Astata costae A. COSTA , 1867

Hans-J, 2025, Astata LATREILLE, 1796 (Hymenoptera, Astatidae) from the Palearctic region with descriptions of nine new taxa and a key to species, Linzer biologische Beiträge 57 (1), pp. 71-168 : 102-104

Astata costae A. COSTA, 1867

Astata costae A. COSTA, 1867: 90 , ♀. Holotype ♀, Italy, Toscana, no specific locality (Napoli).

Astatus gracilis GUSSAKOVSKIJ, 1927: 210, 3. Lectotype 3, Crimea, Sebastopol (ZIN). Designated by PULAWSKI 1965: 571. Synonymized with Astata costae by BEAUMONT 1942: 409.

Astatus parvulus GUSSAKOVSKIJ, 1927: 292. Holotype ♀, Uzbekistan, Mayskoe near Tashkent (ZIN). Synonymized by PULAWSKI 1956: 50. New status as Astata costae parvula by BEAUMONT 1942: 409.

M a t e r i a l e x a m i n e d: ARMENIA: Shumuch 8 km SE, 10.05.2003, 13, Mučka (BOL). BULGARIA: Argutino, 07.1968, 13, A. Gortler ( BOL) ; Lozen , W of Svilengrad, 41°48'N 26°00'E, 140 m, 13.06.2017, 13, B. Halada ( BOL) ; Kiten , 42°14'N 27°46'E, 19.08.1990, 13, Burger ( SDEI) ; Knazevo , 15 km W Elchovo, 42°06'N 26°30'E, 19.06.2008, 233, M. & Z. Halada ( BOL) ; Lozenek , 15 m, 42°13'N 27°47'E, 16.06.2008, 13, M. & Z. Halada ( BOL) ; Melnik , 13.08.1993, 13, M. Halada ( BOL) ; Sandanski , 01- 17.07.1966, 13, 1♀, 06.1969 1♀, Kocourek ( BOL) ; Slancev Brjag , 20- 05.08.1965, 1♀, 25.07.1968, 233, 1♀, 03.08.1968, 1♀, 14.07- 14.08.1971, 533, 4♀♀, 10.06.1972, 1♀, Padr ( BOL) ; Varvara , 06- 12.08.1970, 13, Kocourek ( BOL) ; Zvilengrad 30 km SW, 550 m, 41°43'N 26°02'E, 21.06.2008, 233, M. & Z. Halada ( BOL) . CROATIA: Grbavac , 4 km E Dubrovnik, 42,652°N 18,176°E, 15.09.2018, 1♀, Schmid-Egger ( SE) ; Istria, 15 km SE Pula, Beach SE Litzjan , 44.810°N 13.987°E, 17.07.2021, 13, Schmid-Egger ( SE) ; Istria, Laborica , 20.07.2002, 13, Ma. Halada ( BOL) ; Rab isl., Lopar 3 km W, 22.08.2010, 13, 2♀♀, Halada ( BOL) . CYPRUS: Kiti , 26.08.2000, 13, Schlaefle (WS) ; Yermasoyia River , 18.09.1962, 1♀, ( BOL) . FRANCE: Corse, Algajola, Dünen , 01- 11.07.1979, 233, Wolf, det. Pulawski ( BOL) ; Grepiac , 24.06.2011, 13, Grouet (JA) ; Montredon-des-Corbieres , 09.06.2012, 13, 26.06.2013, 13, Grouet (JA) . GREECE: Greece , Peloponnese, 12 km S of Andritsena, N37.24' E21.50', 500 m, 28.08.2019, 333, J. Halada ( BOL) ; Leptokaria , 09.07.1993, 13, 16.07.1993, 13, Jacobs (JA) ; Peloponissos mer., Mt. Tayetos Oros centr., 28- 31.07.2010, 1♀, J. Halada ( BOL) ; Rhodos, Faliraki , 04.09.1984, 13, Schmid-Egger ( SE) ; Rhodos, Kolimbia , 11.09.1984, 13, Schmid-Egger ( SE) ; Rhodos, Kritinia , 11.09.1984, 13, Schmid-Egger ( SE) ; Santorin, Akrotiri, Red Beach , 09.05.1992, 13, Fiechtner & Leidenroth (JA) ; Zakynthos, Vassilikos , 30.09.2013, 3♀♀, Schlaefle (WS, JA) . HUNGARY: N of Varpalota , 47.212°N 18.130°E, 11.06.2020, 2♀♀, M. Halada ( BOL) ; Pakozd E of Szekesfehervar, 47.222°N 18.521°E, 12.06.2020, 233, M. Halada ( BOL) ; Örkeny 50 km SE Budapest, 05- 09.08.2004, 1♀, Ma. Halada ( BOL) ; Pakozd E of Szekesfehervar, 150 m, 47°13'N 18°31'E, 25.06.2015, 1♀, M. Halada ( BOL) . IRAN: Golestan, Azhad Shahr 5 km S, 300 m, 37°07'N 55°27'E, 11.06.2010, 13, Mi. Halada ( BOL) ; Kerman, Deh Bakri, Gebal Barez Mts. , 1640 m, 29°08'N 57°91'E, 03.06.2010, 13, Mi. Halada ( BOL) ; Golestan, Shakuh-E-Sofla, 2515 m, 23°32'34''N 54°25'10''E, 10.07.2018, 13, Liebig ( LI) ; Golestan, Ziarat , 2299 m, 36°40'12''N 54°33'27''E, 05.07.2018, 333, Liebig ( LI) ; Khorasan N, Bajgiran 27 km SO, 2194 m, Euphorbia , 37.41.57N 58.06.01E, 12.07.2022, 13, W.- H. Liebig ( LI) ; Teheran prov., Elburz, Serkheh , 2266 m, 35.7396°N 52.9925°E, 06.07.2022, 1♀, Schmid-Egger ( SE) ; Azer e Shabestar, Sis , 10 km E Shabestar, 38°26'N 45°86'E, 1540 m, 19.06.2010, 733, 15♀♀, Mi. Halada ( BOL) . ISRAEL: Mt. Hermon, Emeq Sal'am , 33.300°N, 35.786°E, 2040 m, 21.06.2019, 13, A. Dorchin ( BOL) . ITALY: Sardegna, Cagliari , 23.07.1959, 233, 2♀♀, Wolf ( BOL) ; Sicily, Castelbuno env,, Mt. Madonie , 600 m, 14.06.2012, 1♀, J. Halada ( BOL) ; Sicily, Castellamare env., 55 km E Trapani, 09- 12.06.2002, 13, 1♀ J. Halada ( BOL) ; Calabria, Lokri 15 km S, 02.08.1999, 13, J. Halada ( BOL) ; Sicily, Ragusa 35 km SW, beach, 18- 22.06.2002, 13, J. Halada ( BOL) ; Sicily, 55 km E Trapani, Castellammare env., 12.06.2002, 1♀, Halada ( SE), Sicily, Randazzo, Mt. Spagnolo , 1200 m, 06.06.1999, 1♀, Merz ( SE) ; Vieste, Valle Izillo , 44°47'27''N 16°6'37''E, 10.07.2013, 233, Kafka ( BOL) . JORDAN: Shuna north, 29- 30.04.1996, 13, Ma. Halada ( BOL) . KAZAKHSTAN: Dzungarian Alatau , Sharkent 26 km NW ,

44.3737N 79.7773E, 1184 m, 07.06.2024, 13, Schmid-Egger ( SE) ; Lepsi , 46°4'N 78°8'E, 19- 20.06.1995, 1♀, J. Halada ( BOL) ; Balchasch-Lake, Lepsy 22 km NW, 46.377N 78.724E, 289 m, 12.06.2024, 1♀, Schmid-Egger ( SE) ; Lepsi env., 400 m, 46°13'27''N 78°59'27''E, 02.06.2016, 1♀, J. Halada ( BOL) ; Malaysari 144 km N Alma-Ata, 25.06.1992, 233, M. Halada ( BOL). KYRGYZSTAN: Jalal-Abad, Ala-Buka 20 km SSW, 980-1100 m, 41°13'8''N 71°23'39''E, 28- 30.05.2008, 1♀, E. & J. Hüttinger ( BOL) ; Jalal-Abad, Ala-Buka, Umg. Kara-Uhkur u. Sovet-Sai , 41°13'48''N 71°23'39''E, 28- 30.05.2008, 1♀ ; E. & J. Hüttinger ( BOL) ; Alash Forest, Alash-Too Mts. , Ferghan. Mt. R., 08.2000, 13, Gurko ( BOL) ; Burgandi 60 km W Jalal-Abad, 14.06.1995, 13, J. Halada ( BOL) ; Naryn-Tal westl. Naryn , 1850 m, 41.22°N 75.36°E, 16.07.1999, 233, Jacobs (JA). MALTA: Dingli, 300 m, 35°85'N 14°40'E, 25.06.2014, 13, Kafka ( BOL) ; Gozo, Mgarr-ix-Xini, 16.06.1999, 1♀, Merz ( SE). MOROCCO: Ait Baha, Lake 2 km SE, 610 m, 30.051°N 9.108°W, 11.06.2014, 13, Schmid-Egger ( SE) ; Col du Kerdous, 15.06.2003, 13, Schlaefle (WS); Essaouria , 09.05.1995, 1♀, Mi. Halada ( BOL) ; Gorges du Tetra, 29.05.1990, 13, Schlaefle (WS); Guelmin 10 km E, 05.05.1995, 13, Mi. Halada ( BOL) ; Haddada , 65 m, 34.1397°N 6.5352°W, 17.07.2019, 233, El Abdouni &, L Hamroud ( SE) ; Rich 10 km N, 23.05.1995, 13, Mi. Halada ( BOL) ; Tamri , 70 km N Agadir, 08.05.1995, 1♀, Mi. Halada ( BOL) ; Tizi-n-Test S, 1900 m, 30.06.1987, 1♀, M. Schwarz ( BOL). OMAN: SW of Sar , 03- 05.03.2017, 13, M. Snižek ( BOL). ROMANIA: Dunabe Delta, Chilia Veche, 05.2019, 1♀, J. Votypka ( BOL). SLOVAKIA: Chotin, 22.07.1962, 1♀, Padr ( BOL) ; Gbelce , 14.08.1965, 1♀, Kocourek ( BOL) ; Somotor, 07.19??, 13, Kocourek ( BOL) ; Sturovo, 06.1960, 1♀, Kocourek ( BOL). SPAIN: Puigcerda, Schottergrube , 18.07.196,3 13, 1♀, Hamann ( BOL) ; Segovia, 28.06.1989, 1♀, Schlaefle (WS). SWITZERLAND: Wallis, Chandolin , 2150 m, 29.08.1984, 13, Wolf ( BOL) ; Wallis, Zeneggen, 1500 m, 08.2001, 13, 07.2006, 233, Schlaefle (WS, JA). SYRIA: Dibbin 30 km S Suwayda, 15- 17.05.1996, 13, Ma. Halada ( BOL) ; Kafr, Suwayda , 21.06.2000, 233, M. Halada ( BOL) ; Mishirfeh, Ar Raqqa, 04.06.2000, 13 1♀, Deneš ( BOL) ; Rankus 40 km N Damascus, 23.05.1996, 13, Ma. Halada ( BOL). TURKMENISTAN: Annau , Aschabad 20 km E, 03.06.1993, 1♀, M. Halada ( BOL) ; Vanovskij, Firjusa 3 km N, 13- 14.06.1988, 2♀♀, Lelej ( SE). TÜRKIYE: Agri env., 27.06.1993, 233, Mi. Halada ( BOL) ; Camardi , Nigde, 1800 m, 16.08.1991, 13, M. Halada ( BOL) ; Chamaldi-Sai , 41°2'N 71°8'E, 30.05.1995, 13, J. Halada ( BOL) ; Dalyan, Farm, Malaisetrap , 1 m, 36°48'54''N 28°39'04''E, 08- 20.08.2015, 1♀, Dursun ( BOL) ; Denizli, 10 km NE Denizli , N37.56' E29.07', 290 m, 04.07.2006, 13, M. Halada ( BOL) ; Dogubayazit N, Pr-Agri , 1700 m, 03.07.1985, 1♀, W. Schacht ( BOL) ; Ercis 10 km E, Van Golu , 25.06.1997, 13, Halada ( BOL) ; Erzurum 15 km SW, 39°48'N 41°10'E, 02.07.2001, 233, J. Straka ( BOL) ; Erzurum E of, 06.07.2000, 13, M. Halada ( BOL) ; Gevas, Van Gölü , 29.06.1993, 2♀♀, Mi. Halada ( BOL) ; Gözeldere 25 km E, 930 m, 37°32'N 43°49'E, 22.06.2010, 13, 2♀♀, Mi. Halada ( BOL) ; Gündogmus 7 km NE, W Giesandra , 30 km N Alanya, 1730 m, 36.86°N 32.03°E, 02.08.2009, 433, Schmid-Egger ( SE) ; Gurun 20 km E, Mezikiran Gecidi , 10.07.1997, 533, Ma. Halada ( BOL) ; Hakkari, Cemdinli , 1700 m, 20.07.1988, 13, Schmid-Egger ( SE) ; Hakkari, Esendere , 21.07.1988, 2♀♀, Schmid-Egger ( BOL) ; Hakkari, Stadtrand , 1800 m, 24.07.1988, 13, Schmid-Egger ( SE) ; Hakkari, Oramar , 1700 m, 29.06.1985, 1♀, M. Schwarz ( BOL) ; Hakkari, Suvari-Halil-Pass, E Beytüssebap, 2300 m, 03.08.1982, 1♀, Warncke (JA) ; Halfeti ( Birecik ), 31.05.1998, 233 1♀, Ma. Halada ( BOL) ; Ihlarn , Nigde, 09.08.1991, 13, Halada ( BOL) ; Kozluk ( Baykan ), 03.06.1998, 233, Ma. Halada ( BOL) ; Muradiye , 03.07.2000, 233, 05.07.2000, 233, M. Halada ( BOL) ; Muradye 10 km N, 27.08.1997, 13, Ma. Halada ( BOL) ; Nemrut Dag , 2000 m, 16.08.1991, 233, 2♀♀, M. Halada ( BOL) ; Nemrut Dag , Bitlis, 2000 m, 16.08.1991, 233, Ma. Halada ( BOL) ; Pamukkale , 08- 09.06.1964, 1♀, J. Gusenleitner ( BOL) ; Porsuk Baraji, Sebran , 08.07.1993, 13, Mi. Halada ( BOL) ; Salihli / Manisa, 15 km SEE, 170 m, 38°26'N 28°19'E, 02.07.2006, 13, 1♀, J. & M. Halada ( BOL) ; Side E, Dunes , 06.10.2011, 13, J. Halada ( BOL) ; Sille near Konya, 16.06.1968, 1♀, leg J. Schmidt ( BOL) ; Tatvan 10 km N, 24.06.1997, 233, Ma. Halada ( BOL) ; Tatvan 30 km E, 07.07.1997, 533, Ma. Halada ( BOL) ; Tatvan , Van G., 01.07.2000, 13, M. Halada ( BOL) ; Zelve (Nevsehir), 1100 m, 25.08.1991, 1♀, Halada ( BOL). UNITED ARAB EMIRATES: Abu Dhabi pr., New Shwaib , 24°44'N 55°47'E, 280 m, 08.03.2017, 2♀♀, M. & B. Halada ( BOL) ; Sarjan pr., W of Dhaid, 25°17'N 55°51'E, 120 m, 02.03.2017, 1♀, M. & B. Halada ( BOL). UZBEKISTAN: Aman-Kutan-Pass, W- Zeravshan MT. R., 1700 m, 39°18'N 66°54'E, 27.06.1999, 1♀, I. Makogonova ( BOL) ; Papngan 20 km NW Kokand, 41°2'N 70°6'E, 12.05.1994, 1♀, Ma. Halada ( BOL). YEMEN: Wadi Aniz SSW Sana, 14°60'N 44°09'E, 1520 m, 07.10.2005, 1♀, J. Halada ( BOL) .

D e s c r i p t i o n:J. 6-9 mm. Black, tergum I red with large black basal spot, II red, III with red proximal margin. Legs black, tarsi brownish. Middle clypeal lobe distinctly protruding beyond lateral lobe, apical margin straight. Clypeus and frons densely punctate. Face with white appressed setae. Mandible simple; black, sometimes red medially. Gena smooth and shiny, with shallow punctures and white setae. Antennomeres I-II with white setae; III about 5× as long as apically wide, about 1.3× as long as IV. Antennomeres V-IX strongly concave mesally, about 1.5-1.7× as long as maximal width (Fig. 55). Proximal tubercles of tyloids yellowish (Fig. 56). Pronotal collar and mesoscutum dull, very densely punctate. Mesoscutellum densely punctate with smooth and shiny spot anteriorly. Mesopleuron densely punctate, without striae. Propodeal dorsum strongly reticulate, sometimes with distinct oblique longitudinal carinae. Mesosoma except propodeal dorsum with erect white setae. Central setae of metasomal hairbrush dense and about a quarter as long as lateral ones, the latter about as long as those on sternum II. Terga dull, with oblique microstriae. Legs unmodified, with white setae. Forewing with light yellowish tinge, apical margin hyaline. Marginal cell about 3.5-4.0× as long as wide.

♀. 6-9 mm. Head and mesosoma black. Mandible reddish with black base and tip. Metasoma red, sometimes with reddish brown apical segments. Coxae and femora black, tibiae and tarsi reddish, mid- and hindtibiae often brownish darkened. Clypeal lamella distinctly protruding beyond lateral lobe, apical margin straight. Frons in front of anterior ocellus smooth and shiny, with fine scattered punctures, interspaces at least as wide as puncture diameter; between anterior ocellus and eye more scattered punctate, interspaces distinctly wider than puncture diameter ( Fig. 101 ). Vertex and gena smooth and shiny, with very isolated punctures. Antennomere III about 2.4-2.8× as long as apically wide, about 1.1-1.4× as long as IV. Apical antennomeres short, about 1.2-1.5× as long as maximal width ( Fig. 102 ). Mesoscutum smooth and shiny, anterior surface densely punctate, remaining part with isolated punctures. Mesoscutellum smooth and shiny. Mesopleuron strongly punctured, ventral surface nearly impunctate. Propodeal dorsum strongly reticulate, lateral surface with longitudinal carinae. Stigmal area of propodeum straight or slightly convex. Proximal metasomal terga with very fine oblique microstriae. Head, mesosoma (except propodeal dorsum) and legs with sparse and short white setae. Coxae and trochanters unmodified. Forebasitarsus short, with four rake spines; ventral surface with 8-10 undulating spines, fine setae between the spines covering half of tarsomere length. Midbasitarsus straight, ventrally with undulating spines. Forewing with light brownish tinge, apically more strongly. Marginal cell about 2.5-2.8× as long as wide.

D i s t r i b u t i o n: Europe: Switzerland, Slovakia, Hungary, Croatia, France, Portugal ( CROSS et al. 2021), Spain, Italy, Malta, Romania, Bulgaria, Greece, Cyprus, Ukraine ( SHORENKO & KONOVALOV 2010), Crimea ( SHORENKO 2017). Russia: European part, Western Siberia ( NEMKOV 2017). Maghreb: Morocco, Tunisia ( BEN KHEDHER et al. 2020). Middle East: Armenia, Azerbaijan: Nakhichevan ( MOKROUSOV et al. 2019), Türkiye, Israel, Syria, Jordan, Oman, Yemen, Iran. Central Asia: Kazakhstan, Turkmenistan, Kyrgyzstan, Uzbekistan, Tajikistan ( NAZAROVA 1998).

Figures

Figs 95-106. Figs 95-96: thorax dorsal ♀: (95) A. laeta, (96) A. radialis. Figs 97-98: forebasitarsus ♀: (97) A. rubra nov.sp., (98) A. mongolica nov.sp. Figs 99-101: frons ♀: (99) A. rubra nov.sp., (100) A. rubriventris, (101) A. costae. Figs 102-103: antenna ♀: (102) A. costae, (103) A. lusitanica. Figs 104-106: A. ljubomirovi ♀: (104) clypeus, (105) frons, (106) mesopleuron.