Aname truncata, Wilson & Harvey & Simmons & Rix, 2025

Wilson, Jeremy D., Harvey, Mark S., Simmons, Leigh W. & Rix, Michael G., 2025, An integrative systematic revision of the wishbone spiders (Araneae: Anamidae: Aname L. Koch, 1873) of subtropical and tropical eastern Australia, with the description of 55 new species, European Journal of Taxonomy 985, pp. 1-298 : 83-85

Aname truncata sp. nov.

urn:lsid:zoobank.org:act:D2026664-38CE-4036-9196-8F93B8300690

Figs 1 , 8 , 55

Diagnosis

Males of A. truncata sp. nov. are unknown.

Females of A. truncata sp. nov. can be distinguished from all species for which females are known except A. barakula sp. nov., A. braemar sp. nov., and A. lambkinae sp. nov. by the presence of spermathecae with two vesicles, with relatively long and straight lateral vesicles (lateral vesicle length/genitalia width> 0.25) and medial vesicles with few bends (usually one or two), and a narrow sternum (sternum length / width>1.3) ( Fig. 55A–L ). Females of A. truncata can be distinguished from those of A. barakula and A. braemar by the presence of spermathecae with lateral vesicles without distinct, widened crowns ( Fig. 55L ; cf. Figs 51 ). Females of A. truncata can be distinguished from those of A. lambkinae by the presence of spermathecae with less elongate lateral vesicles (lateral vesicle length/ width ~1.6; cf. ~ 3.6 in A. lambkinae ) and straighter medial vesicles without distinct crowns ( Fig. 55L ; cf. Fig. 54 ).

Etymology

The specific epithet ‘ truncata ’ is a Latin adjective meaning ‘truncated’ or ‘cut-off’, in reference to the wide, truncate abdomen of this species relative to others.

Type material

Holotype

AUSTRALIA – Queensland • ♀; Isla Gorge National Park , off Isla Gorge Road; 25°12′ S, 149°59′ E; 395 m a.s.l.; 15 Apr. 2023; J.D. Wilson and M.G. Rix leg.; excavated, open burrow on ground; QMB S118253 .

Paratype

AUSTRALIA – Queensland • 1 ♀; Isla Gorge National Park , off Isla Gorge Road; 25°12′ S, 149°59′ E; 399 m a.s.l.; 15 Apr. 2023; J.D. Wilson and M.G. Rix leg.; excavated, open burrow on ground; QMB S118254 .

Other material examined

AUSTRALIA – Queensland • 1 juv.; Taroom-Bauhinia Downs Road, near Palmgrove National Park ; 24°57′ S, 149°31′ E; 409 m a.s.l.; 15 Apr. 2023; J.D. Wilson and M.G. Rix leg.; excavated, open burrow on ground; QMB S118260 .

Description

Female (holotype, QMB S118253)

GENERAL ( Fig. 55A–L ). Body length 23.37, in good condition.

DORSAL PROSOMA ( Fig. 55A, E–F ). Carapace length 7.96, width 6.13, length/width 1.30, clypeus to fovea length/carapace length 0.66, caput width/carapace width 0.85, carapace red-brown, reflective setae present, heavy on caput, heavy on thorax, fovea procurved, fovea width/carapace length 0.13 ( Fig. 55A, F ); chelicerae dark red-brown, rastellum absent or inconspicuous, chelicerae length/carapace length 0.53 ( Fig. 55A ); eye group rectangular, width/length 2.03, eye tubercle present ( Fig. 55E ).

ABDOMEN ( Fig. 55B, D ). Abdomen length 11.13, dark brown, dorsal pattern absent, with consistent cover of short setae.

VENTRAL PROSOMA ( Fig. 55C, G–I ). Labium cuspules absent ( Fig. 55H ); maxillae heel distinct, cuspules present, count =about 90, extending posteriorly onto heel, extending laterally about 30% of maxillae length ( Fig. 55C, I ); coxae cuspules absent, thorn-like setae on prolateral face present ( Fig. 55C, I ); sternum length/width 1.46, central sternum with consistent covering of moderate setae, row of longer setae around posterior edges, some shorter thorn-like setae around anterior edges ( Fig. 55G–H ); posterior sigilla ovoid, central sternum to posterior sigilla length/sternum length 0.24, posterior sigilla length/ sternum length 0.13 ( Fig. 55G–H ); other sigilla small, round and lateral ( Fig. 55G–H ).

LEG I ( Fig. 55J–K ). Leg I dark red-brown, femur length 5.63, patella length 3.65, tibia length 3.96, metatarsus length 3.63, tarsus length 2.43, total length 19.31, leg I length/carapace length 2.43; scopulae on distal metatarsus and tarsus; spine count Fe D 1, Fe PL 1, Pa PL 2, Ti PL 0, Ti RL 4, Me PL 2, Me RL 3, Ta 0; tibia length/width [TIL/TID] 3.03.

GENITALIA ( Fig. 55D, L ). Epigastric furrow unmodified ( Fig. 55D ); spermathecae with two vesicles each ( Fig. 55L ); lateral vesicle relatively straight, length 0.42, lateral vesicle length/genitalia width 0.30, length/width at base 1.61, crown un-demarcated ( Fig. 55L ); medial vesicle short, projecting medially and curving from medial to lateral angle, medial vesicle length/genitalia width 0.14, length/width 1.67, medial vesicle length/lateral vesicle length 0.45 ( Fig. 55L ).

Distribution and natural history

Aname truncata sp. nov. occurs in central Queensland, in the Brigalow Belt South bioregion. It is known from two locations, both located in or near Palmgrove and Isla Gorge National Parks ( Fig. 8 ). It constructs an open, silk-lined burrow with silk spilling out from the entrance, usually with a slightly built up ‘collar’ of soil around the entrance, and with a hidden secondary ‘wishbone’ entrance ( Fig. 8 ).

Figures

Fig. 1. Phylogeny and general map of eastern Australian Aname L. Koch, 1873. Left: IQ-TREE Maximum Likelihood phylogeny for the genus Aname with species groups sensu Rix et al. (2021) highlighted in grey, species complexes as defined in this study highlighted in colour, and showing the phylogenetic position of many of the species described herein, as well as some potential species (those with nicknames in quotation marks) that may turn out to be distinct, but did not meet our standards for description. Species groups that are not represented in eastern Australia are collapsed. Support values on the phylogeny show the results of 1000 ultrafast bootstrap replicates, and are only shown in cases where support was below 90%. Top-right: a map of subtropical and tropical eastern Australia, showing the locality of all Aname specimens included in this study. Bottom-right: a smaller phylogeny highlighting the higher-level relationships between species groups of Aname as recovered in this study.

Fig. 8. Aname barakula-complex species habitus images, distribution map, and burrow architecture. Points on the map represented by a star are type localities. Note that intraspecific variation occurs in both physical appearance and burrow architecture, and images shown here should be treated only as examples, not used for species diagnosis.

Fig. 55. Aname truncata sp. nov., holotype, ♀ (QMB S118253). A. Cephalothorax, dorsal view. B. Abdomen, dorsal view. C. Cephalothorax, ventral view. D. Abdomen, ventral view. E. Ocular region. F. Fovea. G. Sternal sigilla, left view. H. Sternum and labium. I. Maxilla and coxae I–II, left view.J. Left leg I, prolateral view. K. Left leg I, retrolateral view. L. Internal genitalia, dorsal view. Scale bars: A–B, J =2 mm; L=1 mm.

Fig. 51. Aname barakula sp. nov., paratype, ♀ (QMB S118249). A. Cephalothorax, dorsal view. B. Abdomen, dorsal view. C. Cephalothorax, ventral view. D. Abdomen, ventral view. E. Ocular region. F. Fovea. G. Sternal sigilla, left view. H. Sternum and labium. I. Maxilla and coxae I–II, left view.J. Left leg I, prolateral view. K. Left leg I, retrolateral view. L. Internal genitalia, dorsal view. Scale bars: A–B, J =2 mm; L=1 mm.

Fig. 54. Aname lambkinae sp. nov., ♀ (QMB S118233). A. Cephalothorax, dorsal view. B. Abdomen, dorsal view. C. Cephalothorax, ventral view. D. Abdomen, ventral view. E. Ocular region. F. Fovea. G. Sternal sigilla, left view. H. Sternum and labium. I. Maxilla and coxae I–II, left view. J. Left leg I, prolateral view. K. Left leg I, retrolateral view. L. Internal genitalia, dorsal view. Scale bars: A–B, J =2 mm; L=1 mm.