taxonID	type	description	language	source
03C7879316079338FE7ACF7D0C51FD73.taxon	diagnosis	Diagnosis. A genus of Buliminidae which is characterized by a large, brownish, elongated oval shell without apertural teeth, the palatal insertion of the peristome which does not distinctly approach to the columellar edge, a usually well-developed imperforate penial papilla (which is rudimentary only in Ena dazimonensis sp. n.; see below), a penial appendix, the lack of a penial caecum, a caecum at the middle section of the epiphallus, a more or less short agellum, and a diverticulum to the bursa copulatrix.	en	Hausdorf, B (2001): The genus Ena in Turkey, with remarks on its phylogenetic relationships (Gastropoda: Buliminidae). Journal of Natural History 35: 1627-1638
03C7879316079335FE7CCC820ECAFF3C.taxon	description	Bulimus Nogellii R oth, 1850: 341. Terra typica: ‘ K olchis’. Bulimus Nogelli [sic]: L. PfeiOEer, 1856: 56, pl. 16 gures 7, 8.	en	Hausdorf, B (2001): The genus Ena in Turkey, with remarks on its phylogenetic relationships (Gastropoda: Buliminidae). Journal of Natural History 35: 1627-1638
03C7879316079335FE7CCC820ECAFF3C.taxon	description	Buliminus (Napaeus) ponticus: R etowski, 1889: 249.	en	Hausdorf, B (2001): The genus Ena in Turkey, with remarks on its phylogenetic relationships (Gastropoda: Buliminidae). Journal of Natural History 35: 1627-1638
03C7879316079335FE7CCC820ECAFF3C.taxon	description	Shell (gures 1, 2). Elongated oval; with 7 – 8.25 slightly in ated whorls; moderately thick-walled; teleoconch with coarse growth ridges crossed by irregular incised spiral striae producing a granulated sculpture; brownish corneous; subtranslucent; the body whorl does not ascend towards the aperture; aperture oval, edentate; peristome slightly expanded, re exed and moderately thickened on the inside; palatal insertion of the peristome connected with the columellar edge by a thin callus; rimate or umbilicus open. Measurements. Various localities (n 530): D: 5.2 – 6.7 mm, mean 55.7 Ô 0.3 mm; H: 13.0 – 21.8 mm, mean 515.3 Ô 1.8 mm; D / H: 0.307 – 0.408, mean 50.374 Ô 0.025. Genitalia (gures 5, 8). The penial appendix inserts at the distal section of the penis. The appendix sections A, A and A are distinctly delimited, whereas there 1 2 3 is no marked boundary between the sections A and A. The appendix retractor 4 5 inserts at the proximal half of section A. The penial retractor inserts at the proximal 1 section of the penis. Both retractors insert close to each other at the diaphragm. An imperforate penial papilla inserts at the proximal end of the penis. There is a short groove in the proximal part of the penial papilla facing the opening of the epiphallus. There is a longitudinal fold at each side of the penial papilla. The two folds converge and become higher towards their distal ends, which are detached from the penis wall. Opposite to these structures, there are two indistinct longitudinal ridges, which run towards the opening of the epiphallus. Besides, the penial wall is irregularly sculptured. There is a spur-shaped caecum at the middle section of the very long epiphallus. The agellum is moderately short. The vagina is shorter than the penis. The free oviduct is longer than the vagina. The diverticulum of the bursa copulatrix is more than twice as long as the bursa and its duct proximally of the branching point. km 20, UTM-grid () (. n. sp, dazimonensis Ena and) (. n. sp, Bank and.) Hausdorf squares menkhorsti Ena,) () Roth (nogellii Ena of Distribution. 13. IG F Remarks. According to Forcart (1940: 118), Ena nogellii and Chondrula pontica diOEer in the ratio D / H. Ena nogellii is said to be characterized by a ratio lower than 0.345, whereas the ratio is said to be greater than 0.345 in Chondrula pontica. In 30 specimens measured for this study, the ratio D / H varies almost continuously between 0.307 and 0.408. There are no constant diOEerences between the specimens identi ed as Ena nogellii and those identi ed as Chondrula pontica by Forcart (1940). The two gures of R etowski (1886: pl. 1 gure 9) which Forcart (1940) identi ed as the two distinct species show only a smaller and a larger specimen of Ena nogellii. In Forcart’s (1940: gure 2) gure of the genitalia of Ena nogellii the diverticulum of the bursa copulatrix is shorter than the bursa and its duct proximally of the branching point. A re-examination of Forcart’s specimen has shown that the diverticulum is torn oOE. The rest of the diverticulum is still attached to the spermoviduct. M oreover, the position of the atrium is not correctly indicated in the gure; therefore the vagina appears very long. Most of the 12 examined syntypes of B. ponticus Retowski, 1886 (IZPAN; SMF 63752, 63753, 63754, 103922) belong to Ena nogellii. However, one of the specimens in the Retowski collection in the IZPAN belongs to a Mastus species. Therefore, a lectotype of B. ponticus Retowski, 1886 (IZPAN, coll. Retowski, measurements: D 55.6 mm, H 514.6 mm) has been designated to clarify the application of that name.	en	Hausdorf, B (2001): The genus Ena in Turkey, with remarks on its phylogenetic relationships (Gastropoda: Buliminidae). Journal of Natural History 35: 1627-1638
03C7879316079335FE7CCC820ECAFF3C.taxon	distribution	Distribution and material (gure 13). Ena nogellii is widespread in the Pontic region of Turkey between the V. Bolu in the west and the V. Giresun in the east.	en	Hausdorf, B (2001): The genus Ena in Turkey, with remarks on its phylogenetic relationships (Gastropoda: Buliminidae). Journal of Natural History 35: 1627-1638
03C7879316079335FE7CCC820ECAFF3C.taxon	description	‘ K olchis’ (holotype of B. nogellii Roth: ZSM 19960564). Debris of the Black Sea near K oktebel, Crimea (lectotype (design. nov.) of B. ponticus R etowski: IZPAN; paralectotypes of B. ponticus: IZPAN; SMF 63752, 63753, 63754, 103922).	en	Hausdorf, B (2001): The genus Ena in Turkey, with remarks on its phylogenetic relationships (Gastropoda: Buliminidae). Journal of Natural History 35: 1627-1638
03C78793160A9336FE90CEC00FE8FDF9.taxon	description	Shell (gure 3). Elongated oval; with 6.5 – 8.5 slightly in ated whorls; moderately thick-walled; teleoconch with more or less coarse growth ridges crossed by irregular incised spiral striae producing a granulated sculpture; brownish corneous; subtranslucent; the body whorl does not ascend towards the aperture; aperture oval, edentate; peristome slightly expanded, re exed and moderately thickened on the inside; palatal insertion of the peristome connected with the columellar edge by a thin callus; rimate or umbilicus open. Measurements. 5 km N of Tamdere (n 510): D: 6.9 – 8.0 mm, mean 57.5 Ô 0.3 mm; H: 15.5 – 19.5 mm, mean 517.2 Ô 1.1 mm; D / H: 0.395 – 0.516, mean 50.439 Ô 0.035. Genitalia (gures 6, 9). The penial appendix inserts at the distal section of the penis. The appendix sections A, A and A are distinctly delimited, whereas there 1 2 3 is no marked boundary between the sections A and A. The appendix retractor 4 5 inserts at the proximal section of section A. The penial retractor inserts at the 1 proximal section of the penis. Both retractors insert close to each other at the diaphragm. A very long and slender imperforate penial papilla inserts at the proximal end of the penis. There is a short groove in the proximal part of the penial papilla facing the opening of the epiphallus. There is a V-shaped structure with two longitudinal ridges at the penial wall behind the penial papilla. The lateral and distal wrinkled margins of this structure are detached from the penial wall. Besides, the penial wall is sculptured with irregular longitudinal ridges. There is a spur-shaped caecum at the middle section of the very long epiphallus. The agellum is moderately short. The vagina is shorter than the penis. The free oviduct is longer than the vagina. The diverticulum of the bursa copulatrix is more than twice as long as the bursa and its duct proximally of the branching point. Remarks. Ena menkhorsti diOEers from Ena nogellii (Roth) in the broader shell with a usually slightly weaker sculpture, and in the inner structures of the penis, especially the V-shaped structure behind the very long penial papilla. No constant diOEerences have been found between the proportions of the various sections of the genitalia, which are very variable in both species. The two converging folds in the penis of Ena nogellii, which are distally detached from the penis wall, are in the same position as the V-shaped structure in the penis of Ena menkhorsti and are therefore probably homologous to that structure. A detachment of such structures is not known from other Ena species or related genera. Therefore, the distal detachment of these structures is a synapomorphy of Ena menkhorsti and Ena nogellii, indicating that they are sister species. B. Hausdorf and R. A. Bank (Hoofddorp, The Netherlands) are joint authors of this new species. Etymology. The species is named in honour of Dipl. - Ing. Henk P. M. G. Menkhorst, Krimpen aan de Ijssel, who discovered it. Type locality. V. Giresun: rocks 5 km N of Tamdere, DE 48. Distribution and material (gure 13). Ena menkhorsti is widespread in the eastern Pontic region of Turkey between the Vilayets Tokat and Ordu in the west and the V. Rize in the east. Several specimens of Ena menkhorsti and of Ena nogellii have been found sympatrically at rocks 5 km N of Tamdere in the V. Giresun. V. Ordu: 12 km N of G ürgentepe, CF 92 (M EN). V. G iresun: 5 km N of Tamdere, D E 48 (det. anat.; holotype N N M, leg. H. M enkhorst, 23 July 1990, measurements: D 57.7 mm, H 517.2 mm; MEN; ZMH 2810); Giresun 49 km towards Şebin Karahisar, 800 m alt., DE 49 (det. anat.; NBE); Giresun 75 km towards Şebin K arahisar, 1500 m alt., D E 57 (N BE); 8 km S of D ereli, D F 50 (M EN); 12 km S of G iresun, D F 51 (M EN); 15 km S of G iresun, D F 51 (N BE); 7 km S of the mouth of the DogÏankent Çayõ, DF 83 (NBE); mouth of the DogÏankent Çayõ 30 km towards Torul, DF 91 (NBE). V. Trabzon: 3.5 km N of Tonya, EF 22 (IZPAN); 10 – 13 km SSE of Maçka, EF 51 (IZPAN); 35 km S of branch from OF towards Bayburt, S of Ataköy, F F 00 (NBE). V. Rize: 21 km S of Çamlihemşin, 870 m alt., F F 62 (N BE). V. Tokat: G ökdere, CE 08 (M EN).	en	Hausdorf, B (2001): The genus Ena in Turkey, with remarks on its phylogenetic relationships (Gastropoda: Buliminidae). Journal of Natural History 35: 1627-1638
03C7879316099337FE64CC700C75FE63.taxon	description	Bulimus (Petraeus) nogelli [sic]: N ägele, 1894: 106 [non R oth, 1850].	en	Hausdorf, B (2001): The genus Ena in Turkey, with remarks on its phylogenetic relationships (Gastropoda: Buliminidae). Journal of Natural History 35: 1627-1638
03C7879316099337FE64CC700C75FE63.taxon	description	Measurements. Tokat (n 530): D: 5.6 – 6.7 mm, mean 56.1 Ô 0.3 mm; H: 18.2 – 22.6 mm, mean 520.1 Ô 1.0 mm; D / H: 0.265 – 0.346, mean 50.307 Ô 0.020. Genitalia (gures 7, 10). The penial appendix inserts at the distal section of the penis. There is neither a marked boundary between the appendix sections A and 1 A nor between the sections A, A and A. The appendix retractor inserts at the 2 3 4 5 proximal section of the extraordinarily long section A. The penial retractor inserts 1 at the proximal section of the penis. Both retractors insert close to each other at the diaphragm. There is a horseshoe-shaped ridge with ends which bend inwards in the proximal section of the penis. There is a small, conical hump, which is probably a rudiment of the penial papilla, in the middle of the horseshoe-shaped structure. There is a heel-shaped caecum at the middle section of the very long epiphallus. The agellum is very short. The vagina is more than half as long as the penis. The free oviduct is four times as long as the vagina. The diverticulum of the bursa copulatrix is more than twice as long as the bursa and its duct proximally of the branching point. Remarks. Ena dazimonensis diOEers from Ena nogellii (Roth) and Ena menkhorsti Hausdorf and Bank, sp. n. in the almost smooth shell which has on average more whorls that are hardly in ated, and by the distinctly thickened peristome. The genitalia of Ena dazimonensi s are especially characterized by the heel-shaped epiphallial caecum, the very short agellum and the inner structures of the penis, especially the horseshoe-shaped ridge and the rudimentary penial papilla. The inner structure of the penis of Ena dazimonensis resembles that of Pseudonapaeu s chodschendicu s Schileyko, 1978. Etymology. The species is named after the ancient village Dazimon (the modern Tokat), the type locality of the species. Type locality. V. Tokat: Tokat, rocks above the cemetery, BE 96. Distribution and material (gure 13). Ena dazimonensis is known only from the surroundings of Tokat. V. Tokat: Tokat, BE 96 (det. anat.; holotype ZMH 2807, leg. B. Hausdorf, 5 October 1987, measurements: D 56.1 mm, H 520.4 mm; H AU; N M B 4480 d; SM F 14486, 63751, 104386, 104387, 104388, 202962, 237583, specimens to Nägele, 1894 and Forcart, 1940; ZM H 2808); 3.5 km S of Tokat, BE 96 (HAU); 5 km S of Tokat, BE 96 (M EN).	en	Hausdorf, B (2001): The genus Ena in Turkey, with remarks on its phylogenetic relationships (Gastropoda: Buliminidae). Journal of Natural History 35: 1627-1638
