Cicindela (s. str.) coerulea, Pallas, 1773

Matalin, Andrey V., 2025, An intraspecific structure of Сicindela (s. str.) coerulea Pallas, 1773 (Coleoptera, Cicindelidae), Zootaxa 5583 (2), pp. 201-246 : 238

Identification key to the subspecies of Cicindela (s. str.) coerulea

1(2) White elytral pattern undeveloped or strongly reduced ( Figs 24–41 ). 12.5–16.0 mm...... C. coerulea coerulea Pallas, 1773

2(1) White elytral pattern well developed ( Figs 89–106 , 110–116 , 155–166 , 178–189 , 215–232 ).......................... 3

3(4) White elytral pattern narrow with unconnected marks, humeral lunula divided on two isolated spots, rarely with virtually invisible medial portion, middle band in most specimens with short or long thin oblique branch without rounded apical extension, rarely with slightly developed it, marginal band undeveloped, rarely only marked ( Figs 178–189 ). Aedeagus with relatively long basal portion ( Figs 55, 56 ). 13.5–16.1 mm ...................... C. coerulea shantungensis Mandl, 1970

4(3) White elytral pattern medium-sized or wide with isolated marks, rarely with connected of some marks or with coupling all of them. Aedeagus with relatively short basal portion ( Figs 42–54, 57, 59–64 )....................................... 5

5(6) Medial band virtually transverse or slightly curved with very short oblique branch or without it ( Figs 215–232 ).......... 9

6(5) Medial band distinctly sinuate with well-developed oblique branch ( Figs 89–106 , 112–116 , 155–166 ).................. 7

7(8) White elytral pattern medium-sized, in most specimens with isolated marks ( Figs 89–106 ), rarely with coupling middle band and apical lunula ( Figs 112–114 ) or all marks connected by marginal band ( Figs 115, 116 ), extremely rarely with undeveloped ( Figs 110, 111 ) or clearly reduced ( Figs 107–109 ) marks. Internal sac long and sharply rounded towards to ventral side so В distinctly or virtually contact with the aedeagus tube ( Figs 117–132 ). 12.0– 15.4 mm ... C. coerulea nitida Lichtenstein, 1796

8(7) White elytral pattern often wide and all marks connected by marginal band ( Figs 83 , 155–159, 161, 163 ); in some specimens humeral lunula virtually separated from other marks ( Figs 160, 164–166 ); rarely both humeral and apical lunulae unconnected with relatively long marginal band ( Fig. 162 ). Internal sac short so В does not contact the aedeagus tube ( Figs 167–170 ). 12.1–14.5 mm ............................................................... C. coerulea kaszabi Mandl, 1965

9(10) Labrum shorter, LW/LL = 2.3–2.44. Pronotum narrower, PW / PL = 1.36–1.4, with slightly rounded subparallel lateral sides ( Figs 203–206 ). Middle band narrowly connected with apical lunula ( Figs 215–220 ). 12.2–13.6 mm ...................................................................................... C. coerulea przewalskii Dokhtouroff, 1887

10(9) Labrum longer, LW/LL = 2.0–2.33. Pronotum wider, PW / PL = 1.41–1.57, with virtually straight, convergent at its base with the sides ( Figs 207–214 ). All elytral spots separated from each other ( Figs 221–232 ). 12.1–15.0 mm......................................................................................... C. coerulea grumi Tschitschérine, 1903

Figures

FIGURES 24–41. Cicindela coerulea coerulea, left elytron: 24—Chernoyarka; 25—Maraldy-Romanovka; 26, 38, 39—Zhana- Arka; 27—Meret; 28—Irtysh; 29—Chernyi Yar; 30, 35–37, 40, 41—Biysk; 31—Lokot; 32—Kuludzhanskie sands; 33—Zaysan Lake, Kaznakovka; 34—Ryzaevskiy; 28—neotype; 24, 25, 28, 31–33, 36, 37, 39, 41—males; 26, 27, 29, 30, 34, 35, 38, 40— females.

FIGURES 89–106. Cicindela coerulea nitida, left elytron, typical pattern: 89—Hanka Lake; 90—Wenchuan (image by Keyi Wang); 91, 94, 100, 104, 106—Minusinsk; 92—Olekminsk; 93—Muren; 95—Mandakh; 96, 101—Luna; 97—Changai; 98— Jiulian Mt.; 99—central Yakutia; 102, 103—Mondy; 105—Nalaikh; 89, 91, 94–96, 100–103, 105, 106—males; 90, 92, 93, 97–99, 104—females.

FIGURES 107–116. Cicindela coerulea nitida, left elytron, atypical pattern: 107—Altai, Seminsky Mnt. pass; 108—Chita; 109, 110—Ongudai; 111—Amur; 112–114—Transbaikalia, Selenga fl. (images by Matthias Seidel and Helena Shaverdo); 115, 116—Mandal Gobi; 112–114—paratypes of C. coerulea selengensis; 107, 108, 111, 113, 16—males; 109, 110, 112, 114, 115— females.

FIGURES 155–166. Cicindela coerulea kaszabi, left elytron: 155—60 km W v. Somon Bulgan; 156—Bayan Dalay; 157— Beger-Nur; 158—Tsagan-Gol River; 159–162—Orok-Nuur; 163, 164—Barkull; 165, 166—Karlyk-Tag; 155—holotype; 155, 156, 158, 163, 165, 166—males; 157, 159–162, 164—females.

FIGURES 178–189. Cicindela coerulea shantungensis, left elytron: 178—Kiautschou (image by Matthias Seidel and Helena Shaverdo); 179—Peking; 180, 181—canyon of Moudjik (= Si-He) River; 182—Pin-Faun-Cha sands; 183—Can zhou (= Cangzhou); 184, 185—Yangmadianzi (images by Xinxin Xiong); 186—Shan-si; 187–189—Xiaojiagang (images by Xinxin Xiong); 178–paratype; 179, 180, 182–185, 187, 188—males; 178, 181, 186, 189—females.

FIGURES 215–232. Cicindela coerulea ssp., left elytron: 215–220—C. coerulea przewalskii, Amdo; 221–232—C. coerulea grumi (221–226—р. БадО-хО и ХЫ-хО-дО; 227–230—C. coerulea kozlovi, ДабасуН ГОби; 231, 232—Qinghai, Wulan); 215, 227—lectotypes; 216–226, 228–230—paralectotypes; 215, 216, 222, 225–229, 231—males; 217–221, 223, 224, 230, 232— females.

FIGURES 42–64. Cicindela coerulea ssp, aedeagus, left lateral view: 42–44—C. coerulea coerulea (42—Irtysh; 43— Kuludzhanskie sands; 44— Chernoyarka); 45–54—C. coerulea nitida (45—Nertshinsk; 46—Tere-Khol Lake; 47—Minusinsk; 48—Buir-Nuur; 49—Arkhara; 50—Yakutsk; 51—Sergeevka; 52—Nalaikh; 53—Jiulian Mt. (image by Keyi Wang); 54— Mandal Cobi); 55, 56—C. coerulea shantungensis (55—Peking; 56—Pin-Faun-Cha sands); 57–61—C. coerulea kaszabi (57, 58—60 km W v. Somon Bulgan; 59—60 km WNW of Bayan-Dalay; 60, 61—Barkull); 62—C. coerulea przewalskii, Amdo; 63—C. coerulea grumi, р. БадО-хО и ХЫ-хО-дО; 64—C. coerulea kozlovi, ДабасуН ГОби; 58—internal sack (without scale); 42—neotype; 45, 62—lectotypes; 57, 58—holotype; 63, 64—paralectotypes.

FIGURES 117–124. Cicindela coerulea nitida, internal sack, right lateral view: 117—Khodyn Lake; 118—Abakan; 119— Mondy; 120—Olekminsk; 121—Dzabyn-Gutu-Khudzhirt; 122—Natalino; 123—Blagoveshchensk; 124—Innokentievka; without scales. The abbreviations of humps and sclerites see in the text.

FIGURES 125–132. Cicindela coerulea nitida, internal sack, left lateral view: 125—Khodyn Lake; 126—Abakan; 127— Mondy; 128—Olekminsk; 129—Dzabyn-Gutu-Khudzhirt; 130—Natalino; 131—Blagoveshchensk; 132—Innokentievka; without scales. The abbreviations of humps and sclerites see in the text.

FIGURES 76–88. Cicindela coerulea spp., habitus, dorsal view: 76–82—C. coerulea nitida (76, 77—Nertshinsk (images by Jörg Gebert); 78—Khodyn Lake; 79—Kiran; 80, 81—Selenga fl. (images by Matthias Seidel and Helena Shaverdo); 82— Chita); 83–88—C. coerulea kaszabi (83, 84—60 km W v. Somon Bulgan; 85— Bayan Dalay; 86—Gulin-Tal; 87—Beger-Nur; 88—Tsagan-Gol River); 76, 77—lectotype of C. optata; 80, 81—paratype of C. coerulea selengensis; 83, 84—holotype; 77, 81, 84—labels; 78, 80, 83, 86, 88—males; 76, 79, 82, 85, 87—females.

FIGURES 167–170. Cicindela coerulea kaszabi, internal sack: 167, 169—Gulin-Tal; 168, 170—Barkull; 167, 168—right lateral view; 169, 170—left lateral view; without scales. The abbreviations of humps and sclerites see in the text.

FIGURES 203–214. Cicindela coerulea ssp., pronotum: 203–206—C. coerulea przewalskii, Amdo; 207–214—C. coerulea grumi (207–210—р. БадО-хО и ХЫ-хО-дО; 211, 213—C. coerulea kozlovi, ДабасуН ГОби; 212, 214—Qinghai, Wulan); 203, 211—lectotypes; 204–210, 213—paralectotypes; 203, 204, 207, 208, 211, 212—males; 205, 206, 209, 210, 213, 214— females.