovo , 26.07.1956, 13, 08.08.1959, 13, 16.07.1962, 13, 27.07.1962, 233, 01.08.1962, 13, Padr ( BOL) ; Sturovo , 06.05.1965, 13, Kocourek ( BOL) ;

Sturovo , 28.07.1993, 233, M. Halada ( BOL) ; Vinicky, 06.1952, 13, Kocourek ( BOL). SPAIN: Palamos , Catal., 07.1994, 13, P. Stary ( BOL) ; Puigcerda , 18.07.1963, 333, 1♀, Hamann ( BOL, JA) ; Pyrenees, Rio Cinqueta-Tal, 32 km SW Bagneres-de-Luchon, 42.40°N 0.30°E, 1200 m, 18.07.1996, 13, Geller-Grimm ( SE). SWITZERLAND: Lenk , Wallis, 27.07.2004, 13, Schlaefle ( BOL) ; Sion, 07.1962, 1♀, Schlaefle (WS). SYRIA: Homs 40 km W, 18.05.2000, 13, M. Halada ( BOL) ; Jisr ash Shugur, 10- 11.05.1996, 1♀, Ma. Halada ( BOL). TAJIKISTAN: Dusanbe 40 km vych. Javrov , 24.06.1981, 13, M. Kocourek ( BOL) ; Dusanbe env., 25.06.1981, 13, M. Kocourek ( BOL). TUNISIA: Monastir Skanes, 35°45'N 10°44'E, 10.05.2001, 13, 14.05.2001, 13, Rosa, det. Pulawski ( SE). TÜRKIYE: Akcali 35 km S Hakkari, 1700 m, 37°71'N 44°03'E, 21.06.2010, 233, Mi. Halada ( BOL) ; Akyaka , pasture and salty meadow, 3 m, 37°03'N 28°20'E, 09.2012, 1♀, V. Bartak ( BOL) ; Alanya 20 km E, 16.06.1997, 13, Ma. Halada ( BOL) ; Avgadi , 30 km NW of Erdemli, 1300 m, 20.06.1996, 1♀, Jelinek ( BOL) ; Buglan Gecidi 40 km NW Mus, 1600 m, 38°56'N 41°09'E, 24.06.2010, 533, Mi. Halada ( BOL) ; Denizli 10 km NE, 290 m, 37°56'N 29°07'E, 04.07.2006, 633, J. Halada ( BOL) ; Denizli 35 km SSE, 970 m, 37°37'N 29°17'E, 05.07.2006, 13, M. Halada ( BOL) ; Denizli, Isikli Golu , 22.06.1998, 13, J. Halada ( BOL) ; Erkenek 60 km SW Malatya, 1300 m, 26.06.2000, 13, M. Halada ( BOL) ; Erzurum E of, 06.07.2000, 533, M. Halada ( BOL) ; Gaziantep 30 km NW, 2001 333, Snižek ( BOL) ; Gurun 20 km E, Mezikiran Gecidi , 10.07.1997, 13, Ma. Halada ( BOL) ; Hakkari, Esendere , 21.07.1988, 13, 1♀, Schmid-Egger ( SE, JA) ; Hakkari, Gözeldere , 20 km E, 930 m, 37°32'N 43°49'E, 22.06.2010, 233, Mi. Halada ( BOL) ; Isparta, Karakus Dagi centr., 1460 m, 38°15'N 30°39'E, 11.07.2006, 13, M. Halada ( BOL) ; Karadut, Nemrut Dagi , 02.07.1993, 13, Mi. Halada ( BOL) ; Kopesiz , 25 km E Malatya, 08.07.1997, 13, Ma. Halada ( BOL) ; Kutahya 20 km NNE, 13.07.2006, 1♀, M. Halada ( BOL) ; Kuzucebelen , Mersin, 28.05.1998, 13, Ma. Halada ( BOL) ; Malatya 15 km E, 27.06.2000, 13, M. Halada ( BOL) ; Mugla, University Campus , stony step, 710 m, 37°09'39''N 28°22'20''E, 2013, 1♀, Anonym ( BOL) ; Nemrut Dag , Bitlis, 2000 m, 16.08.1991, 13, M. Halada ( BOL) ; Nemrut Dagi, Karadut , 02.07.1993, 333, Deneš ( BOL) ; Pasli , 50 km S Kars, 01.07.1997, 2♀♀, Ma. Halada ( BOL) ; Pazarcik , Kahramanmaras, 24.06.2000, 13, M. Halada ( BOL) ; Refahye 15 km W, W of Erzincan, 1600 m, 07.07.2000, 233, M. Halada ( BOL) ; Sahlili , Manisa 15 km SEE, 170 m, 38°26'N 28°19'E, 02.07.2006, 13, M. Halada ( BOL) ; Salihli , Manisa, 15 km SEE, 170 m, 38°26'N 28°19'E, 02.07.2006, 233, J. Halada ( BOL) ; Salihli , Manisa, 40 km NW, 150 m, 38°40'N 27°45'E, 28.06.2006, 1♀, J. Halada ( BOL) ; Silifke , 26.05.2001, 13, Snižek ( BOL) ; Tatvan 30 km E, 07.07.1997, 333, Ma. Halada ( BOL) ; Tatvan , Van G., 01.07.2000, 13, M. Halada ( BOL) ; Van 20 km W, 05.07.1997, 13, 3♀♀, Ma. Halada ( BOL). TURKMENISTAN: Annau , Aschabad 20 km E, 03.06.1993, 13, M. Halada ( BOL) ; Aschabad 15 km N, 25- 31.05.1993, 633, 1♀, M. Halada ( BOL) ; Aschabad env., 10.06.1993, 13, M. Halada ( BOL) ; Firyuza , 40 km W Aschabad, 06.06.1993, 833, M. Halada ( BOL) ; Kizl-Arvat 50 km S Chajagala, Kopet dag, 37°22'N 55°57'E, 17.05.1993, 13, M. Halada ( BOL). UZBEKISTAN: Circik 40 km vych. Taskent , 04.07.1981, 633, M. Kocourek ( BOL) ; Ddjizak 5 km W, 23.05.1994, 13, J. Halada ( BOL) ; Papngan 20 km NW Kokand, 41°2'N 70°6'E, 12.05.1994, 233, Ma. Halada ( BOL) .

D e s c r i p t i o n: 3. 8-10 mm. Head and mesosoma black; metasomal tergum I red with black basal spot, II red, III black with read basilateral spots; sterna predominantly black; some specimens from Syria with nearly complete black metasoma ( var. melanotica ). Clypeus and frons densely punctate. Middle clypeal lobe hardly protruding beyond lateral lobe, anterior margin straight (Fig. 5). Face with appressed white setae. Mandible black, simple. Gena smooth and shiny, with long white setae. Antennomere I on inner side with white setae, III about 4× as long as apically wide, about 1.2-1.3× as long as IV; V-VIII slightly concave basally, with maximal width apically (Fig. 14). Pronotal collar with oblique rugae. Mesoscutum densely punctate, with smooth and shiny posteromedian spot and short white setae. Mesoscutellum smooth and shiny, laterally and distally densely punctate. Mesopleuron densely punctate, with white setae. Dorsal surface of propodeum reticulate, glabrous; declivity and lateral surface rugose-punctate, with long white setae. Metasomal tergum I with white setae, with oblique microstriae; sterna with white setae. Hairbrush very short, lateral setae not longer than central ones, all together not longer than one third of those on sternum II (Fig. 25). Legs black, foretibia on inner side and all tarsi brown. Midcoxa on inner side concave, hindcoxa unmodified, trochanters and femora with white setae. Forewing with pale brownish tinge. Marginal cell about 3.5-4.0× as long as wide.

♀. 8-10 mm. Head, antenna and mesosoma black, metasomal segments I-III red; some specimens from Syria with nearly complete black metasoma ( var. melanotica ). Coxae, trochanters and hindfemur black, fore- and midfemur black with reddish tip, foretibia and -tarsus reddish, midtibia and -tarsus reddish brown, hindtibia and -tarsus brown. Face with white setae. Anterior margin of clypeal lamella slightly curved. Clypeus and face densely punctate, with smooth and shiny heart-shaped spot in front of anterior ocellus. Frons in front of anterior ocellus as well as surface between ocelli and eye shiny, with strong scattered punctures ( Fig. 88 ). Gena smooth and shiny, with isolated shallow punctures and white setae. Mandible black with reddish middle, lower margin simple. Antennomere III about 2.4-2.6× as long as apically wide, about 1.2-1.3× as long as IV. Pronotal collar rugose-punctate. Mesoscutum anteriorly densely punctate, with white setae and some dark bristles between it; posterior surface smooth and shiny, laterally with punctures. Mesoscutellum smooth and shiny, with punctures posterolaterally. Ventral surface of mesopleuron shiny with strong punctures, lateral surface with longitudinal striae. Propodeal dorsum glabrous, irregularly reticulate; declivity rugose-punctate; lateral surface anteriorly smooth, posteriorly with longitudinal rugae; with short white setae. Stigmal area of propodeum concave. Metasomal terga with oblique microstriae. Sternum II and posterior margins of following sterna with white setae. Pygidial plate narrow, lateral margin concave. Forebasitarsus with four rake spines, ventrally with nine spines evenly prolonged towards the tip (Fig. 79). Ventral surface of midbasitarsus with short spines basally and long ones distally. Forewing brownish tinged, apical margin with diffuse brown spot. Marginal cell about 3.0-3.5× as long as wide.

R e m a r k s: Specimens from Syria with nearly complete black metasoma are described by PULAWSKI (1973) as Astata stecki melanotica. All specimens examined here agree with the typical coloration. The black form is treated here as color variation.

D i s t r i b u t i o n: Europe: Germany, Poland ( OLSZEWSKI et al. 2021), Switzerland, Czechia, Slovakia, Austria ( DOLLFUSS 1987), Hungary, Portugal ( CROSS et al. 2021), Spain, France, Italy, Croatia, Bulgaria, Albania, Greece, Cyprus, Ukraine ( SHORENKO & KONOVALOV 2010), Crimea. Russia: European part, Western Siberia ( NEMKOV 2017), Russia: Eastern Siberia, Far East. Maghreb: Morocco, Tunisia. Middle East: Egypt, Armenia, Azerbaijan: Nakhichevan ( MOKROUSOV et al. 2019), Türkiye, Syria, Jordan, Iran. Central Asia: Kazakhstan, Turkmenistan, Kyrgyzstan, Uzbekistan, Tajikistan, Mongolia. South Asia: India Kashmir (type locality).

Figures

Figs 85-94. Figs 85-87: pygidial plate ♀: (85) A. diversipes, (86) A. rufipes, (87) A. apostata. Figs 88-89: frons ♀: (88) A. kashmirensis, (89) A. minor. Fig. 90: A. pontica ♀: vertex. Figs 91-92: A. quettae ♀: (91) head frontal, (92) forewing. Figs 93-94: head frontal ♀: (93) A. laeta, (94) A. radialis.