taxonID	type	description	language	source
03C687E9FFB4BF7BA0F6091DCD50F845.taxon	type_taxon	Type species: Dactylopius pini Kuwana 1902.	en	Tanaka, Hirotaka, Kamitani, Satoshi (2022): Review ofthegenus CrisicoccusFerris (Hemiptera: Coccomorpha: Pseudococcidae) in Japan with description of a new species, and the identity of a South Korean mealybug misidentified as Crisicoccus matsumotoi (Shiraiwa 1935). Zootaxa 5209 (5): 555-572, DOI: 10.11646/zootaxa.5209.5.3
03C687E9FFB4BF7BA0F6091DCD50F845.taxon	diagnosis	Genus diagnosis (adapted and modified from Williams 2004). Body of adult female normally broadly oval; anal lobe developed, each lobe with an anal lobe bar and bar seta present on ventral surface. Antennae each usually with 7 or 8 segments. Legs well developed, translucent pores usually present (rarely absent) on hind coxae, present also on hind tibiae and sometimes on hind femora. Tarsal digitules knobbed. Circulus normally present. Anterior and posterior ostioles present. Anal ring normal, usually bearing 6 setae. Cerarii numbering 0 – 17 pairs, each containing 2 conical setae or sometimes reduced to a single conical seta; often cerarii on head and thorax indistinct; preocular cerarii (C 2) absent. Auxiliary setae rarely present except on anal lobe cerarii. Body setae often flagellate, sometimes conical or spiniform. Trilocular pores normal. Multilocular disc pores present, usually confined to venter. Oral rim tubular ducts absent. Oral collar tubular ducts present on venter and sometimes also on dorsum.	en	Tanaka, Hirotaka, Kamitani, Satoshi (2022): Review ofthegenus CrisicoccusFerris (Hemiptera: Coccomorpha: Pseudococcidae) in Japan with description of a new species, and the identity of a South Korean mealybug misidentified as Crisicoccus matsumotoi (Shiraiwa 1935). Zootaxa 5209 (5): 555-572, DOI: 10.11646/zootaxa.5209.5.3
03C687E9FFB4BF7BA0F6091DCD50F845.taxon	discussion	Remarks. According to Williams (2004), Crisicoccus is considered a convenient genus to include any species possessing well-developed anal lobe bars that do not fit the features of other mealybug genera with anal lobe bars. Crisicoccus is quite similar to Formicococcus Takahashi 1928 and Planococcus Ferris 1950 and so it may be difficult to distinguish these genera. Further morphological and molecular phylogenetic studies of these three genera are greatly needed.	en	Tanaka, Hirotaka, Kamitani, Satoshi (2022): Review ofthegenus CrisicoccusFerris (Hemiptera: Coccomorpha: Pseudococcidae) in Japan with description of a new species, and the identity of a South Korean mealybug misidentified as Crisicoccus matsumotoi (Shiraiwa 1935). Zootaxa 5209 (5): 555-572, DOI: 10.11646/zootaxa.5209.5.3
03C687E9FFB7BF7FA0F60C9CC93FFDD3.taxon	vernacular_names	[Japanese common name: Matsumoto-kona-kaigaramushi]	en	Tanaka, Hirotaka, Kamitani, Satoshi (2022): Review ofthegenus CrisicoccusFerris (Hemiptera: Coccomorpha: Pseudococcidae) in Japan with description of a new species, and the identity of a South Korean mealybug misidentified as Crisicoccus matsumotoi (Shiraiwa 1935). Zootaxa 5209 (5): 555-572, DOI: 10.11646/zootaxa.5209.5.3
03C687E9FFB7BF7FA0F60C9CC93FFDD3.taxon	materials_examined	Material examined. Neotype, here designated: JAPAN / Tokyo, Tachikawa, / Fujimi-cho, / on Zelkova serrata, / 8. v. 1962, / coll. S. Kawai; an adult female mounted with other 9 adult females on a slide (KTUA). The individual at the left end of the middle row of specimens was selected as the neotype and is clearly indicated on the slide with a black ink dot (Fig. 1). Other material. JAPAN: Tokyo, Chuo-ku, Hamarikyuteien, on Mallotus japonicus, 15. v. 1972, coll. S. Kawai, 6 adult females mounted on 2 slides (KTUA); Shizuoka prefecture, Shimizu, Okitsu, on Diospyros kaki, 8. vii. 1976, coll. S. Kawai, an adult female mounted singly (KTUA); Shimane prefecture, Izumo, on Vitis sp., 25. v. 2001, coll. S. Narai, an adult female mounted singly (KTUA); Ibaraki prefecture, Kasama, Ago, on Pyrus pyrifolia var. culta, 27. iv. 2022, coll. T. Tsunoda, 6 adult females mounted singly (3 ELKU, 3 EUMJ).	en	Tanaka, Hirotaka, Kamitani, Satoshi (2022): Review ofthegenus CrisicoccusFerris (Hemiptera: Coccomorpha: Pseudococcidae) in Japan with description of a new species, and the identity of a South Korean mealybug misidentified as Crisicoccus matsumotoi (Shiraiwa 1935). Zootaxa 5209 (5): 555-572, DOI: 10.11646/zootaxa.5209.5.3
03C687E9FFB7BF7FA0F60C9CC93FFDD3.taxon	description	Updated description Appearance in life. Adult female 3 ‒ 4 mm long, dark purple covered with a white powdery wax. Projections of the wax secretion from body margin not so developed and limited to a few segments of posterior part of body. The body contents of this species turn blue black to dark green in 10 % potassium hydroxide solution (Kawai 1980, translated by HT). Slide-mounted adult female (Fig. 2) (n = 15). Body elongate oval, 3.6 (1.9 – 4.5) mm long and 2.2 (1.2 – 3.0) mm wide; derm membranous; segmentation recognizable, but not well developed. Anal lobes well developed, each with a long apical seta, 188 – 242 (145 – 270) µm long. Each lobe with a narrow and slightly faint anal lobe bar on ventral surface, but bar sometimes difficult to see in some specimens. Antenna 368 – 369 (368 – 500) µm long, with 8 segments and many flagellate setae; subapical segment with 1 fleshy seta and apical segment with 3 fleshy setae. Eyespot present on margin, not associated with discoidal pores. Legs well developed, with many flagellate setae; hind leg measurements (in µm): trochanter + femur 269 – 270 (269 – 394) long; hind tibia + tarsus 281 – 286 (281 – 377); claw 39 – 42 (34 – 45), without a denticle. Ratio of lengths of hind tibia + tarsus to trochanter + femur about 1: 1.04 – 1.06 (1: 0.89 – 1.06); ratio of lengths of hind tibia to tarsus 1: 1.78 – 1.83 (1: 1.78 – 2.60). Paired setose tarsal digitules present, subequal in length to the minutely knobbed claw digitules. Hind coxa with translucent pores on both surfaces; hind tibia usually with translucent pores on posterior surface; hind trochanter, femur and tarsus without translucent pores. Labium 170 (127 – 185) µm long, mostly shorter than clypeolabral shield. Circulus oval to quadrate, located between abdominal segments III and IV and divided by an intersegmental line, 70 (50 – 170) µm long and 90 (48 – 165) µm wide. Ostioles present, each with inner edges of lips weakly sclerotized; anterior ostioles each with a total for both lips of 22 – 27 (11 – 59) trilocular pores and 5 (2 – 7) setae; each posterior ostiole with a total for both lips of 24 – 27 (19 – 62) trilocular pores and 5 – 8 (2 – 8) setae. Anal ring 94 (90 – 113) µm wide, bearing 6 (6 – 8) setae, each seta 112 – 145 (60 – 170) µm long. Cerarii usually numbering 4 or 5 (4 – 8) pairs, mostly present on posterior abdominal segments, rarely present also on thoracic segments. Anal lobe cerarii (C 18) each containing 2 conical cerarian setae, each seta 17 – 20 (9 – 23) µm long and about 7 (5 – 8) µm wide at base, 4 – 5 (4 – 8) auxiliary setae and a concentration of trilocular pores. Penultimate cerarii (C 17) each containing 2 conical setae, 3 (0 – 8) auxiliary setae and a few trilocular pores. Cerarii situated further forward generally each with 2 conical setae with a few trilocular pores and auxiliary setae. Dorsum. Setae flagellate, each 17 – 70 µm long, distributed usually segmentally; longest setae present on head. Trilocular pores each 3 – 4 µm wide, evenly distributed. Oral rim tubular ducts and oral collar tubular ducts absent. Discoidal pores each about 2 – 3 (1 – 4) µm wide, sparsely distributed. Venter. Setae relatively long and flagellate, each 32 – 90 (19 – 168) µm long; setae on head longest. Multilocular disc pores, each 6 – 8 (6 – 9) µm wide, present in medial areas of abdominal segments IV ‒ IX, arranged in single row on each posterior area of abdominal segments IV ‒ V, in 2 or 3 rows on each posterior area of abdominal segments VI – VII, and in 1 – 2 rows on anterior areas of abdominal segments VI – VII but arranged randomly on abdominal segments VIII and IX. Trilocular pores, each same size as those on dorsum, evenly distributed. Oral collar tubular ducts of 2 sizes present: (i) large-type ducts, each about 3 – 4 µm in diameter, mostly wider than a trilocular pore, present on marginal to submarginal areas of all abdominal and thoracic segments and on medial areas of abdominal segments IV – VII, also forming transverse bands across abdominal segments IV – VII; and (ii) small-type ducts, each about 2 – 3 μm in diameter, mostly smaller than a trilocular pore, present on medial areas of abdominal segments IV – VII intermixed with large-type ducts, forming transverse bands across abdominal segments IV – VII. Discoidal pores, same size as those on dorsum, sparsely present.	en	Tanaka, Hirotaka, Kamitani, Satoshi (2022): Review ofthegenus CrisicoccusFerris (Hemiptera: Coccomorpha: Pseudococcidae) in Japan with description of a new species, and the identity of a South Korean mealybug misidentified as Crisicoccus matsumotoi (Shiraiwa 1935). Zootaxa 5209 (5): 555-572, DOI: 10.11646/zootaxa.5209.5.3
03C687E9FFB7BF7FA0F60C9CC93FFDD3.taxon	biology_ecology	Host plants in Japan. Buxaceae: Buxus microphylla (Kawai 1980); Cannabaceae: Celtis sinensis (Kawai 1980); Ebenaceae: Diospyros kaki (Kawai 1980; Kawai 2003); Euphorbiaceae: Mallotus japonicus (Kawai 1980); Fabaceae: Wisteria floribunda (Kawai 2003); Juglandaceae: Juglans mandshurica (Kawai 1980); Moraceae: Ficus carica (Kawai 1980; Kawai 2003); Platanaceae: Platanus orientalis (Kawai 1980); Rosaceae: Chaenomeles speciosa (Kawai 1980), Photinia glabra (Kawai 2003), Pyrus pyrifolia var. culta (Kawai 1980; Kawai 2003); Sapindaceae: Acer palmatum (Kawai 1980), Acer spp. (Kawai 2003); Ulmaceae: Zelkova serrata (Kawai 1980); Vitaceae: Vitis spp. (Kawai 2003).	en	Tanaka, Hirotaka, Kamitani, Satoshi (2022): Review ofthegenus CrisicoccusFerris (Hemiptera: Coccomorpha: Pseudococcidae) in Japan with description of a new species, and the identity of a South Korean mealybug misidentified as Crisicoccus matsumotoi (Shiraiwa 1935). Zootaxa 5209 (5): 555-572, DOI: 10.11646/zootaxa.5209.5.3
03C687E9FFB7BF7FA0F60C9CC93FFDD3.taxon	discussion	Remarks. The original description of C. seruratus by Kanda (1933) was based mostly on highly variable morphological characteristics (including proportions of antennal segment lengths) that have little taxonomic value. Most of Kanda’s specimens are deposited in Osaka Museum of Natural History, Japan (OMNH); however, a lengthy search was made for type specimens of this species in almost all of the Coccomorpha collections in Japan (including Osaka Museum of Natural History) but none could be found; we therefore conclude that all the type specimens of C. seruratus described by Kanda have been lost. The designation of a neotype of this species was necessary because the original description is not informative, complicated taxonomic problems are associated with the species, and the type material has been lost. One of the specimens used in this redescription is designated the neotype (above) for taxonomic stability. The neotype specimen was obtained from a tree of Zelkova serrata in the Kanto region of Japan, which is the same area where the primary type series for C. seruratus were collected. The morphology of this type specimen agrees well with the original description. Currently, in Japan C. seruratus is recognized as C. matsumotoi. Its morphology differs significantly from “ C. matsumotoi ” reported by Ezzat & McConnell (1956) and Williams (2004), by having the following morphological characteristics (contrasting characteristics of the material incorrectly identified as “ C. matsumotoi ” are stated in parentheses): (i) a small number of cerarii, numbering fewer than 8 pairs (cerarii numbering 15 – 16 pairs); and (ii) oral collar tubular ducts of two different sizes present on venter (venter with only one size of oral collar tubular ducts). Therefore, in this study, we describe “ C. matsumotoi ” reported by Ezzat & McConnell (1956) and Williams (2004) as a distinct, new species, Crisicoccus ezzati sp. nov. (see below). Crisicoccus seruratus has been collected on the roots of Zelkova serrata (= Abelicea serrata), which is the host plant of Kanda’s original collection, but he used the name Abelicea serrata and he misspelt the scientific name of the host as “ Obelicea serurata ”. Kawai's (1980) view that C. matsumotoi and C. seruratus are the same species seems to be correct; however, he incorrectly designated C. seruratus (the older name) as a junior synonym of C. matsumotoi (García Morales et al. 2016). Above, we formally synonymize Pseudococcus matsumotoi Siraiwa 1935 as a junior synonym of Pseudococcus seruratus Kanda. We were unable to examine any types of Pseudococcus astericola Shinji 1936; however, Kanda (1941) synonymized this species under Pseudococcus seruratus Kanda and we follow his synonymy. In Korea, Paik (1978) and Kwon et al. (2003) regarded C. seruratus and “ C. matsumotoi ” as distinct species; however, the species they recognized as “ C. matsumotoi ” in Korea was a misidentification of Spilococcus pacificus (see below). Crisicoccus species do not have oral rim tubular ducts on the body surface, whereas Spilococcus species have at least a few so the genera can be easily distinguished from each other. Crisicoccus seruratus shows similarities to C. melaleucae Williams 1985 (described from Queensland, Australia), such as possessing a small number of cerarii (≤ eight pairs), oral collar tubular ducts of two sizes, and lacking cerarii on the head; however, it differs from C. melaleucae in having relatively long dorsal setae, each approximately 17 – 70 µm long, whereas C. melaleucae has relatively short dorsal setae, each approximately 8 – 16 µm long.	en	Tanaka, Hirotaka, Kamitani, Satoshi (2022): Review ofthegenus CrisicoccusFerris (Hemiptera: Coccomorpha: Pseudococcidae) in Japan with description of a new species, and the identity of a South Korean mealybug misidentified as Crisicoccus matsumotoi (Shiraiwa 1935). Zootaxa 5209 (5): 555-572, DOI: 10.11646/zootaxa.5209.5.3
03C687E9FFB0BF7DA0F60E60C9A0FC97.taxon	vernacular_names	[Japanese common name: Nashi-kona-kaigaramushi]	en	Tanaka, Hirotaka, Kamitani, Satoshi (2022): Review ofthegenus CrisicoccusFerris (Hemiptera: Coccomorpha: Pseudococcidae) in Japan with description of a new species, and the identity of a South Korean mealybug misidentified as Crisicoccus matsumotoi (Shiraiwa 1935). Zootaxa 5209 (5): 555-572, DOI: 10.11646/zootaxa.5209.5.3
03C687E9FFB0BF7DA0F60E60C9A0FC97.taxon	materials_examined	Material examined. Holototype, here designated: Pseudococcus / Crisicoccus / matsumotoi / (Shir.) / On Pears / Japan: at Seattle / Berryhill. Scott. Collr’s / Sept. 12. 1940 / Seattle 9045. 1 adult female mounted singly on a slide (USNM).	en	Tanaka, Hirotaka, Kamitani, Satoshi (2022): Review ofthegenus CrisicoccusFerris (Hemiptera: Coccomorpha: Pseudococcidae) in Japan with description of a new species, and the identity of a South Korean mealybug misidentified as Crisicoccus matsumotoi (Shiraiwa 1935). Zootaxa 5209 (5): 555-572, DOI: 10.11646/zootaxa.5209.5.3
03C687E9FFB0BF7DA0F60E60C9A0FC97.taxon	description	Description based on the holotype only. Appearance in life. Not seen. Slide-mounted adult female (Fig. 3). Body elongate oval, 2.0 mm long and 1.0 mm wide; derm membranous; segmentation recognizable, but not well developed. Anal lobes well developed, ventral surface with rather faint anal lobe bar and a long apical seta, 226 – 254 µm long. Antenna 399 – 410 µm long, with 8 segments and many flagellate setae; subapical segment with 1 fleshy seta and apical segment with 3 fleshy setae. Eyespot present on margin, not associated with discoidal pores. Legs well developed, with many flagellate setae; hind leg measurements (in µm): trochanter + femur 304 – 307 long; hind tibia + tarsus 304 – 312; claw 34 – 35, without a denticle. Ratio of lengths of hind tibia + tarsus to trochanter + femur about 1: 0.99 – 1.03; ratio of lengths of hind tibia to tarsus 1: 2.12 – 2.23. Paired setose tarsal digitules present, subequal in length to the minutely knobbed claw digitules. Hind coxa with translucent pores on both surfaces; hind tibia with translucent pores on posterior surface; hind trochanter, femur and tarsus without translucent pores. Labium 218 µm long, shorter than clypeolabral shield. Circulus usually quadrate, located between abdominal segments III and IV, 76 µm long and 109 µm wide. Ostioles present, each with inner edges of lips weakly sclerotized; anterior ostioles each with a total for both lips of 23 trilocular pores and 2 setae; each posterior ostiole with a total for both lips of 25 – 28 trilocular pores and 3 – 5 setae. Anal ring 95 µm wide, bearing 6 setae, each seta 108 – 138 µm long. Cerarii numbering 15 – 16 pairs. Anal lobe cerarii (C 18) usually each containing 2 conical cerarian setae, each seta 20 – 22 µm long and about 5 – 6 µm wide at base, 6 – 7 auxiliary setae and a concentration of trilocular pores. Penultimate cerarii (C 17) each containing 2 conical setae and a few auxiliary setae and trilocular pores. Cerarii situated further forward generally each with 2 – 4 conical setae, usually with flagellate tips, and trilocular pores. Dorsum. Setae flagellate, each 14 – 48 µm long, distributed segmentally; longest setae present on head. Trilocular pores each 3 – 4 µm wide, evenly distributed. Oral rim tubular ducts and oral collar tubular ducts absent. Discoidal pores each about 2 µm wide, sparsely distributed. Venter. Setae relatively long and flagellate, each 18 – 89 µm long; setae on medial area of posterior abdominal segments longest. Multilocular disc pores, each 6 – 8 µm wide, present in medial areas of abdominal segments IV ‒ IX, arranged in 1 (or rarely 2) rows on each posterior area of abdominal segments IV – VII but arranged randomly on abdominal segments VIII and IX. Trilocular pores, each same size as those on dorsum, evenly distributed. Oral collar tubular ducts all of 1 size, each narrower than a trilocular pore, about 2 – 3 µm wide, present on medial areas of abdominal segments IV – VII and submarginal to marginal areas of abdominal segments II – VIII, usually forming transverse bands across segments; additionally, a few ducts present on submarginal to marginal areas of thoracic segments. Discoidal pores, same size as those on dorsum, sparsely present.	en	Tanaka, Hirotaka, Kamitani, Satoshi (2022): Review ofthegenus CrisicoccusFerris (Hemiptera: Coccomorpha: Pseudococcidae) in Japan with description of a new species, and the identity of a South Korean mealybug misidentified as Crisicoccus matsumotoi (Shiraiwa 1935). Zootaxa 5209 (5): 555-572, DOI: 10.11646/zootaxa.5209.5.3
03C687E9FFB0BF7DA0F60E60C9A0FC97.taxon	biology_ecology	Host plant in Japan. Rosaceae: Pyrus pyrifolia var. culta.	en	Tanaka, Hirotaka, Kamitani, Satoshi (2022): Review ofthegenus CrisicoccusFerris (Hemiptera: Coccomorpha: Pseudococcidae) in Japan with description of a new species, and the identity of a South Korean mealybug misidentified as Crisicoccus matsumotoi (Shiraiwa 1935). Zootaxa 5209 (5): 555-572, DOI: 10.11646/zootaxa.5209.5.3
03C687E9FFB0BF7DA0F60E60C9A0FC97.taxon	discussion	Remarks. Crisicoccus ezzati sp. nov. was first reported as C. matsumotoi by Ezzat & McConnell (1956), based on specimens from Japan intercepted at a US plant quarantine station and three specimens collected in Fukuoka prefecture, Japan. Some specimens collected later from India and Philippines were reported by Williams (2004). In this study, we examined a specimen identified by Ezzat & McConnell (1956) and found that the species is quite different from that currently considered to be C. matumotoi (= Crisicoccus seruratus) in Japan. Crisicoccus ezzati has (contrasting characteristics of C. seruratus are stated in parentheses): (i) cerarii numbering 15 – 16 pairs (no more than eight pairs of cerarii); and (ii) only one size of oral collar tubular ducts on the venter (two sizes of oral collar tubular ducts on the venter). Judging the validity of this species being labelled “ C. matsumotoi ” is difficult because the type specimens for C. matsumotoi are lost (García Morales et al. 2016). However, in this study, we followed Dr Shozo Kawai, Professor Emeritus of Tokyo University of Agriculture, who studied Japanese mealybugs taxonomically for a long time and treated the species currently recognized as C. matsumotoi in Japan as the true C. matsumotoi (= C. seruratus). Therefore, the species described by Ezzat & McConnell (1956) and Williams (2004) lacks a valid name and is here described as a new species. Crisicoccus ezzati bears some similarities to C. orchidiradicis (Takahashi 1951) from Malaysia in having a large number of cerarii (≥ 15 pairs) and anterior cerarii with relatively narrow, elongated cerarian setae. However, it differs by the following morphological characteristics (contrasting characteristics of C. orchidiradicis are given in parentheses): (i) 15 – 16 pairs of cerarii (17 pairs); (ii) translucent pores present on hind coxae (without translucent pores on hind coxae), and (iii) anterior cerarii lacking auxiliary setae (all cerarii containing auxiliary setae).	en	Tanaka, Hirotaka, Kamitani, Satoshi (2022): Review ofthegenus CrisicoccusFerris (Hemiptera: Coccomorpha: Pseudococcidae) in Japan with description of a new species, and the identity of a South Korean mealybug misidentified as Crisicoccus matsumotoi (Shiraiwa 1935). Zootaxa 5209 (5): 555-572, DOI: 10.11646/zootaxa.5209.5.3
03C687E9FFB0BF7DA0F60E60C9A0FC97.taxon	etymology	Etymology. The new species is named after Dr Yehia Mahmoud Ezzat, who was one of the first to report on it.	en	Tanaka, Hirotaka, Kamitani, Satoshi (2022): Review ofthegenus CrisicoccusFerris (Hemiptera: Coccomorpha: Pseudococcidae) in Japan with description of a new species, and the identity of a South Korean mealybug misidentified as Crisicoccus matsumotoi (Shiraiwa 1935). Zootaxa 5209 (5): 555-572, DOI: 10.11646/zootaxa.5209.5.3
03C687E9FFB2BF73A0F60FA4CD5CF9C3.taxon	vernacular_names	[Japanese common name: Azarea-kona-kaigaramushi]	en	Tanaka, Hirotaka, Kamitani, Satoshi (2022): Review ofthegenus CrisicoccusFerris (Hemiptera: Coccomorpha: Pseudococcidae) in Japan with description of a new species, and the identity of a South Korean mealybug misidentified as Crisicoccus matsumotoi (Shiraiwa 1935). Zootaxa 5209 (5): 555-572, DOI: 10.11646/zootaxa.5209.5.3
03C687E9FFB2BF73A0F60FA4CD5CF9C3.taxon	materials_examined	Material examined. Japan: Tokyo, Akikawa-shi, Nobe, on Rhododendron amagianum, 5. viii. 1972, coll. S. Kawai, 3 adult females mounted singly and 2 adult females mounted together on a slide (KTUA); Ibaraki Prefecture, Tsukuba, Kannon-dai, on Rhododendron x pulchrum, 21. iv. 2021, coll. J. Tabata, 4 adult females mounted singly (2 ELKU, 2 EUMJ).	en	Tanaka, Hirotaka, Kamitani, Satoshi (2022): Review ofthegenus CrisicoccusFerris (Hemiptera: Coccomorpha: Pseudococcidae) in Japan with description of a new species, and the identity of a South Korean mealybug misidentified as Crisicoccus matsumotoi (Shiraiwa 1935). Zootaxa 5209 (5): 555-572, DOI: 10.11646/zootaxa.5209.5.3
03C687E9FFB2BF73A0F60FA4CD5CF9C3.taxon	description	Updated description Appearance in life. Adult female 3 ‒ 4 mm long, dark purple to purple-brown, covered with a white powdery wax. Projections of wax secretion from body margin short and indistinct on cephalothorax, slightly longer on a few segments of posterior part of body. Body contents of this species turn blue-black to dark green in 10 % potassium hydroxide solution (Kawai 1980, translated by HT). Slide-mounted adult female (Fig. 4) (n = 9). Body elongate oval, 1.9 – 3.1 mm long and 0.9 – 1.9 mm wide; derm membranous; segmentation recognizable but not well developed. Anal lobes well developed, ventral surface with an anal lobe bar and a long apical seta, 154 – 239 µm long. Antenna 334 – 432 µm long, with 8 segments and many flagellate setae; subapical segment with 1 fleshy seta, apical segment with 3 fleshy setae. Eyespot present on margin, not associated with discoidal pores. Legs well developed, with many flagellate setae; hind leg measurements (in µm): trochanter + femur 248 – 320 long; hind tibia + tarsus 283 – 350; claw 26 – 36, without a denticle. Ratio of lengths of hind tibia + tarsus to trochanter + femur about 1: 1.04 – 1.16; ratio of lengths of hind tibia to tarsus 1: 1.79 – 2.40. Paired setose tarsal digitules present, subequal in length to the minutely knobbed claw digitules. Hind coxa with translucent pores on both surfaces; hind tibia sometimes with translucent pores on posterior surface; hind trochanter, femur and tarsus without translucent pores. Labium 120 – 152 µm long, shorter than clypeolabral shield. Circulus usually oval but rarely quadrate, located between abdominal segments III and IV or rarely on abdominal segment IV, usually divided by an intersegmental line, 40 – 113 µm long and 84 – 120 µm wide. Ostioles present, each with inner edges of lips weakly sclerotized; anterior ostioles each with a total for both lips of 17 – 29 trilocular pores and 2 – 7 setae; each posterior ostiole with a total for both lips of 16 – 44 trilocular pores and 4 – 7 setae. Anal ring 84 – 112 µm wide, bearing 6 setae, each seta 104 – 160 µm long. Cerarii numbering 6 – 12 pairs, normally present on posterior abdominal segments, rarely present also on thoracic segments. Anal lobe cerarii (C 18) each containing mostly 2 (rarely 3) conical cerarian setae, each 14 – 22 µm long and about 5 – 7 µm wide at base; 3 – 6 auxiliary setae, and a concentration of trilocular pores. Penultimate cerarii (C 17) each containing 2 conical setae, 0 auxiliary setae and a few trilocular pores. Cerarii situated further forward generally each with 2 conical cerarian setae and a few trilocular pores. Dorsum. Setae spiniform, mostly straight, each 7 – 30 µm long, usually distributed segmentally; longest setae present on head or posterior abdominal segments. Trilocular pores each 3 – 4 µm wide, evenly distributed. Oral rim tubular ducts and oral collar tubular ducts absent. Discoidal pores each about 2 µm wide, sparsely distributed. Venter. Setae relatively long and flagellate, each 11 – 140 µm long; setae on head longest. Multilocular disc pores, each 6 – 9 µm wide, present in medial areas of abdominal segments IV ‒ IX, arranged in single rows on each posterior area of abdominal segments IV ‒ V, 2 or 3 rows on each posterior area of abdominal segments VI – VII, and in a single row on each anterior area of abdominal segments VI – VII but arranged randomly on abdominal segments VIII and IX. Trilocular pores, each of same size as those on dorsum, evenly distributed. Oral collar tubular ducts all of 1 size, each about 2 – 4 µm wide, present on medial areas of abdominal segments IV – VII and usually forming transverse bands across segments; also relatively slightly stouter tubular ducts present on submarginal to marginal areas of abdominal segments I – VIII and thoracic segments. Discoidal pores, same width as those on dorsum, sparsely present.	en	Tanaka, Hirotaka, Kamitani, Satoshi (2022): Review ofthegenus CrisicoccusFerris (Hemiptera: Coccomorpha: Pseudococcidae) in Japan with description of a new species, and the identity of a South Korean mealybug misidentified as Crisicoccus matsumotoi (Shiraiwa 1935). Zootaxa 5209 (5): 555-572, DOI: 10.11646/zootaxa.5209.5.3
03C687E9FFB2BF73A0F60FA4CD5CF9C3.taxon	biology_ecology	Host plants in Japan. Ebenaceae: Diospyros kaki (Kawai 1980; 2003); Ericaceae: Rhododendron amagianum (Kawai 1980), Rhododendron macrosepalum (Kawai 1980), Rhododendron spp. (Kawai 2003), Rhododendron x pulchrum; Fabaceae: Albizia julibrissin (Kawai 1980); Fagaceae: Castanopsis cuspidata (Kawai 1980); Magnoliaceae: Magnolia kobus (Kawai 1980); Oleaceae: Ligustrum lucidum (Kawai 1980; 2003); Rosaceae: Eriobotrya japonica (Kawai 1980; 2003), Pyracantha angustifolia (Kawai 1980), Pyrus pyrifolia var. culta (Kawai 1980; 2003); Sapindaceae: Acer buergerianum (Kawai 1980; 2003); Schisandraceae: Illicium anisatum (Kawai 1980); Salicaceae: Salix babylonica var. babylonica (Kawai 1980); Taxaceae: Cephalotaxus harringtonia (Kanda 1943), Taxus cuspidata (Kawai 1980).	en	Tanaka, Hirotaka, Kamitani, Satoshi (2022): Review ofthegenus CrisicoccusFerris (Hemiptera: Coccomorpha: Pseudococcidae) in Japan with description of a new species, and the identity of a South Korean mealybug misidentified as Crisicoccus matsumotoi (Shiraiwa 1935). Zootaxa 5209 (5): 555-572, DOI: 10.11646/zootaxa.5209.5.3
03C687E9FFB2BF73A0F60FA4CD5CF9C3.taxon	discussion	Remarks. Crisicoccus azaleae is similar to C. pini (Kuwana 1902) in having a small number of cerarii (<13 pairs) and oral collar tubular ducts of one size only. However, it differs from C. pini in the following morphological characteristics (contrasting characteristics of C. pini are given in parentheses): (i) a circulus usually present between abdominal segments III and IV on the venter (circulus always lacking); (ii) presence of spiniform and mostly straight dorsal setae (dorsal setae are mostly flagellate and slightly curved); and (iii) thoracic segments with a few ventral oral collar tubular ducts (thoracic segments without ventral oral collar tubular ducts). In particular, C. azaleae is characterized by straight, spiniform dorsal setae, which are significantly different from the curved flagellate setae in C. pini. Danzig & Gavrilov-Zimin (2015) stated that “ Crisicoccus azaleae (Tinsley 1898) is very similar, probably conspecific with C. pini ”. However, both species are clearly different and are treated here as wellseparated species.	en	Tanaka, Hirotaka, Kamitani, Satoshi (2022): Review ofthegenus CrisicoccusFerris (Hemiptera: Coccomorpha: Pseudococcidae) in Japan with description of a new species, and the identity of a South Korean mealybug misidentified as Crisicoccus matsumotoi (Shiraiwa 1935). Zootaxa 5209 (5): 555-572, DOI: 10.11646/zootaxa.5209.5.3
03C687E9FFBCBF76A0F60A50CD02FD06.taxon	vernacular_names	[Japanese common name: Matsu-kona-kaigaramushi]	en	Tanaka, Hirotaka, Kamitani, Satoshi (2022): Review ofthegenus CrisicoccusFerris (Hemiptera: Coccomorpha: Pseudococcidae) in Japan with description of a new species, and the identity of a South Korean mealybug misidentified as Crisicoccus matsumotoi (Shiraiwa 1935). Zootaxa 5209 (5): 555-572, DOI: 10.11646/zootaxa.5209.5.3
03C687E9FFBCBF76A0F60A50CD02FD06.taxon	materials_examined	Material examined. Japan: Shizuoka Prefecture, Shimizu, Okitsu, on Pinus sp., 2. vi. 1972, coll. S. Kawai, 6 adult females mounted on 2 slides (KTUA); Fukuoka Prefecture, Fukuoka, Nishi-ku, Ikinomatsubara, on Pinus thunbergii, 9. v. 2021, coll. H. Tanaka, 14 adult females mounted singly (7 ELKU, 7 EUMJ).	en	Tanaka, Hirotaka, Kamitani, Satoshi (2022): Review ofthegenus CrisicoccusFerris (Hemiptera: Coccomorpha: Pseudococcidae) in Japan with description of a new species, and the identity of a South Korean mealybug misidentified as Crisicoccus matsumotoi (Shiraiwa 1935). Zootaxa 5209 (5): 555-572, DOI: 10.11646/zootaxa.5209.5.3
03C687E9FFBCBF76A0F60A50CD02FD06.taxon	description	Updated description Appearance in life (Fig. 5). Adult female 2 ‒ 4 mm long, light orange, covered with white powdery wax. Wax projections from body margin short and indistinct on cephalothorax, and slightly longer on a few segments on posterior part of body. Slide-mounted adult female (Fig. 6) (n = 20). Body elongate oval, 2.6 – 4.0 mm long and 1.2 – 2.0 mm wide; derm membranous; segmentation recognizable, but not well developed. Anal lobes well developed, ventral surface with slightly faint, narrow anal lobe bar and a long apical seta, 130 – 228 µm long. Antenna 328 – 451 µm long, with 8 segments and many flagellate setae; subapical segment with 1 fleshy seta and apical segment with 3 fleshy setae. Eyespots present on margin, not associated with discoidal pores. Legs well developed, with many flagellate setae; hind leg measurements (in µm): trochanter + femur 243 – 348 long; hind tibia + tarsus 255 – 379; claw 24 – 34, without a denticle. Ratio of lengths of hind tibia + tarsus to trochanter + femur about 1: 0.91 – 1.15; ratio of lengths of hind tibia to tarsus 1: 1.90 – 2.87. Paired setose tarsal digitules present, subequal in length to the minutely knobbed claw digitules. Hind coxa with translucent pores on both surfaces; hind tibia occasionally with translucent pores on posterior surface; hind trochanter, femur and tarsus without translucent pores. Labium 98 – 165 µm long, shorter than clypeolabral shield. Circulus lacking. Ostioles present, each with inner edges of lips weakly sclerotized; anterior ostioles each with a total for both lips of 12 – 27 trilocular pores and 2 – 8 setae; each posterior ostiole with a total for both lips of 13 – 30 trilocular pores and 2 – 7 setae. Anal ring 68 – 88 µm wide, bearing 6 setae, each seta 98 – 174 µm long. Cerarii numbering 4 – 7 pairs, all present on abdominal segments. Anal lobe cerarii (C 18) mostly each containing 2 cerarian setae, each seta 14 – 40 µm long and about 4 – 8 µm wide at base, 1 – 6 auxiliary setae and a concentration of trilocular pores. Penultimate cerarii (C 17) each containing 2 conical setae, no auxiliary setae and a few trilocular pores. Cerarii situated further forward generally each with 2 conical setae with a few trilocular pores. Dorsum. Setae relatively short and flagellate, each 6 – 39 µm long, usually distributed segmentally; longest setae present on head. Trilocular pores each 3 – 4 µm wide, evenly distributed. Oral rim tubular ducts absent. Oral collar tubular ducts present or absent, if present, each 2 – 3 µm wide, located on marginal to submarginal area of posterior abdominal segments. Discoidal pores each about 1 – 2 µm wide, sparsely distributed. Venter. Setae relatively long and flagellate, each 7 – 122 µm long; setae on head longest. Multilocular disc pores, each 6 – 10 µm wide, present in medial areas of abdominal segments IV ‒ IX, but occasionally absent from a few anterior segments, arranged in 1 – 2 rows on posterior area of abdominal segment VII; 0 – 1 row on anterior area of abdominal segment VII; in 1 row on each posterior area of abdominal segments IV ‒ VI but arranged randomly on abdominal segments VIII and IX. Trilocular pores, each same size as those on dorsum, evenly distributed. Oral collar tubular ducts all 1 size, each about 2 – 3 µm wide, present on medial areas of abdominal segments III – VII and usually forming transverse bands across segments, also present on submarginal to marginal areas of posterior abdominal segments. Discoidal pores, same size as those on dorsum, sparsely present.	en	Tanaka, Hirotaka, Kamitani, Satoshi (2022): Review ofthegenus CrisicoccusFerris (Hemiptera: Coccomorpha: Pseudococcidae) in Japan with description of a new species, and the identity of a South Korean mealybug misidentified as Crisicoccus matsumotoi (Shiraiwa 1935). Zootaxa 5209 (5): 555-572, DOI: 10.11646/zootaxa.5209.5.3
03C687E9FFBCBF76A0F60A50CD02FD06.taxon	biology_ecology	Host plants in Japan. Pinaceae: Pinus densiflora, P. parviflora and P. thunbergii (Kawai 1972; 1980; 2003).	en	Tanaka, Hirotaka, Kamitani, Satoshi (2022): Review ofthegenus CrisicoccusFerris (Hemiptera: Coccomorpha: Pseudococcidae) in Japan with description of a new species, and the identity of a South Korean mealybug misidentified as Crisicoccus matsumotoi (Shiraiwa 1935). Zootaxa 5209 (5): 555-572, DOI: 10.11646/zootaxa.5209.5.3
03C687E9FFBCBF76A0F60A50CD02FD06.taxon	discussion	Remarks. Crisicoccus pini is similar to C. azaleae in having a small number of cerarii (<13 pairs) and only one size of oral collar tubular ducts. The differential diagnosis of these species is provided in “ Remarks ” under C. azaleae above.	en	Tanaka, Hirotaka, Kamitani, Satoshi (2022): Review ofthegenus CrisicoccusFerris (Hemiptera: Coccomorpha: Pseudococcidae) in Japan with description of a new species, and the identity of a South Korean mealybug misidentified as Crisicoccus matsumotoi (Shiraiwa 1935). Zootaxa 5209 (5): 555-572, DOI: 10.11646/zootaxa.5209.5.3
03C687E9FFB9BF76A0F6085FC93AF907.taxon	type_taxon	Type species: Dactylopius gutierreziae Cockerell 1896.	en	Tanaka, Hirotaka, Kamitani, Satoshi (2022): Review ofthegenus CrisicoccusFerris (Hemiptera: Coccomorpha: Pseudococcidae) in Japan with description of a new species, and the identity of a South Korean mealybug misidentified as Crisicoccus matsumotoi (Shiraiwa 1935). Zootaxa 5209 (5): 555-572, DOI: 10.11646/zootaxa.5209.5.3
03C687E9FFB9BF76A0F6085FC93AF907.taxon	diagnosis	Genus diagnosis (modified from McKenzie 1967). Body of adult female elongate oval to broadly oval; anal lobe developed, each lobe without anal lobe bar and bar seta, usually with 2 conical setae, 1 or more slender auxiliary setae, and a concentration of trilocular pores. Other cerarii usually numbering 6 – 17 pairs, each containing 2 conical setae and a slight concentration of trilocular pores. Antennae each usually with 8 segments, slender. Legs well developed, translucent pores often present on hind coxae and hind tibiae. Tarsal digitules knobbed. Circulus present or absent. Anterior and posterior ostioles usually present. Anal ring normal, usually bearing 6 setae. Body setae often flagellate, sometimes conical or spiniform. Trilocular pores normal. Multilocular disc pores present, usually confined to venter. Oral rim tubular ducts present at least on dorsum. Oral collar tubular ducts present on venter at least.	en	Tanaka, Hirotaka, Kamitani, Satoshi (2022): Review ofthegenus CrisicoccusFerris (Hemiptera: Coccomorpha: Pseudococcidae) in Japan with description of a new species, and the identity of a South Korean mealybug misidentified as Crisicoccus matsumotoi (Shiraiwa 1935). Zootaxa 5209 (5): 555-572, DOI: 10.11646/zootaxa.5209.5.3
03C687E9FFB9BF76A0F6085FC93AF907.taxon	discussion	Remarks. The genus Spilococcus is similar to Paracoccus Ezzat & McConnell 1956 in having oral rim tubular ducts and a small number of cerarii (<17) and in some cases it may be difficult to distinguish them; at least one species currently placed in Spilococcus should be transferred to Paracoccus (see “ Remarks ” under Spilococcus pacificus). Further morphological and molecular phylogenetic studies including these two genera are greatly needed. The separation of Crisicoccus from Spilococcus is discussed in “ Remarks ” under C. seruratus above.	en	Tanaka, Hirotaka, Kamitani, Satoshi (2022): Review ofthegenus CrisicoccusFerris (Hemiptera: Coccomorpha: Pseudococcidae) in Japan with description of a new species, and the identity of a South Korean mealybug misidentified as Crisicoccus matsumotoi (Shiraiwa 1935). Zootaxa 5209 (5): 555-572, DOI: 10.11646/zootaxa.5209.5.3
03C687E9FFB9BF77A0F60B12C9A0FD6A.taxon	discussion	(adapted and modified from Kaydan & Szita 2017)	en	Tanaka, Hirotaka, Kamitani, Satoshi (2022): Review ofthegenus CrisicoccusFerris (Hemiptera: Coccomorpha: Pseudococcidae) in Japan with description of a new species, and the identity of a South Korean mealybug misidentified as Crisicoccus matsumotoi (Shiraiwa 1935). Zootaxa 5209 (5): 555-572, DOI: 10.11646/zootaxa.5209.5.3
03C687E9FFB8BF75A0F60EE6CEBBFA9E.taxon	materials_examined	Material examined. South Korea: Busan, on Acer palmatum, 20. v. 2001, no collector indicated, 1 adult female mounted singly (APQA); Gunwi (GB), on Pyrus ussuriensis, 13. ix. 2012, coll. Suh, S. J., 2 adult females mounted on a slide (APQA).	en	Tanaka, Hirotaka, Kamitani, Satoshi (2022): Review ofthegenus CrisicoccusFerris (Hemiptera: Coccomorpha: Pseudococcidae) in Japan with description of a new species, and the identity of a South Korean mealybug misidentified as Crisicoccus matsumotoi (Shiraiwa 1935). Zootaxa 5209 (5): 555-572, DOI: 10.11646/zootaxa.5209.5.3
03C687E9FFB8BF75A0F60EE6CEBBFA9E.taxon	description	Updated description Appearance in life. Body orange in life (Danzig & Gavrilov-Zimin 2015). Slide-mounted adult female (Fig. 7) (n = 3): Body elongate oval, 2.7 – 3.3 mm long and 1.4 – 1.6 mm wide; derm membranous; segmentation recognizable, but not well developed. Anal lobes well developed; dorsal surface of each lobe with a slightly sclerotized area; ventral surface with well-developed anal lobe bar and long apical seta, 208 – 271 µm long. Antenna 360 – 450 µm long, with 8 segments and many flagellate setae; subapical segment with 1 fleshy seta and apical segment with 3 fleshy setae. Eyespots present on margin, not associated with discoidal pores. Legs well developed, with many flagellate setae; hind leg measurements (in µm): trochanter + femur 260 – 338 long; hind tibia + tarsus 280 – 395; claw 34 – 40, without a denticle. Ratio of lengths of hind tibia + tarsus to trochanter + femur about 1: 1.01 – 1.17; ratio of lengths of hind tibia to tarsus 1: 1.55 – 2.17. Paired setose tarsal digitules present, subequal in length to the minutely knobbed claw digitules. Hind coxa with translucent pores on both surfaces; hind tibia with or without translucent pores on posterior surface; hind trochanter, femur and tarsus without translucent pores. Labium 165 – 186 µm long, usually same length as clypeolabral shield. Circulus usually quadrate, located between abdominal segment III and IV, 68 – 170 µm long and 87 – 125 µm wide. Ostioles present, each with inner edges of lips weakly sclerotized; anterior ostioles each with a total for both lips of 16 – 29 trilocular pores and 3 – 5 setae; each posterior ostiole with a total for both lips of 19 – 49 trilocular pores and 2 – 5 setae. Anal ring 86 – 98 µm wide, bearing 6 setae, each seta 110 – 140 µm long. Cerarii numbering 6 – 8 pairs, all present on abdominal segments. Anal lobe cerarii (C 18) mostly each containing 2 conical cerarian setae, each seta 12 – 21 µm long and about 4 – 7 µm wide at base, 4 – 7 auxiliary setae and a concentration of trilocular pores. Penultimate cerarii (C 17) each containing 2 conical setae, no auxiliary setae and some trilocular pores. Cerarii situated further forward generally each with 2 conical setae and usually a few trilocular pores. Dorsum. Setae flagellate, each 8 – 45 µm long, distributed segmentally; longest setae present on head. Trilocular pores each 3 – 4 µm wide, evenly distributed. Oral collar tubular ducts absent. A few oral rim tubular ducts present on submarginal to medial areas of head, thoracic and abdominal segments. Discoidal pores each about 2 – 3 µm wide, sparsely distributed. Venter. Setae relatively long and flagellate, each 18 – 136 µm long; setae on head longest. Multilocular disc pores, each 6 – 9 µm wide, present in medial areas of abdominal segments IV ‒ IX; a few also present on head and thoracic segments, arranged in 1 to 3 rows on posterior areas of abdominal segments VI – VII but arranged randomly on abdominal segments VIII and IX. Trilocular pores, each same size as those on dorsum, evenly distributed. Oral collar tubular ducts of 2 sizes present: (i) large-type ducts, each about 3 – 4 µm in diameter, mostly wider than a trilocular pore, present on marginal to submarginal areas of all abdominal segments and anterior of thorax; and (ii) small-type ducts, each about 1 – 2 μm in diameter, present on medial areas of abdominal segments IV – VII, forming transverse bands across most posterior abdominal segments. Discoidal pores, each about 1 – 2 µm wide, sparsely distributed.	en	Tanaka, Hirotaka, Kamitani, Satoshi (2022): Review ofthegenus CrisicoccusFerris (Hemiptera: Coccomorpha: Pseudococcidae) in Japan with description of a new species, and the identity of a South Korean mealybug misidentified as Crisicoccus matsumotoi (Shiraiwa 1935). Zootaxa 5209 (5): 555-572, DOI: 10.11646/zootaxa.5209.5.3
03C687E9FFB8BF75A0F60EE6CEBBFA9E.taxon	biology_ecology	Host plants in South Korea. Moraceae: Broussonetia kazinoki (Paik 1978; Suh 2020), Rosaceae: Malus pumila (Kwon et al. 2003; Suh 2020), Pyrus pyrifolia var. cultiva (Kwon & Han 2003; Suh 2020), P. ussuriensis (Rosaceae) (Kwon et al. 2003; Suh 2020); and Sapindaceae: Acer palmatum (Paik 1978; Suh 2020).	en	Tanaka, Hirotaka, Kamitani, Satoshi (2022): Review ofthegenus CrisicoccusFerris (Hemiptera: Coccomorpha: Pseudococcidae) in Japan with description of a new species, and the identity of a South Korean mealybug misidentified as Crisicoccus matsumotoi (Shiraiwa 1935). Zootaxa 5209 (5): 555-572, DOI: 10.11646/zootaxa.5209.5.3
03C687E9FFB8BF75A0F60EE6CEBBFA9E.taxon	discussion	Remarks. In South Korea, this species was first reported as “ C. matsumotoi ” by Paik (1978) and later by Paik (2000), Kwon et al. (2003), and Suh (2020). However, the morphology of the Korean specimens examined in this study, which were identified by South Korean mealybug experts Suh, S. J. and Park, M. J. as “ C. matsumotoi ” is quite different from that of the species currently regarded as C. matsumotoi (= Crisicoccus seruratus) in Japan, the native country of the type specimen. The Korean material could be identified as a member of the genus Spilococcus, namely S. pacificus. The true C. matsumotoi (= C. seruratus) is a well-known significant pest of fruit crops in Japan (Kawai 1980; 2003) and, based on our investigation, is probably absent from South Korea. Thus, this rectification of the misidentifications will be beneficial to agriculture and international plant quarantine. Where mentioned above, Suh (2020) just cited an earlier paper by South Korean researchers; she was not responsible for the misidentifications. In this study, we could not check all the specimens of C. matsumotoi that were used by the South Korean researchers; possibly a few of them could be the true C. matsumotoi (= C. serruratus) from South Korea. However, the morphological characteristics of the Korean specimens we examined in this study are consistent with those of “ C. matsumotoi ” in the key provided by Kwon et al. (2003), such as having some oral rim tubular ducts on the dorsum. We now consider that most or all specimens reported as “ C. matsumotoi ” from South Korea are consistent with S. pacificus. We also believe that the specimens used by other researchers should be re-examined in the future, especially as Suh (2020) cited Kwon & Han (2003) as having recorded S. pacificus on Acer palmatum in Korea, whereas Kwon et al. (2003) mentioned “ C. matsumotoi ” on the same host plant. All the Korean specimens examined in this study have oral-rim tubular ducts on the dorsum and anal lobe bars on the ventral surfaces of the anal lobes. According to Kaydan & Szita (2017), the species should be transferred either to the genus Maconellicoccus Ezzat or to Paracoccus Ezzat & McConnell (probably, to the genus Paracoccus with minor generic diagnostic modifications); however, in the present study, none of the type specimens were examined so the species is left in Spilococcus for now.	en	Tanaka, Hirotaka, Kamitani, Satoshi (2022): Review ofthegenus CrisicoccusFerris (Hemiptera: Coccomorpha: Pseudococcidae) in Japan with description of a new species, and the identity of a South Korean mealybug misidentified as Crisicoccus matsumotoi (Shiraiwa 1935). Zootaxa 5209 (5): 555-572, DOI: 10.11646/zootaxa.5209.5.3
03C687E9FFBABF75A0F609ACC9A0F97D.taxon	discussion	(adapted from Kwon et al. 2003)	en	Tanaka, Hirotaka, Kamitani, Satoshi (2022): Review ofthegenus CrisicoccusFerris (Hemiptera: Coccomorpha: Pseudococcidae) in Japan with description of a new species, and the identity of a South Korean mealybug misidentified as Crisicoccus matsumotoi (Shiraiwa 1935). Zootaxa 5209 (5): 555-572, DOI: 10.11646/zootaxa.5209.5.3
