Eoecophylla quilchenensis, Archibald, Mathewes, and Perfilieva, 2024

Archibald, S. Bruce, Mathewes, Rolf W. & Perfilieva, Ksenia S., 2024, Fossil weaver ants (Hymenoptera, Formicidae, Oecophyllini) of the early Eocene Okanagan Highlands of far-western North America, The Canadian Entomologist (e 2) 156, pp. 1-16 : 5-10

publication ID	https://doi.org/10.4039/tce.2023.27
persistent identifier	https://treatment.plazi.org/id/0394A47F-0F48-7462-FF17-F9F9FB3CFA44
treatment provided by	Felipe
scientific name	Eoecophylla quilchenensis, Archibald, Mathewes, and Perfilieva
status	sp. nov.

Treatment

Eoecophylla quilchenensis, Archibald, Mathewes, and Perfilieva View in CoL , n. sp.

ZooBank Registration number: urn:lsid:zoobank.org:pub:BDF6655F-884D-4452-A8D8-24A338A67151

Figs. 1–5 View Figure 1 View Figure 2 View Figure 3 View Figure 4 View Figure 5 .

Material. All Quilchena insect fossils were collected by RWM, J. Mathewes, G. Guthrie, and Simon Fraser University undergraduate students over many years and are now transferred from the Simon Fraser University collection to the Beaty Biodiversity Museum.

Queens (type A of Archibald and Mathewes 2000): all preserved in dorsal aspect. Holotype: BBM-PAL-P000016 ( SFU number Q-0409AB; Figs. 1C–D View Figure 1 , 4A View Figure 4 ): body with a forewing and hind wing. Paratypes: BBM-PAL-P000017 (Q-0011; Fig. 1A–B View Figure 1 ): well-preserved body, no wings; BBM-PAL-P000019 (Q-0517; Figs. 2C–D View Figure 2 , 4D View Figure 4 ): much of the body, poorly preserved, well-preserved forewing with partial wing beneath it, other forewing disarticulated from the body. Other specimens: BBM-PAL-P000018 (Q-0006; Figs. 2A–B View Figure 2 , 4C View Figure 4 ): part of gaster, much of forewing, perhaps part of hind wing; BBM-PAL-P000020 (Q-0271; Figs. 3 View Figure 3 , 4B View Figure 4 ): poorly preserved body, two forewings, one rather complete; BBM-PAL-P000021 (Q-0001; Fig. 4E–F View Figure 4 ): forewing, mostly complete; BBM-PAL-P000022 (Q-0412; Fig. 4G–H View Figure 4 ): part of forewing, missing distal portion; BBM-PAL-P000023 (Q-0457; Fig. 4I–J View Figure 4 ): much of forewing missing distal portion. BBM-PAL- P000024 (Q-0492; Fig. 4K–L View Figure 4 ): partial forewing .

Male (type B of Archibald and Mathewes 2000). Allotype: BBM-PAL-P000025 (Q-0007; Fig. 5E–F View Figure 5 ): almost complete, wings with some venation preserved. Paratypes: BBM-PAL-P000026 (Q-0008; Fig. 5A–B View Figure 5 ): almost complete, well preserved, but no wings; BBM-PAL-P000027 (Q-0009; Fig. 5C–D View Figure 5 ): rather well preserved, but no wings, almost no legs. Other specimens: BBM-PAL- P000028 (Q-0013; Fig. 5G View Figure 5 ): mostly poorly preserved, parts of four faint wings; BBM-PAL- P000029 (Q-0258; Fig. 5H View Figure 5 ): body, some leg parts, no wings; BBM-PAL-P000030 (Q-0456; Fig. 5I View Figure 5 ): body, most legs, no wings; BBM-PAL-P000031 (Q-0485; Fig. 5J View Figure 5 ): poorly preserved body, no wings. The following are not figured. BBM-PAL-P000032 (Q-0002): complete but with poorly preserved wings; BBM-PAL-P000033 (Q-0010): partially preserved, no wings, no head, some legs; BBM-PAL-P000034 (Q-0012): poorly preserved, faint wings; BBM-PAL-P000035 (Q-0014): poorly preserved body with parts of two legs, no wings; BBM-PAL-P000036 (Q-0019): poorly preserved body, no legs, faint wing(s?); BBM-PAL-P000037 (Q-0021): well-preserved body, no wings; BBM-PAL-P000038 (Q-0366): part of body, leg bits, no wings; BBM-PAL-P000039 (Q-0410): poorly preserved body, no wings; BBM-PAL-P000040 (Q-0453): body, some legs, no wings; BBM-PAL- P000041 (Q-0510): poorly preserved body, no legs, no wings; BBM-PAL-P000042 (RS-160):

poorly preserved body, no wings; BBM-PAL-P000043 (RS-281): poorly preserved body, some leg parts, no wings.

Diagnosis. As for the genus, above.

Description. Holotype queen BBM-PAL-P000016: preserved in dorsal aspect ( Figs. 1C–E View Figure 1 , 4A View Figure 4 ). BL ca. 14.5. Head: HL ca. 1.7, HW ca. 2.0; mandibles large, MdL ca. 1.0, subtriangular, masticatory margin indistinctly preserved; eyes, antennae poorly preserved; occipital corners well developed, rounded. Alitrunk indistinctly preserved. Forewing: FwL ca. 20.0; FwW indistinct, ca. 5.5; 1M, 1Rs nearly aligned; pterostigma long, narrow, similar lengths basad, distad crossvein 2r-rs; 2r-rs angled, anterior basad posterior; 5Rs curved towards anterior margin; stub of m-cu present, extending almost halfway between Rs M, Cu; Rs M distinctly bent at stub; 1Rs M, 2Rs M distinctly angled; 2Rs M, 2r-rs, 5Rs, 4M form a ×; fragment of cu-a preserved on left forewing, better-preserved on BBM-PAL-P000018 forewing, see Fig. 2B View Figure 2 . Hind wing: 1M, 2M free, 2M well developed, rs-m curved, portions of 2M Cu, R, 2Rs preserved, A not preserved, nor region where jugal lobe might be present. Waist of one segment; petiole poorly preserved. Legs: fragments, poorly preserved. Gaster: indistinct, without constriction between AIII, AIV (first two segments); GL ca. 5.4; GW ca. 5.0.

Paratype queen BBM-PAL-P000019: preserved in dorsal aspect ( Figs. 2C–D View Figure 2 , 4D View Figure 4 ). BL ca. 19.0. Head, alitrunk indistinctly preserved but with subtriangular mandibles bearing probably 9? or more teeth. Forewing: FwL ca. 19.0 preserved, missing apical portion; FwW ca. 5.5; 1M, 1 Rs aligned; pterostigma long, narrow, apparently similar lengths basad, distad crossvein 2r-rs; 2r-rs angled, anterior basad posterior; 5 Rs curved towards anterior margin; stub of m-cu present, extends less than halfway between Rs M, Cu; 1 Rs M, 2 Rs M distinctly angled; 2 Rs M, 2r-rs, 5 Rs, 4M meet to form an ×. Hind wings absent. Legs: Fragment present, mesofemur, metafemur ca. 3.5 mm long. Waist of one segment; petiole indistinct. Without constriction between AIII, AIV. Gaster.:GL ca. 8.0 as preserved; GW ca. 5.5 .

Paratype queen BBM-PAL-P000017: preserved in dorsal aspect ( Fig. 1A–B View Figure 1 ). BL 16.1. Head: with mandibles subtriangular, otherwise indistinct; HL MdL 2.9, HW 2.6; occipital corners well developed, rounded. Alitrunk: AL 5.3. Wings: absent. Legs: fragments present. Petiole: PtL ca. 1.6. Gaster: GL 6.7, GW 5.8.

Allotype male BBM-PAL-P000025: preserved in dorsal aspect ( Fig. 5E–F View Figure 5 ). BL 11.2. Head: square with rounded corners; HL 1.5; HW 1.7; mandibles poorly preserved; eyes indistinct. Alitrunk: AL 4.9, AW ca. 2.6. Forewing: mostly poorly preserved, 2Rs M, 2r-rs, 5Rs, 4M meet to form ×; 5Rs curved towards anterior margin. Hind wing: not known. Legs: long. Waist: single segmented; petiole without scale. Gaster: GL ca. 4.1, GW 3.5, large, semi-spherical without constriction between first two segments.

Paratype male BBM-PAL-P000026: preserved in dorsal aspect ( Fig. 5A–B View Figure 5 ). BL 10.4. Head: HL 1.4, HW 1.4; occipital corners well developed, rounded; MdL 0.6. Alitrunk: AL 4.3. Legs: long, metafemur ca. 3.3, metatibia ca. 3.0. Waist single-segmented, petiole without scale, PtL 1.1, PtW 0.8. Gaster: GL 4.0, GW 3.1.

Paratype male BBM-PAL-P000027: head in dorsolateral, alitrunk in lateral, petiole, gaster in dorsal aspect ( Fig. 5C–D View Figure 5 ). Alitrunk: AL 4.0, AH 3.0. Waist: single-segmented; petiole without scale, PtL 1.1, PtW 0.8. Gaster: GL 4.0, GW 3.5.

Etymology. The specific epithet is a toponym referring to the species’ only known locality.