taxonID	type	description	language	source
03C687D9FFC2FFF4A5C6FF60823AE71E.taxon	diagnosis	Diagnosis (modified from Takano et al. 2021). Fore tarsus with heavily sclerotized peg-like setae in both sexes (ch. 35 - 1). Sclerotized spermathecal capsule absent (ch. 89 - 1).	en	Zhang, Guang, Gao, Jian-Jun, Takano, Kohei Takenaka, Yafuso, Masako, Suwito, Awit, Meleng, Paulus Ak, Toda, Masanori J. (2023): Phylogenetic classification and palm-inflorescence anthophily of the Colocasiomyia zeylanica species group (Diptera: Drosophilidae), with descriptions of five new species. Zootaxa 5278 (2): 201-238, DOI: 10.11646/zootaxa.5278.2.1, URL: http://dx.doi.org/10.11646/zootaxa.5278.2.1
03C687D9FFC2FFF4A5C6FDCB8415E2A5.taxon	diagnosis	Diagnosis (modified from Fartyal et al. 2013). Pedicel with uniformly short, stout setae only, lacking prominently long one (s) (ch. 2 - 1; Fig. 3, “ Fig. 6 ” in Grimaldi 1991). Two peg-like setae present only on fore tarsomere II (ch. 36 - 0, ch. 38 - 0; Fig. 9 A – E 1, “ Fig. 1 C ” in Okada 1986, “ Fig. 9 ” in Grimaldi 1991). Hypandrium forming like tube surrounding phallus (ch. 66 - 2; Fig. 11). Gonocoxites absent (ch. 69 - 4; Fig. 11). Shared characters. Distance between antennal scapes wider than half of scape diameter (ch. 1 - 1; Fig. 2 A – E 2, “ Fig. 1 A ” in Okada 1986, “ Fig. 6 ” in Grimaldi 1991). Antennal, first flagellomere grayish yellow, with small invaginated organ but no hollow organ (ch. 3 - 0; Fig. 3, “ Fig. 6 ” in Grimaldi 1991); arista entirely densely pubescent, without branches (ch. 5 - 1; Fig. 3, “ Fig. 1 A, B ” in Okada 1986, “ Fig. 6 ” in Grimaldi 1991). Eye dark red (Fig. 14). Frons blackish brown; frontal vittae mat. Supracervical setae distally more or less curved, longer than inner, occipital setae (ch. 11 - 0, ch. 12 - 0; Fig. 4). Palpus pale grayish yellow, flat or convex in ventrodistal portion (ch. 16 - 0; Fig. 5, “ Fig. 1 B ” in Okada 1986, “ Fig. 7 ” in Grimaldi 1991). All cibarial sensilla very small or minute; anterior sensilla irregularly arranged in square or triangle (ch. 18 - 1, ch. 19 - 1; Fig. 5 A – E 2, “ Fig. 7 ” in Grimaldi 1991). Proboscis without a pair of lateral, spherical bumps covered with numerous, short, stout setae (ch. 22 - 0; Fig. 5, “ Fig. 7 ” in Grimaldi 1991). Additional, presutural, dorsocentral setae absent (Fig. 6, “ Fig. 1 A ” in Okada 1986). Basal and apical, scutellar setae nearly parallel (Fig. 6). Halter grayish brown. Wing hyaline; veins yellow (Fig. 8). Fore tarsomere II slightly elongated below (ch. 34 - 1; Fig. 9 A – E 1, “ Fig. 1 C ” in Okada 1986, “ Fig. 9 ” in Grimaldi 1991). Preapical setae absent on all tibiae (ch. 41 - 1; Fig. 9, “ Fig. 1 A ” in Okada 1986). Dorsomedian, terminal, down-curved seta on tarsomere Vs not longer than claw (ch. 42 - 0; Fig. 9). Mid tibia apicoventrally with 2 (1 long and 1 small) stout setae (ch. 39 - 0; Fig. 9 A – E 2). Spiracles simple, small (ch. 44 - 0). Male sternite VI pubescent entirely, without process (ch. 48 - 0, ch. 50 - 0; Fig. 10 A – E 1). Epandrium anteroventrally much elongated, not fused to cercus, without peg-like, stout seta at posteroventral apex; apodeme narrow lobe on anteromedial to - dorsal margin or absent (ch. 55 - 1, ch. 57 - 0, ch. 58 - 0, ch. 62 - 0; Figs 11 - 1 A – C 1,2, 11 - 2 D, E). Hypandrial setae absent (ch. 67 - 1; Figs 11 - 1 A – C 3,4, 11 - 2 D, E). Postgonites fused to aedeagal sheath, forming together phallal-sheath tube distally more or less separated into a pair of lateral lobes (ch. 71 - 1, ch. 72 - 1, ch. 73 - 0; Figs 11 - 1 A – C 3,4, 11 - 2 D, E). Phallapodeme mostly rod-like (ch. 77 - 0; Figs 11 - 1 A – C 3,4, 11 - 2 D, E; “ Fig. 10 ” in Grimaldi 1991). Phallal guide absent (ch. 78 - 0; Figs 11 - 1 A – C 4, 11 - 2 D, E). Epiproct nearly entirely pubescent (ch. 79 - 0; Fig. 12 A – E 1). Epigynium pubescent only on dorsal portion (ch. 82 - 0; Fig. 12 A – E 1). Hypogynial, anteroventral bridge absent (ch. 85 - 1; Fig. 12 A – E 2); valves separated from each other, distally differentiated into more or less narrow process from basal, broad portion, with only trichoid sensilla (ch. 86 - 0, ch. 87 - 1, ch. 88 - 1; Fig. 12). Included taxa. Colocasiomyia zeylanica species subgroup; and C. oligochaeta subgroup.	en	Zhang, Guang, Gao, Jian-Jun, Takano, Kohei Takenaka, Yafuso, Masako, Suwito, Awit, Meleng, Paulus Ak, Toda, Masanori J. (2023): Phylogenetic classification and palm-inflorescence anthophily of the Colocasiomyia zeylanica species group (Diptera: Drosophilidae), with descriptions of five new species. Zootaxa 5278 (2): 201-238, DOI: 10.11646/zootaxa.5278.2.1, URL: http://dx.doi.org/10.11646/zootaxa.5278.2.1
03C687D9FFC3FFF5A5C6FF28845DE016.taxon	diagnosis	Diagnosis. Male sternites V and VI fused to each other (ch. 46 - 1; Fig. 10 A – C 1). Hypandrial phragma vertical plate-like (ch. 68 - 1; Fig. 11 - 1 A – C 3,4). Shared characters. Antennal arista distally narrowing (ch. 4 - 0; Fig. 3 A – C, “ Fig. 1 A, B ” in Okada 1986, “ Fig. 6 ” in Grimaldi 1991). Eye with stout, dense, interfacetal setulae (ch. 7 - 1; Fig. 2 A – C 1). Proclinate, orbital setae present (ch. 8 - 0; Fig. 2 A – C 2, “ Fig. 1 A, B ” in Okada 1986, “ Fig. 6 ” in Grimaldi 1991). Inner, vertical setae longest on head, in line with ipsilateral orbitals (Fig. 2 A – C 2). Outer, vertical setae present (ch. 9 - 0; Fig. 4 A – C, “ Fig. 1 A, B ” in Okada 1986, “ Fig. 6 ” in Grimaldi 1991). Facial carina shorter than pedicel + 1 st flagellomere (Fig. 2 A – C 2). Gena blackish brown. Upper, postocular setae down-curved, except uppermost ones (ch. 10 - 0; Fig. 4 A – C). Patch of supracervical setae present between dorsomedial and - lateral, tentorial apodemes (ch. 13 - 0; Fig. 4 A – C). Dorsomedial, tentorial apodeme reaching 1 / 4 to vertex; dorsolateral apodemes divergent, nearly straight or distally slightly curved outward (ch. 14 - 0; Fig. 4 A – C). Palpus apically with single, prominent seta (ch. 17 - 1; Fig. 5 A – C 1). Clypeus not thicker than 1 / 2 length of cibarium (Fig. 5 A – C 1). Cibarial, medial sensilla in parallel rows as wide as sensilla campaniformia; posterior sensillum 1 per side (ch. 20 - 0, ch. 21 - 2; Fig. 5 A – C 2). Thorax glossy, blackish brown (Fig. 14 A – C). Postpronotal lobe with 2 prominent setae subequal in length (ch. 25 - 0; Fig. 6 A – C). Anterior, dorsocentral setae situated posteriorly to transverse suture (ch. 26 - 0; Fig. 6 A – C). A pair of prescutellar, acrostichal setae present (ch. 27 - 0; Fig. 6 A – C). Acrostichal setulae in 6 rows (ch. 28 - 1; Fig. 6 A – C). Costal setae in middle row from 2 nd costal section to proximal portion of 3 rd section all apically blunt, heavy, peg-like; sparse setae in upper row small, weak, trichoid (ch. 30 - 0, ch. 31 - 0; Fig. 8 A – C 1). Patch covered with only minute pubescence absent on anterodorsal portion of fore tarsomere I (ch. 33 - 0; Fig. 9 A – C 1). Hind tibia without apicoventral, stout setae (ch. 40 - 0; Fig. 9 A – C 3). Claws shorter than tarsomere V (ch. 43 - 0; Fig. 9 A – C). Abdominal sternites grayish brown; I narrow, transverse plate more or less protruded or expanded posteromedially (ch. 45 - 1; Fig. 10 A – C). Epandrium largely pubescent on dorsal to lateral portion; ventral lobe not differentiated (ch. 53 - 0, ch. 56 - 0; Fig. 11 - 1 A – C 1). Surstylus plate-like, fused to epandrium but not to cercus (ch. 59 - 0, ch. 60 - 1, ch. 63 - 0; Fig. 11 - 1 A – C 1,2). Membranous aedeagus absent (ch. 70 - 2; Fig. 11 - 1 A – C 3,4). Phallapodeme fused to phallal sheath (ch. 75 - 1; Fig. 11 - 1 A – C 4). Hypoproct nearly entirely pubescent (ch. 80 - 0; Fig. 12 A – C 1). Epigynium dorsally neither constricted nor separated (ch. 81 - 0; Fig. 12 A – C 1). Hypogynial valves with trichoid sensilla on both basal and distal portions (Fig. 12 A – C). Included species. Colocasiomyia zeylanica *, C. nepalensis *, C. pinangae sp. nov., C. besaris sp. nov., and C. luciphila sp. nov. * Two known species were assigned to this species subgroup based on their original descriptions and illustrations (Okada 1986; Grimaldi 1991); although the two diagnostic characters are not confirmed for them, they share the following homoplastic synapomorphies with the other members of this subgroup: eye with stout, interfacetal setulae (ch. 7 - 1); and claws shorter than tarsomere V (ch. 43 - 0; “ Fig. 1 A, C ” in Okada 1986).	en	Zhang, Guang, Gao, Jian-Jun, Takano, Kohei Takenaka, Yafuso, Masako, Suwito, Awit, Meleng, Paulus Ak, Toda, Masanori J. (2023): Phylogenetic classification and palm-inflorescence anthophily of the Colocasiomyia zeylanica species group (Diptera: Drosophilidae), with descriptions of five new species. Zootaxa 5278 (2): 201-238, DOI: 10.11646/zootaxa.5278.2.1, URL: http://dx.doi.org/10.11646/zootaxa.5278.2.1
03C687D9FFC3FFF6A5C6FAC3824CE335.taxon	description	(Figs 2 – 12 A, 14 A)	en	Zhang, Guang, Gao, Jian-Jun, Takano, Kohei Takenaka, Yafuso, Masako, Suwito, Awit, Meleng, Paulus Ak, Toda, Masanori J. (2023): Phylogenetic classification and palm-inflorescence anthophily of the Colocasiomyia zeylanica species group (Diptera: Drosophilidae), with descriptions of five new species. Zootaxa 5278 (2): 201-238, DOI: 10.11646/zootaxa.5278.2.1, URL: http://dx.doi.org/10.11646/zootaxa.5278.2.1
03C687D9FFC3FFF6A5C6FAC3824CE335.taxon	diagnosis	Diagnosis. Surstylus with 5 – 7 stout, short, peg-like teeth (Fig. 11 - 1 A 2). Pregonites apically round, with coarse serrations on ventral margin (Fig. 11 - 1 A 4). Epigynium with 11 – 12 sensilla per side near lower posterior margin (Fig. 12 A 1). Distal, narrow portion of hypogynial valve slightly shorter than basal, broad portion (Fig. 12 A).	en	Zhang, Guang, Gao, Jian-Jun, Takano, Kohei Takenaka, Yafuso, Masako, Suwito, Awit, Meleng, Paulus Ak, Toda, Masanori J. (2023): Phylogenetic classification and palm-inflorescence anthophily of the Colocasiomyia zeylanica species group (Diptera: Drosophilidae), with descriptions of five new species. Zootaxa 5278 (2): 201-238, DOI: 10.11646/zootaxa.5278.2.1, URL: http://dx.doi.org/10.11646/zootaxa.5278.2.1
03C687D9FFC3FFF6A5C6FAC3824CE335.taxon	description	Description (♁ and ♀). Head. Antennal pedicel blackish brown; first flagellomere approximately 1.7 times as long as pedicel; arista approximately 2.1 times as long as first flagellomere (Fig. 3 A). Distance between antennal scapes narrower than scape diameter (Fig. 2 A 2). Facial carina wider than 1 st flagellomere (Fig. 2 A 2). Supracervical setae 4 – 7 and postoculars 17 – 18 per side (Fig. 4 A). Cibarial projections at anterolateral corners as long as width of anterior margin; medial sensilla 2 – 4 per side (Fig. 5 A 2). Prementum and neighboring lateral membrane with 7 – 8 setae per side (Fig. 5 A 2). Labellum with 10 pseudotracheae per side (Fig. 5 A 1). Thorax. Two katepisternal, prominent setae longer than acrostichal setulae (Fig. 7 A). Legs grayish brown, except for fore tibiae and all tarsi pale grayish yellow (Fig. 14 A). Fore tarsomere I slightly longer than tarsomeres II and III combined; mid I as long as II + III + IV; hind I slightly longer than II + III + IV (Fig. 9 A). Fore tarsomere II with 1 large and 1 small pegs; latter 1 / 3 as long as former (Fig. 9 A 1). Abdomen. Tergites blackish brown (Fig. 14 A). Male sternites III and IV as wide as long; V + VI distally deeply bilobed (Fig. 10 A 1). Female sternites III – V as wide as long; VI wider than long; VII wider than long plate bilobed in distal half (Fig. 10 A 2). Female tergite VII caudomedially deeply notched. Male terminalia. Epandrium with 2 – 4 long setae only on lateral to ventral portion (Fig. 11 - 1 A 1). Cercus nearly entirely pubescent except for small, anteroventral, expanded flap, with approximately 37 setae (Fig. 11 - 1 A 1). Phallal sheath not pubescent (Fig. 11 - 1 A 4). Female terminalia. Perineal membrane with patch of dense, distinct warts. Measurements (holotype / range in 6 ♁ and 5 ♀ paratypes, in mm): BL (straight distance from anterior edge of pedicel to tip of abdomen) = 1.45 / ♁ 1.29 – 1.54, ♀ 1.43 – 1.62; ThL (distance from anterior notal margin to apex of scutellum) = 0.58 / ♁ 0.59 – 0.67, ♀ 0.60 – 0.67; WL (distance from humeral cross vein to wing apex) = 1.22 / ♁ 1.25 – 1.40, ♀ 1.23 – 1.44; WW (maximum wing width) = 0.60 / ♁ 0.58 – 0.66, ♀ 0.59 – 0.67. Indices (holotype / range in 6 ♁ and 5 ♀, or less if noted, paratypes, in ratio): FW / HW (frontal width / head width) = 0.55 / 0.53 – 0.60; ch / o (maximum width of gena / maximum diameter of eye) = 0.36 / 0.29 – 0.40; prorb (proclinate orbital seta length / posterior reclinate orbital seta length) = 0.85 / 0.84 – 0.97 (2 ♁, 1 ♀); rcorb (anterior reclinate orbital seta length / posterior reclinate orbital seta length) = 0.30 / 0.23 – 0.40 (3 ♁, 1 ♀); vb (subvibrissal seta length / vibrissa length) = 0.33 / 0.22 – 0.32; dcl (anterior dorsocentral seta length / posterior dorsocentral seta length) = 0.46 / 0.44 – 0.71 (4 ♁, 4 ♀); presctl (prescutellar acrostichal seta length / posterior dorsocentral seta length) = 0.42 / 0.25 – 0.40 (6 ♁, 4 ♀); sctl (basal scutellar seta length / apical scutellar seta length) = 0.54 / 0.54 – 0.63 (6 ♁, 4 ♀); sterno (anterior katepisternal seta length / posterior katepisternal seta length) = 1.00 / 0.94 – 1.26; orbito (distance between proclinate and posterior reclinate orbital setae / distance between inner vertical and posterior reclinate orbital setae) = 0.35 / 0.32 – 0.51; dcp (distance between ipsilateral dorsocentral setae / distance between anterior dorsocentral setae) = 0.50 / 0.57 – 0.65; sctlp (distance between ipsilateral scutellar setae / distance between apical scutellar setae) = 1.07 / 0.86 – 1.03; C (2 nd costal section between subcostal break and R 2 + 3 / 3 rd costal section between R 2 + 3 and R 4 + 5) = 2.04 / 1.70 – 2.16; 4 c (3 rd costal section between R 2 + 3 and R 4 + 5 / M 1 between r-m and dm-m) = 1.46 / 1.30 – 1.54; 4 v (M 1 between dm-m and wing margin / M 1 between r-m and dm-m) = 2.86 / 2.48 – 2.78; 5 x (M 4 between dm-m and wing margin / dm-m between M 1 and M 4) = 3.18 / 2.50 – 3.35; ac (3 rd costal section between R 2 + 3 and R 4 + 5 / distance between distal ends of R 4 + 5 and M 1) = 2.63 / 2.24 – 3.04; M (M 4 between dm-m and wing margin / M 1 between r-m and dm-m) = 1.11 / 0.87 – 1.08; C 3 F (length of heavy setation in 3 rd costal section / length of 3 rd costal section) = 0.37 / 0.37 – 0.49.	en	Zhang, Guang, Gao, Jian-Jun, Takano, Kohei Takenaka, Yafuso, Masako, Suwito, Awit, Meleng, Paulus Ak, Toda, Masanori J. (2023): Phylogenetic classification and palm-inflorescence anthophily of the Colocasiomyia zeylanica species group (Diptera: Drosophilidae), with descriptions of five new species. Zootaxa 5278 (2): 201-238, DOI: 10.11646/zootaxa.5278.2.1, URL: http://dx.doi.org/10.11646/zootaxa.5278.2.1
03C687D9FFC3FFF6A5C6FAC3824CE335.taxon	materials_examined	Holotype. ♁ (MZB), “ Cibodas, West Java, Indonesia, 12. xii. 2004, K. T. Takano / ex inflorescence of Pinanga coronata ”. Paratypes. Indonesia: 24 ♁, 26 ♀, same data as the holotype (MZB, SEHU); 10 ♁, 10 ♀ (# 05160 – 79), same data as the holotype except for 24. xii. 2003 (KIZ).	en	Zhang, Guang, Gao, Jian-Jun, Takano, Kohei Takenaka, Yafuso, Masako, Suwito, Awit, Meleng, Paulus Ak, Toda, Masanori J. (2023): Phylogenetic classification and palm-inflorescence anthophily of the Colocasiomyia zeylanica species group (Diptera: Drosophilidae), with descriptions of five new species. Zootaxa 5278 (2): 201-238, DOI: 10.11646/zootaxa.5278.2.1, URL: http://dx.doi.org/10.11646/zootaxa.5278.2.1
03C687D9FFC3FFF6A5C6FAC3824CE335.taxon	distribution	Distribution. Indonesia (West Java).	en	Zhang, Guang, Gao, Jian-Jun, Takano, Kohei Takenaka, Yafuso, Masako, Suwito, Awit, Meleng, Paulus Ak, Toda, Masanori J. (2023): Phylogenetic classification and palm-inflorescence anthophily of the Colocasiomyia zeylanica species group (Diptera: Drosophilidae), with descriptions of five new species. Zootaxa 5278 (2): 201-238, DOI: 10.11646/zootaxa.5278.2.1, URL: http://dx.doi.org/10.11646/zootaxa.5278.2.1
03C687D9FFC3FFF6A5C6FAC3824CE335.taxon	discussion	Remarks. This species resembles C. nepalensis, but can be distinguished from it in having the shorter arista (in C. nepalensis: “ 2.5 × the length of flagellomere I ”; Grimaldi 1991), cibarial medial sensilla in parallel rows as wide as sensilla campaniformia (narrower than the latter; “ Fig. 7 ” in Grimaldi 1991), two equally long katepisternal setae (“ Katepisternum … with 1 large seta ”), and two pegs much different in size on the fore tarsomere II (“ Tarsal segment on foreleg … bearing a pair of black, spine-like setae; medial one slightly longer ”).	en	Zhang, Guang, Gao, Jian-Jun, Takano, Kohei Takenaka, Yafuso, Masako, Suwito, Awit, Meleng, Paulus Ak, Toda, Masanori J. (2023): Phylogenetic classification and palm-inflorescence anthophily of the Colocasiomyia zeylanica species group (Diptera: Drosophilidae), with descriptions of five new species. Zootaxa 5278 (2): 201-238, DOI: 10.11646/zootaxa.5278.2.1, URL: http://dx.doi.org/10.11646/zootaxa.5278.2.1
03C687D9FFC3FFF6A5C6FAC3824CE335.taxon	etymology	Etymology. Referring to the host-plant genus Pinanga.	en	Zhang, Guang, Gao, Jian-Jun, Takano, Kohei Takenaka, Yafuso, Masako, Suwito, Awit, Meleng, Paulus Ak, Toda, Masanori J. (2023): Phylogenetic classification and palm-inflorescence anthophily of the Colocasiomyia zeylanica species group (Diptera: Drosophilidae), with descriptions of five new species. Zootaxa 5278 (2): 201-238, DOI: 10.11646/zootaxa.5278.2.1, URL: http://dx.doi.org/10.11646/zootaxa.5278.2.1
03C687D9FFC0FFF7A5C6F9E5838AE1F9.taxon	description	(Figs 2 – 12 B, 14 B)	en	Zhang, Guang, Gao, Jian-Jun, Takano, Kohei Takenaka, Yafuso, Masako, Suwito, Awit, Meleng, Paulus Ak, Toda, Masanori J. (2023): Phylogenetic classification and palm-inflorescence anthophily of the Colocasiomyia zeylanica species group (Diptera: Drosophilidae), with descriptions of five new species. Zootaxa 5278 (2): 201-238, DOI: 10.11646/zootaxa.5278.2.1, URL: http://dx.doi.org/10.11646/zootaxa.5278.2.1
03C687D9FFC0FFF7A5C6F9E5838AE1F9.taxon	diagnosis	Diagnosis. Surstylus with 8 – 10 stout, longer, peg-like teeth; dorsalmost one thickest (Fig. 11 - 1 B 2). Pregonites apically pointed, smooth on ventral margin (Fig. 11 - 1 B 4). Epigynium with 26 – 28 small sensilla per side near lower posterior margin (Fig. 12 B 1). Distal, narrow portion of hypogynial valve longer than basal, broad portion (Fig. 12 B).	en	Zhang, Guang, Gao, Jian-Jun, Takano, Kohei Takenaka, Yafuso, Masako, Suwito, Awit, Meleng, Paulus Ak, Toda, Masanori J. (2023): Phylogenetic classification and palm-inflorescence anthophily of the Colocasiomyia zeylanica species group (Diptera: Drosophilidae), with descriptions of five new species. Zootaxa 5278 (2): 201-238, DOI: 10.11646/zootaxa.5278.2.1, URL: http://dx.doi.org/10.11646/zootaxa.5278.2.1
03C687D9FFC0FFF7A5C6F9E5838AE1F9.taxon	description	Description (♁ and ♀). Head. Antennal pedicel blackish brown; first flagellomere approximately 1.6 times as long as pedicel; arista approximately 2.1 times as long as first flagellomere (Fig. 3 B). Distance between antennal scapes as wide as scape diameter (Fig. 2 B 2). Facial carina wider than 1 st flagellomere (Fig. 2 B 2). Supracervical setae 8 – 10 and postoculars 18 – 20 per side (Fig. 4 A). Cibarial projections at anterolateral corners as long as width of anterior margin; medial sensilla approximately 2 – 3 per side (Fig. 5 B 2). Prementum and neighboring lateral membrane with 9 – 10 setae per side (Fig. 5 B 2). Labellum with 11 pseudotracheae per side (Fig. 5 B 1). Thorax. Two katepisternal, prominent setae longer than acrostichal setulae (Fig. 7 B). Legs grayish brown, except for all tarsi pale grayish yellow (Fig. 14 B). Fore tarsomere I slightly longer than tarsomeres II and III combined; mid I as long as II + III + IV; hind I slightly longer than II + III + IV (Fig. 9 B). Fore tarsomere II with 1 large and 1 small pegs; latter 1 / 3 as long as former (Fig. 9 B 1). Abdomen. Tergites blackish brown (Fig. 14 B). Male sternites III and IV as wide as long; V + VI distally deeply bilobed (Fig. 10 B 1). Female sternites III – V as wide as long; VI wider than long; VII wider than long plate bilobed in distal half (Fig. 10 B 2). Female tergite VII mid-dorsally separated into 2 lateral plates. Male terminalia. Epandrium with 2 – 4 long setae only on lateral to ventral portion (Fig. 11 - 1 B 1). Cercus nearly entirely pubescent except for roundish, ventral flap, with approximately 40 setae (Fig. 11 - 1 B 1). Phallal sheath partly pubescent (Fig. 11 - 1 B 4). Female terminalia. Perineal membrane with patch of dense, distinct warts. Measurements (holotype / range in 2 ♁ and 1 ♀ paratypes, in mm): BL = 1.51 / ♁ 1.35 – 1.55, ♀ 1.70; ThL = 0.67 / ♁ 0.72 – 0.73, ♀ 0.72; WL = 1.39 / ♁ 1.46 – 1.55, ♀ 1.47; WW = 0.66 / ♁ 0.72, ♀ 0.72. Indices (holotype / range in 2 ♁ and 1 ♀, or less if noted, paratypes, in ratio): FW / HW = 0.54 / 0.53 – 0.56; ch / o = 0.38 / 0.32 – 0.40; prorb = 0.83 / 0.82 – 0.97; rcorb = 0.30 / 0.25 – 0.32; vb = 0.21 / 0.24 – 0.29; dcl = 0.51 / 0.52 – 0.54 (1 ♁ and 1 ♀); presctl = 0.30 / 0.35 – 0.41; sctl = 0.75 / 0.71 – 0.73 (1 ♁ and 1 ♀); sterno = 1.07 / 1.28 – 1.42; orbito = 0.35 / 0.38 – 0.45; dcp = 0.58 / 0.68 – 0.77; sctlp = 0.81 / 0.91 – 1.07; C = 2.10 / 1.82 – 2.18; 4 c = 1.32 / 1.22 – 1.39; 4 v = 2.54 / 2.33 – 2.61; 5 x = 2.78 / 1.82 – 3.23; ac = 2.50 / 2.36 – 2.87; M = 0.99 / 0.91 – 1.02; C 3 F = 0.41 / 0.46 – 0.54.	en	Zhang, Guang, Gao, Jian-Jun, Takano, Kohei Takenaka, Yafuso, Masako, Suwito, Awit, Meleng, Paulus Ak, Toda, Masanori J. (2023): Phylogenetic classification and palm-inflorescence anthophily of the Colocasiomyia zeylanica species group (Diptera: Drosophilidae), with descriptions of five new species. Zootaxa 5278 (2): 201-238, DOI: 10.11646/zootaxa.5278.2.1, URL: http://dx.doi.org/10.11646/zootaxa.5278.2.1
03C687D9FFC0FFF7A5C6F9E5838AE1F9.taxon	materials_examined	Holotype. ♁ (MZB), “ Cibodas, West Java, Indonesia, 12. xii. 2004, K. T. Takano / ex inflorescence of Pinanga coronata ”. Paratypes. Indonesia: 12 ♁, 4 ♀, same data as the holotype (MZB, SEHU); 2 ♁ (# 05200 – 01), 1 ♀ (# 05226), same data as the holotype except for 24. xii. 2003 (KIZ).	en	Zhang, Guang, Gao, Jian-Jun, Takano, Kohei Takenaka, Yafuso, Masako, Suwito, Awit, Meleng, Paulus Ak, Toda, Masanori J. (2023): Phylogenetic classification and palm-inflorescence anthophily of the Colocasiomyia zeylanica species group (Diptera: Drosophilidae), with descriptions of five new species. Zootaxa 5278 (2): 201-238, DOI: 10.11646/zootaxa.5278.2.1, URL: http://dx.doi.org/10.11646/zootaxa.5278.2.1
03C687D9FFC0FFF7A5C6F9E5838AE1F9.taxon	distribution	Distribution. Indonesia (West Java).	en	Zhang, Guang, Gao, Jian-Jun, Takano, Kohei Takenaka, Yafuso, Masako, Suwito, Awit, Meleng, Paulus Ak, Toda, Masanori J. (2023): Phylogenetic classification and palm-inflorescence anthophily of the Colocasiomyia zeylanica species group (Diptera: Drosophilidae), with descriptions of five new species. Zootaxa 5278 (2): 201-238, DOI: 10.11646/zootaxa.5278.2.1, URL: http://dx.doi.org/10.11646/zootaxa.5278.2.1
03C687D9FFC0FFF7A5C6F9E5838AE1F9.taxon	discussion	Remarks. This species resembles C. pinangae sp. nov., but can be distinguished from it by the diagnostic characters.	en	Zhang, Guang, Gao, Jian-Jun, Takano, Kohei Takenaka, Yafuso, Masako, Suwito, Awit, Meleng, Paulus Ak, Toda, Masanori J. (2023): Phylogenetic classification and palm-inflorescence anthophily of the Colocasiomyia zeylanica species group (Diptera: Drosophilidae), with descriptions of five new species. Zootaxa 5278 (2): 201-238, DOI: 10.11646/zootaxa.5278.2.1, URL: http://dx.doi.org/10.11646/zootaxa.5278.2.1
03C687D9FFC0FFF7A5C6F9E5838AE1F9.taxon	etymology	Etymology. The specific name derives from “ besar ” (meaning “ large ”) of Indonesian, referring to its larger body size among the members of the zeylanica group.	en	Zhang, Guang, Gao, Jian-Jun, Takano, Kohei Takenaka, Yafuso, Masako, Suwito, Awit, Meleng, Paulus Ak, Toda, Masanori J. (2023): Phylogenetic classification and palm-inflorescence anthophily of the Colocasiomyia zeylanica species group (Diptera: Drosophilidae), with descriptions of five new species. Zootaxa 5278 (2): 201-238, DOI: 10.11646/zootaxa.5278.2.1, URL: http://dx.doi.org/10.11646/zootaxa.5278.2.1
03C687D9FFC1FFF8A5C6FAA9820BE1F9.taxon	description	(Figs 2 – 12 C, 14 C)	en	Zhang, Guang, Gao, Jian-Jun, Takano, Kohei Takenaka, Yafuso, Masako, Suwito, Awit, Meleng, Paulus Ak, Toda, Masanori J. (2023): Phylogenetic classification and palm-inflorescence anthophily of the Colocasiomyia zeylanica species group (Diptera: Drosophilidae), with descriptions of five new species. Zootaxa 5278 (2): 201-238, DOI: 10.11646/zootaxa.5278.2.1, URL: http://dx.doi.org/10.11646/zootaxa.5278.2.1
03C687D9FFC1FFF8A5C6FAA9820BE1F9.taxon	diagnosis	Diagnosis. Antennal, first flagellomere approximately 2.4 times as long as pedicel; arista approximately 1.6 times as long as first flagellomere (Fig. 3 C). Fore tarsomere II with 2 pegs; smaller peg slightly longer than 1 / 2 of larger one (Fig. 9 C 1). Male sternites III and IV longer than wide; V + VI concave on distal margin (Fig. 10 C 1). Female sternites III – VI longer than wide; VII much wider than long, slightly convex on distal margin (Fig. 10 C 2). Surstylus with 6 – 7 stout, longer, peg-like teeth nearly uniform in thickness (Fig. 11 - 1 C 1,2). Pregonites apically roundish, smooth on ventral margin (Fig. 11 - 1 C 4 a). Epigynium with 2 – 6 small sensilla per side near lower posterior margin (Fig. 12 C 1). Distal, narrow portion of hypogynial valve half as long as basal, broad portion (Fig. 12 C).	en	Zhang, Guang, Gao, Jian-Jun, Takano, Kohei Takenaka, Yafuso, Masako, Suwito, Awit, Meleng, Paulus Ak, Toda, Masanori J. (2023): Phylogenetic classification and palm-inflorescence anthophily of the Colocasiomyia zeylanica species group (Diptera: Drosophilidae), with descriptions of five new species. Zootaxa 5278 (2): 201-238, DOI: 10.11646/zootaxa.5278.2.1, URL: http://dx.doi.org/10.11646/zootaxa.5278.2.1
03C687D9FFC1FFF8A5C6FAA9820BE1F9.taxon	description	Description (♁ and ♀). Head. Antennal pedicel grayish brown. Distance between antennal scapes as wide as scape diameter (Fig. 2 C 2). Facial carina narrower than 1 st flagellomere (Fig. 2 C 2). Supracervical setae 6 – 8 and postoculars 13 – 18 per side (Fig. 4 C). Cibarial projections at anterolateral corners slightly longer than width of anterior margin; medial sensilla approximately 3 – 4 per side (Fig. 5 C 2). Prementum and neighboring lateral membrane with 7 – 8 setae per side (Fig. 5 C 2). Labellum with 11 – 13 pseudotracheae per side (Fig. 5 C 1). Thorax. Prescutellar, acrostichal setae only slightly longer than acrostichal setulae (Fig. 6 C). Only one prominent katepisternal seta longer than acrostichal setulae (Fig. 7 C). Legs grayish brown, except for all tarsi pale grayish yellow (Fig. 14 C). Fore, mid and hind tarsomere Is as long as II + III, II + III + IV and II + III + IV + V, respectively (Fig. 9 C). Abdomen. Tergites dark grayish brown (Fig. 14 C). Female tergite VII mid-dorsally separated into 2 lateral plates. Male terminalia. Epandrium with approximately 4 long setae only on lateral to ventral portion (Fig. 11 - 1 C 1). Cercus entirely pubescent, round on ventral margin, with 28 – 29 setae (Fig. 11 - 1 C 1). Phallal sheath partly pubescent (Fig. 11 - 1 C 4, b). Female terminalia. Perineal membrane without patch of dense, distinct warts. Measurements (holotype / range in 1 ♁ and 3 ♀ paratypes, in mm): BL = 1.51 / ♁ 1.59, ♀ 1.40 – 1.92; ThL = 0.60 / ♁ 0.60, ♀ 0.48 – 0.75; WL = 1.22 / ♁ 1.23, ♀ 1.06 – 1.55; WW = 0.57 / ♁ 0.61, ♀ 0.50 – 0.73. Indices (holotype / range in 1 ♁ and 3 ♀, or less if noted, paratypes, in ratio): FW / HW = 0.48 / 0.49 – 0.55; ch / o = 0.29 / 0.33 – 0.38; prorb = 0.86 / 0.82 – 1.08; rcorb = 0.32 / 0.29 – 0.40; vb = 0.17 / 0.18 – 0.25 (3 ♀); dcl = 0.39 / 0.35 – 0.40 (1 ♁ and 2 ♀); presctl = 0.20 / 0.21 – 0.25 (1 ♁ and 2 ♀); sctl = 0.56 / 0.52 – 0.60; orbito = 0.45 / 0.33 – 0.47; dcp = 0.54 / 0.49 – 0.59; sctlp = 0.93 / 0.88 – 1.03; C = 1.57 / 1.49 – 1.64; 4 c = 1.61 / 1.39 – 1.64; 4 v = 2.39 / 2.20 – 2.63; 5 x = 2.61 / 2.62 – 2.75; ac = 3.27 / 2.49 – 3.09; M = 0.90 / 0.78 – 0.92; C 3 F = 0.60 / 0.49 – 0.72.	en	Zhang, Guang, Gao, Jian-Jun, Takano, Kohei Takenaka, Yafuso, Masako, Suwito, Awit, Meleng, Paulus Ak, Toda, Masanori J. (2023): Phylogenetic classification and palm-inflorescence anthophily of the Colocasiomyia zeylanica species group (Diptera: Drosophilidae), with descriptions of five new species. Zootaxa 5278 (2): 201-238, DOI: 10.11646/zootaxa.5278.2.1, URL: http://dx.doi.org/10.11646/zootaxa.5278.2.1
03C687D9FFC1FFF8A5C6FAA9820BE1F9.taxon	materials_examined	Holotype. ♁ (RDID), “ Lambir Hills National Park, Sarawak, Malaysia, 25 – 26. i. 2012, ex light trap (LT- 35), M. J. Toda ”. Paratypes. Malaysia: 5 ♀, same data as the holotype (SEHU); 1 ♀, same except for 26 – 27. i. 2012, 1 ♀, same except for 23. i. 2012, ex florescent light, 1 ♀, same except for 25. i. 2012, ex florescent light (RDID); 1 ♁ (# 05225), same data as the holotype, 1 ♀ (# 05223), same except for 23 – 24. i. 2012 (Lot # 229), 1 ♀ (# 05224), same except for 24 – 25. i. 2012 (Lot # 230) (KIZ).	en	Zhang, Guang, Gao, Jian-Jun, Takano, Kohei Takenaka, Yafuso, Masako, Suwito, Awit, Meleng, Paulus Ak, Toda, Masanori J. (2023): Phylogenetic classification and palm-inflorescence anthophily of the Colocasiomyia zeylanica species group (Diptera: Drosophilidae), with descriptions of five new species. Zootaxa 5278 (2): 201-238, DOI: 10.11646/zootaxa.5278.2.1, URL: http://dx.doi.org/10.11646/zootaxa.5278.2.1
03C687D9FFC1FFF8A5C6FAA9820BE1F9.taxon	distribution	Distribution. Malaysia (Sarawak).	en	Zhang, Guang, Gao, Jian-Jun, Takano, Kohei Takenaka, Yafuso, Masako, Suwito, Awit, Meleng, Paulus Ak, Toda, Masanori J. (2023): Phylogenetic classification and palm-inflorescence anthophily of the Colocasiomyia zeylanica species group (Diptera: Drosophilidae), with descriptions of five new species. Zootaxa 5278 (2): 201-238, DOI: 10.11646/zootaxa.5278.2.1, URL: http://dx.doi.org/10.11646/zootaxa.5278.2.1
03C687D9FFC1FFF8A5C6FAA9820BE1F9.taxon	discussion	Remarks. This species resembles C. nepalensis in having only one prominent, katepisternal seta and two pegs less different in length on the fore tarsomere II, but can be distinguished from it by the arista 1.6 times as long as first flagellomere (in C. nepalensis: “ 2.5 × the length of flagellomere I ”; Grimaldi 1991), cibarial medial sensilla in parallel rows as wide as sensilla campaniformia (narrower than the latter; “ Fig. 7 ” in Grimaldi 1991), the facial carina narrower than 1 st flagellomere (wider than the latter; “ Fig. 6 ” in Grimaldi 1991), and the absence of membranous aedeagus (“ Distiphallus fusiform, apically with fine spicules, covered with membrane ”). On the other hand, the following characters suggest some relatedness to the oligochaeta species subgroup described below: only one prominent, katepisternal seta; prescutellar, acrostichal setae less differentiated; and male sternites III and IV and female III – VI longer than wide.	en	Zhang, Guang, Gao, Jian-Jun, Takano, Kohei Takenaka, Yafuso, Masako, Suwito, Awit, Meleng, Paulus Ak, Toda, Masanori J. (2023): Phylogenetic classification and palm-inflorescence anthophily of the Colocasiomyia zeylanica species group (Diptera: Drosophilidae), with descriptions of five new species. Zootaxa 5278 (2): 201-238, DOI: 10.11646/zootaxa.5278.2.1, URL: http://dx.doi.org/10.11646/zootaxa.5278.2.1
03C687D9FFC1FFF8A5C6FAA9820BE1F9.taxon	etymology	Etymology. Referring to its nature of being attracted to light.	en	Zhang, Guang, Gao, Jian-Jun, Takano, Kohei Takenaka, Yafuso, Masako, Suwito, Awit, Meleng, Paulus Ak, Toda, Masanori J. (2023): Phylogenetic classification and palm-inflorescence anthophily of the Colocasiomyia zeylanica species group (Diptera: Drosophilidae), with descriptions of five new species. Zootaxa 5278 (2): 201-238, DOI: 10.11646/zootaxa.5278.2.1, URL: http://dx.doi.org/10.11646/zootaxa.5278.2.1
03C687D9FFCEFFF9A5C6FAA9854AE605.taxon	diagnosis	Diagnosis. Antennal arista apically truncate, stick-shaped (ch. 4 - 1; Fig. 3 D, E). Proclinate, orbital setae absent (ch. 8 - 1; Fig. 2 D, E). Anterior, dorsocentral setae absent (ch. 26 - 2; Fig. 6 D, E). Patch of supracervical setae present over basal portion of dorsolateral, tentorial apodeme (ch. 13 - 1; Fig. 4 D, E). Dorsolateral, tentorial apodemes distally strongly curved outward (ch. 14 - 1; Fig. 4 D, E). Palpus without prominent seta (ch. 17 - 2; Fig. 5 D 1, E 1). 5 x index> 4.0 (ch. 32 - 2; Fig. 8 D 2, E 2). Hind tibia apicoventrally with two stout setae (ch. 40 - 1; Fig. 9 D 3, E 3). Epandrium with neither pubescence nor setae (ch. 53 - 1, ch. 54 - 1; Fig. 11 - 2 D, E). Shared characters. Antennal pedicel grayish brown (Fig. 14 D, E). Eye with fine, sparse, interfacetal setulae (ch. 7 - 0; Fig. 2 D 1, E 1). Outer, vertical setae absent (ch. 9 - 1; Fig. 4 D, E). Inner (convergent) vertical setae dislocated outside line of ipsilateral orbitals; posterior reclinate orbitals longest on head (Fig. 2 D 2, E 2). Upper, postocular setae upright or up-curved (ch. 10 - 1; Fig. 4 D, E). Dorsomedial, tentorial apodeme reaching 1 / 5 to vertex (Fig. 4 D, E). Facial carina wider than 1 st flagellomere, as long as pedicel + 1 st flagellomere (Fig. 2 D 2, E 2). Clypeus thicker than 1 / 2 length of cibarium (Fig. 5 D 1, E 1). Cibarial, medial sensilla 2 per side; posterior sensilla absent (ch. 21 - 3; Fig. 5 D 2, E 2). Labellum with 7 pseudotracheae (ch. 24 - 0; Fig. 5 D 1, E 1). Postpronotal lobe with 1 prominent seta (ch. 25 - 2; Fig. 6 D, E). Prescutellar, acrostichal setae absent (ch. 27 - 1; Fig. 6 D, E). Only one prominent katepisternal seta longer than acrostichal setulae (ch. 29 - 2; Fig. 7 D, E). Costal setae in middle row from 2 nd costal section to proximal portion of 3 rd section all weak, trichoid; sparse setae in upper row stout, longer, trichoid (ch. 30 - 2, ch. 31 - 1; Fig. 8 D 1, E 1). Patch covered with only minute pubescence present on anterodorsal portion of fore tarsomere I (ch. 33 - 1; Fig. 9 D 1, E 1). Fore tarsomere II with 1 large and 1 small pegs; latter 1 / 3 as long as former (ch. 37 - 1; Fig. 9 D 1, E 1). Mid tarsomere I as long as tarsomeres II, III and IV combined; hind I slightly longer than II + III + IV (Fig. 9 D 2,3, E 2,3). Claws as long as tarsomere V (ch. 43 - 1; Fig. 9 D, E). Abdominal tergites blackish brown (Fig. 14 D, E). All sternites longer than wide and I fused to II in both sexes (ch. 45 - 2; Fig. 10 D, E); male V not fused to VI, and VI completely bilobed (ch. 46 - 0, ch. 47 - 1; Fig. 10 D 1, E 1); female sternite VII more or less convex on distal margin (ch. 51 - 2; Fig. 10 D 2, E 2). Female tergite VII mid-dorsally separated into 2 triangular, lateral plates (ch. 52 - 0). Epandrial, ventral lobe differentiated (ch. 56 - 1; Fig. 11 - 2 D, E). Surstylus absent (ch. 59 - 4; Fig. 11 - 2 D, E). Hypandrial phragma anterior, strongly sclerotized rod-like portion (ch. 68 - 2; Fig. 11 - 2 D, E). Membranous aedeagus long, tube-like (ch. 70 - 1; Fig. 11 - 2 D, E). Phallal sheath not pubescent (ch. 74 - 0; Fig. 11 - 2 D, E). Phallapodeme articulated to and not shorter than phallal sheath (ch. 75 - 0, ch. 76 - 0; Fig. 11 - 2 D, E). Hypoproct partly pubescent (ch. 80 - 1; Fig. 12 D 1, E 1). Epigynium dorsally separated into 2 lateral lobes (ch. 81 - 1; Fig. 12 D 1, E 1). Perineal membrane with patch of dense, distinct warts (ch. 84 - 1; Fig. 12 E). Hypogynial valve with trichoid sensilla on only distal portion; distal, narrow portion as long as basal, broad portion (Fig. 12 D, E).	en	Zhang, Guang, Gao, Jian-Jun, Takano, Kohei Takenaka, Yafuso, Masako, Suwito, Awit, Meleng, Paulus Ak, Toda, Masanori J. (2023): Phylogenetic classification and palm-inflorescence anthophily of the Colocasiomyia zeylanica species group (Diptera: Drosophilidae), with descriptions of five new species. Zootaxa 5278 (2): 201-238, DOI: 10.11646/zootaxa.5278.2.1, URL: http://dx.doi.org/10.11646/zootaxa.5278.2.1
03C687D9FFCEFFF9A5C6FAA9854AE605.taxon	discussion	Remarks. This species subgroup is well demarcated by a number of non-homoplastic synapomorphies (Fig. 13) defined as the diagnostic characters, especially the peculiar chaetotaxy that the proclinate orbital, outer vertical, and anterior dorsocentral setae are lacking. Such an anomaly from the basic chaetotaxy is rare in the Drosophilidae, but is seen in a few species of Colocasiomyia: C. crassipes (de Meijere) and C. micheliae of the crassipes group lack the outer vertical and dorsocentral setae (Grimaldi 1991; Yafuso et al. 2000); and C. ecornuta Toda & Takano of the cristata group lacks the outer vertical setae (Takano et al. 2021). Included taxa. Colocasiomyia oligochaeta sp. nov., and C. grimaldii sp. nov.	en	Zhang, Guang, Gao, Jian-Jun, Takano, Kohei Takenaka, Yafuso, Masako, Suwito, Awit, Meleng, Paulus Ak, Toda, Masanori J. (2023): Phylogenetic classification and palm-inflorescence anthophily of the Colocasiomyia zeylanica species group (Diptera: Drosophilidae), with descriptions of five new species. Zootaxa 5278 (2): 201-238, DOI: 10.11646/zootaxa.5278.2.1, URL: http://dx.doi.org/10.11646/zootaxa.5278.2.1
03C687D9FFCFFFFAA5C6FCC7857DE5E9.taxon	description	(Figs 2 – 12 D, 14 D)	en	Zhang, Guang, Gao, Jian-Jun, Takano, Kohei Takenaka, Yafuso, Masako, Suwito, Awit, Meleng, Paulus Ak, Toda, Masanori J. (2023): Phylogenetic classification and palm-inflorescence anthophily of the Colocasiomyia zeylanica species group (Diptera: Drosophilidae), with descriptions of five new species. Zootaxa 5278 (2): 201-238, DOI: 10.11646/zootaxa.5278.2.1, URL: http://dx.doi.org/10.11646/zootaxa.5278.2.1
03C687D9FFCFFFFAA5C6FCC7857DE5E9.taxon	diagnosis	Diagnosis. Antennal arista approximately 1.8 times as long as first flagellomere (Fig. 3 D). Distance between antennal scapes as wide as scape diameter (Fig. 2 D 2). Acrostichal setulae in 6 rows (Fig. 6 D). All legs entirely grayish yellow to yellow except for grayish brown tarsomere Vs (Fig. 14 D). Cercus slightly projected at apicoventral corner (Fig. 11 - 2 D). Long, tube-like, membranous aedeagus ornamented with relatively sparse and weak spinules (Fig. 11 - 2 D). Hypogynial valves with dorsal flaps fused into small, ligulate plate at distal end of basal portion (Fig. 12 D).	en	Zhang, Guang, Gao, Jian-Jun, Takano, Kohei Takenaka, Yafuso, Masako, Suwito, Awit, Meleng, Paulus Ak, Toda, Masanori J. (2023): Phylogenetic classification and palm-inflorescence anthophily of the Colocasiomyia zeylanica species group (Diptera: Drosophilidae), with descriptions of five new species. Zootaxa 5278 (2): 201-238, DOI: 10.11646/zootaxa.5278.2.1, URL: http://dx.doi.org/10.11646/zootaxa.5278.2.1
03C687D9FFCFFFFAA5C6FCC7857DE5E9.taxon	description	Description (♁ and ♀). Head. Antennal, first flagellomere approximately 1.7 times as long as pedicel (Fig. 3 D). Gena grayish yellow (Fig. 14 D). Supracervical setae 6 – 9 and postoculars 13 – 15 per side (Fig. 4 D). Cibarial projections at anterolateral corners shorter than width of anterior margin; medial sensilla in parallel rows wider than sensilla campaniformia (Fig. 5 D 2). Prementum and neighboring lateral membrane with approximately 5 setae per side (Fig. 5 D). Thorax. Scutum and scutellum glossy, blackish brown; pleura grayish brown (Fig. 14 D). Legs. Fore tarsomere I as long as II + III (Fig. 9 D 1). Abdomen. Sternites grayish brown. Male terminalia. Cercus pubescent except for anterior to ventral margin, with approximately 32 setae. Female terminalia. Epigynium with 5 – 6 sensilla per side near lower posterior margin (Fig. 12 D 1). Measurements (holotype / range in 5 ♁ and 5 ♀, or less if noted, paratypes, in mm): BL = 1.51 / ♁ 1.30 – 1.62, ♀ 1.42 – 1.64 (4 ♀); ThL = 0.69 / ♁ 0.59 – 0.75, ♀ 0.59 – 0.71; WL = 1.35 / ♁ 1.20 – 1.44, ♀ 1.14 – 1.38; WW = 0.66 / ♁ 0.62 – 0.76, ♀ 0.54 – 0.72. Indices (holotype / range in 5 ♁ and 5 ♀, or less if noted, paratypes, in ratio): FW / HW = 0.56 / 0.56 – 0.62; ch / o = 0.34 / 0.34 – 0.39; rcorb = 0.44 / 0.47 – 0.60 (4 ♁ and 3 ♀); vb = 0.33 / 0.27 – 0.38; sctl = 0.62 / 0.61 – 0.74 (4 ♁ and 5 ♀); sctlp = 1.10 / 0.89 – 1.17; C = 1.44 / 1.28 – 1.47; 4 c = 1.74 / 1.69 – 2.13; 4 v = 2.57 / 2.60 – 3.15; 5 x = 4.61 / 4.02 – 5.11; ac = 3.05 / 2.72 – 3.57; M = 1.15 / 1.10 – 1.34.	en	Zhang, Guang, Gao, Jian-Jun, Takano, Kohei Takenaka, Yafuso, Masako, Suwito, Awit, Meleng, Paulus Ak, Toda, Masanori J. (2023): Phylogenetic classification and palm-inflorescence anthophily of the Colocasiomyia zeylanica species group (Diptera: Drosophilidae), with descriptions of five new species. Zootaxa 5278 (2): 201-238, DOI: 10.11646/zootaxa.5278.2.1, URL: http://dx.doi.org/10.11646/zootaxa.5278.2.1
03C687D9FFCFFFFAA5C6FCC7857DE5E9.taxon	materials_examined	Holotype. ♁ (MZB), “ Cibodas, West Java, Indonesia, 12. xii. 2004, K. T. Takano / ex inflorescence of Pinanga coronata ”. Paratypes. Indonesia: 5 ♁, 10 ♀, same data as the holotype, 7 ♁, 20 ♀, same data as the holotype except for 24. xii. 2003 (MZB, SEHU); 10 ♁, 10 ♀ (# 05180 – 99), same data as the holotype except for 24. xii. 2003 (KIZ).	en	Zhang, Guang, Gao, Jian-Jun, Takano, Kohei Takenaka, Yafuso, Masako, Suwito, Awit, Meleng, Paulus Ak, Toda, Masanori J. (2023): Phylogenetic classification and palm-inflorescence anthophily of the Colocasiomyia zeylanica species group (Diptera: Drosophilidae), with descriptions of five new species. Zootaxa 5278 (2): 201-238, DOI: 10.11646/zootaxa.5278.2.1, URL: http://dx.doi.org/10.11646/zootaxa.5278.2.1
03C687D9FFCFFFFAA5C6FCC7857DE5E9.taxon	distribution	Distribution. Indonesia (West Java).	en	Zhang, Guang, Gao, Jian-Jun, Takano, Kohei Takenaka, Yafuso, Masako, Suwito, Awit, Meleng, Paulus Ak, Toda, Masanori J. (2023): Phylogenetic classification and palm-inflorescence anthophily of the Colocasiomyia zeylanica species group (Diptera: Drosophilidae), with descriptions of five new species. Zootaxa 5278 (2): 201-238, DOI: 10.11646/zootaxa.5278.2.1, URL: http://dx.doi.org/10.11646/zootaxa.5278.2.1
03C687D9FFCFFFFAA5C6FCC7857DE5E9.taxon	etymology	Etymology. Referring to the lack of some setae from the head and thorax.	en	Zhang, Guang, Gao, Jian-Jun, Takano, Kohei Takenaka, Yafuso, Masako, Suwito, Awit, Meleng, Paulus Ak, Toda, Masanori J. (2023): Phylogenetic classification and palm-inflorescence anthophily of the Colocasiomyia zeylanica species group (Diptera: Drosophilidae), with descriptions of five new species. Zootaxa 5278 (2): 201-238, DOI: 10.11646/zootaxa.5278.2.1, URL: http://dx.doi.org/10.11646/zootaxa.5278.2.1
03C687D9FFCCFFFAA5C6FEBD8272E080.taxon	description	(Figs 2 – 12 E, 14 E)	en	Zhang, Guang, Gao, Jian-Jun, Takano, Kohei Takenaka, Yafuso, Masako, Suwito, Awit, Meleng, Paulus Ak, Toda, Masanori J. (2023): Phylogenetic classification and palm-inflorescence anthophily of the Colocasiomyia zeylanica species group (Diptera: Drosophilidae), with descriptions of five new species. Zootaxa 5278 (2): 201-238, DOI: 10.11646/zootaxa.5278.2.1, URL: http://dx.doi.org/10.11646/zootaxa.5278.2.1
03C687D9FFCCFFFAA5C6FEBD8272E080.taxon	diagnosis	Diagnosis. Antennal arista approximately 1.6 times as long as first flagellomere (Fig. 3 E). Distance between antennal scapes narrower than scape diameter (Fig. 2 E 2). Acrostichal setulae in 8 rows (Fig. 6 E). All legs dark grayish yellow except for mid and hind tarsomeres I – IVs grayish yellow (Fig. 14 E). Cercus roundish on ventral margin (Fig. 11 - 2 E). Long, tube-like, membranous aedeagus ornamented with dense, stronger spinules (Fig. 11 - 2 E). Hypogynial valves with roundish, dorsal flaps not fused to each other at distal end of basal portion (Fig. 12 E).	en	Zhang, Guang, Gao, Jian-Jun, Takano, Kohei Takenaka, Yafuso, Masako, Suwito, Awit, Meleng, Paulus Ak, Toda, Masanori J. (2023): Phylogenetic classification and palm-inflorescence anthophily of the Colocasiomyia zeylanica species group (Diptera: Drosophilidae), with descriptions of five new species. Zootaxa 5278 (2): 201-238, DOI: 10.11646/zootaxa.5278.2.1, URL: http://dx.doi.org/10.11646/zootaxa.5278.2.1
03C687D9FFCCFFFAA5C6FEBD8272E080.taxon	description	Description (♁ and ♀). Head. Antennal, first flagellomere approximately 1.3 times as long as pedicel (Fig. 3 E). Gena dark grayish brown (Fig. 14 E). Supracervical setae 4 – 5 and postoculars 17 – 18 per side (Fig. 4 E). Cibarial projections at anterolateral corners as long as width of anterior margin; medial sensilla in parallel rows as wide as sensilla campaniformia (Fig. 5 E 2). Prementum and neighboring lateral membrane with 6 – 7 setae per side (Fig. 5 E). Thorax glossy, blackish brown (Fig. 14 E). Legs. Fore tarsomere I slightly shorter than II + III (Fig. 9 E 1). Abdomen. Sternites dark grayish brown. Male terminalia. Cercus nearly entirely pubescent, with 32 – 34 setae. Female terminalia. Epigynium with approximately 7 small sensilla per side near lower posterior margin (Fig. 12 D 1). Measurements (holotype / range in 5 ♁ and 5 ♀, or less if noted, paratypes, in mm): BL = 1.92 / ♁ 1.45 – 1.70, ♀ 1.65 – 1.88 (4 ♀); ThL = 0.81 / ♁ 0.66 – 0.82, ♀ 0.78 – 0.84; WL = 1.56 / ♁ 1.30 – 1.54, ♀ 1.46 – 1.61; WW = 0.74 / ♁ 0.60 – 0.72, ♀ 0.69 – 0.76. Indices (holotype / range in 5 ♁ and 5 ♀, or less if noted, paratypes, in ratio): FW / HW = 0.58 / 0.55 – 0.60; ch / o = 0.40 / 0.39 – 0.46; rcorb = 0.38 / 0.43 – 0.55 (4 ♁ and 1 ♀); vb = 0.26 / 0.32 – 0.43; sctl = 0.56 / 0.58 – 0.70 (3 ♁ and 2 ♀); sctlp = 1.14 / 1.02 – 1.19; C = 1.46 / 1.30 – 1.43; 4 c = 1.71 / 1.67 – 2.25; 4 v = 2.94 / 2.52 – 3.58; 5 x = 7.14 / 5.07 – 7.56; ac = 2.98 / 3.12 – 3.63; M = 1.25 / 1.14 – 1.64.	en	Zhang, Guang, Gao, Jian-Jun, Takano, Kohei Takenaka, Yafuso, Masako, Suwito, Awit, Meleng, Paulus Ak, Toda, Masanori J. (2023): Phylogenetic classification and palm-inflorescence anthophily of the Colocasiomyia zeylanica species group (Diptera: Drosophilidae), with descriptions of five new species. Zootaxa 5278 (2): 201-238, DOI: 10.11646/zootaxa.5278.2.1, URL: http://dx.doi.org/10.11646/zootaxa.5278.2.1
03C687D9FFCCFFFAA5C6FEBD8272E080.taxon	materials_examined	Holotype. ♁ (MZB), “ Cibodas, West Java, Indonesia, 12. xii. 2004, K. T. Takano / ex inflorescence of Pinanga coronata ”. Paratypes. Indonesia: 9 ♁, 23 ♀, same data as the holotype, 14 ♁, 13 ♀, same data as the holotype except for 24. xii. 2003 (MZB, SEHU); 10 ♁, 10 ♀ (# 05202 – 21), same data as the holotype except for 24. xii. 2003 (KIZ).	en	Zhang, Guang, Gao, Jian-Jun, Takano, Kohei Takenaka, Yafuso, Masako, Suwito, Awit, Meleng, Paulus Ak, Toda, Masanori J. (2023): Phylogenetic classification and palm-inflorescence anthophily of the Colocasiomyia zeylanica species group (Diptera: Drosophilidae), with descriptions of five new species. Zootaxa 5278 (2): 201-238, DOI: 10.11646/zootaxa.5278.2.1, URL: http://dx.doi.org/10.11646/zootaxa.5278.2.1
03C687D9FFCCFFFAA5C6FEBD8272E080.taxon	distribution	Distribution. Indonesia (West Java).	en	Zhang, Guang, Gao, Jian-Jun, Takano, Kohei Takenaka, Yafuso, Masako, Suwito, Awit, Meleng, Paulus Ak, Toda, Masanori J. (2023): Phylogenetic classification and palm-inflorescence anthophily of the Colocasiomyia zeylanica species group (Diptera: Drosophilidae), with descriptions of five new species. Zootaxa 5278 (2): 201-238, DOI: 10.11646/zootaxa.5278.2.1, URL: http://dx.doi.org/10.11646/zootaxa.5278.2.1
03C687D9FFCCFFFAA5C6FEBD8272E080.taxon	discussion	Remarks. This species resembles C. oligochaeta sp. nov., but can be distinguished from it by the diagnostic characters.	en	Zhang, Guang, Gao, Jian-Jun, Takano, Kohei Takenaka, Yafuso, Masako, Suwito, Awit, Meleng, Paulus Ak, Toda, Masanori J. (2023): Phylogenetic classification and palm-inflorescence anthophily of the Colocasiomyia zeylanica species group (Diptera: Drosophilidae), with descriptions of five new species. Zootaxa 5278 (2): 201-238, DOI: 10.11646/zootaxa.5278.2.1, URL: http://dx.doi.org/10.11646/zootaxa.5278.2.1
03C687D9FFCCFFFAA5C6FEBD8272E080.taxon	etymology	Etymology. Patronym, in honor of Dr. David Grimaldi at the American Museum of Natural History, who made early contributions to the systematics of Colocasiomyia.	en	Zhang, Guang, Gao, Jian-Jun, Takano, Kohei Takenaka, Yafuso, Masako, Suwito, Awit, Meleng, Paulus Ak, Toda, Masanori J. (2023): Phylogenetic classification and palm-inflorescence anthophily of the Colocasiomyia zeylanica species group (Diptera: Drosophilidae), with descriptions of five new species. Zootaxa 5278 (2): 201-238, DOI: 10.11646/zootaxa.5278.2.1, URL: http://dx.doi.org/10.11646/zootaxa.5278.2.1
