Harmatia tokodensis, Kovács & Vicián, 2019

Kovács, Zoltán & Vicián, Zoltán, 2019, Muricidae (Neogastropoda) from the Middle Eocene of the Hungarian Paleogene Basin, Fragmenta Palaeontologica Hungarica 36, pp. 31-51 : 43

Harmatia tokodensis n. sp.

( Fig. 25 )

2020 Harmatia sp. – DULAI, p. 202, fig. G.

Holotype – HNHM M.62.7135, SL 72 (P1 spine not included) (Collected by Endre Szőts in 1956).

Type strata and locality – Upper Lutetian–Lower Bartonian grey marl, Tokod, Hungary.

Derivation of name – The name refers to the type locality.

Material – Holotype.

Diagnosis – Trivaricate Harmatia species with rounded teleoconch whorls, ovate aperture, long siphonal canal, narrow primary spiral cords, and long primary cord spines.

Description – Medium-sized shell, protoconch missing, subfusiform teleoconch. Spire of three shouldered, rounded whorls. Last whorl 89% of the total length of the preserved shell (without P1 spine). Aperture ovate, internal part not seen. Siphonal canal without cord spines, long, narrow and slightly curved, penultimate siphonal canal present. Spiral sculpture of moderately developed, narrow primary cords (P1–P4 visible on last whorl), axial sculpture of three spiny varices per whorl without intervarical ribs. Well-developed, long and open P1–P6 spines on last whorl, P1 spine slightly adapically curved, no intervarical ribs.

Remarks – The morphology of the new species corresponds to the diagnosis of genus Harmatia (see MERLE et al. 2011). The genus has been known from the Early Oligocene in Hungary and Austria with two species; the Middle Eocene occurrence of H. tokodensis n. sp. extends the stratigraphic range. Both H. stephani ( NOSZKY 1940, pl. 2, fig. 4; refigured by MERLE et al. 2011, text-fig. 63/1), and H. guembeli longispina ( NOSZKY 1940, pl. 1, fig. 31; refigured by MERLE et al. 2011, text-fig. 63/2) (Kiscellian, Budapest Újlak) are distinguishable by much lower spire, and H. stephani bears almost straight siphonal canal. The spire and sculpture of H. guembeli guembeli (Dreger) (Kiscellian, Bad Häring, Budapest Újlak) resemble that of H. tokodensis n. sp., the former, however, differs by strongly curved long siphonal canal. This feature cannot be seen on the fragmentary type specimen ( DREGER 1892, pl. 4, fig. 5; refigured by LÖFFLER 1999, pl. 12, fig. 1, and MERLE et al. 2011, text-fig. 63/3), but clearly visible on other specimens illustrated by LÖFFLER (1999, pl. 12, figs 2–5).

Figures

Fig. 25. Harmatia tokodensis n. sp., HNHM M.62.7135., holotype, Tokod, SL 72, apertural view (Photo: M. Szabó). Scale bar: 10 mm