identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
03C7D003BB135D2DD0F804A641ABFDED.text	03C7D003BB135D2DD0F804A641ABFDED.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hemerodromia aliaexstriata Plant 2020	<div><p>Hemerodromia aliaexstriata sp. nov.</p><p>(Figs 1–2, 12)</p><p>Diagnosis. A predominantly orange yellow species with a dark median stripe on the scutum becoming wider posteriorly, a yellow scutellum and with dark marks between the front coxae and on the katepisternum. The male cercus is broad in lateral view with bluntly pointed dorsoapical and ventroapical processes and the epandrium is narrow and elongate, extending beyond the tip of the cercus.</p><p>Description. Male: body length 2.6 mm; wing length 2.0 mm. Head. Rather elongate, ~1.5X long as wide; upper occiput slightly bulging, making smooth angle where meeting vertex. Black with paler dusting. Antenna and mouthparts whitish. One pair of minute reclinate ocellar setulae; 3–4 pairs of minute vertical setulae only about as long as ocellars. Antenna with postpedicel ~1.8–2.0X long as wide, stylus somewhat shorter. Thorax. Not quadrate anteriorly; anterior margin of scutum gradually curving in lateral view. Ground colour orange yellow; scutum with median brownish black stripe, narrow anteriorly, becoming as wide as base of scutellum posteriorly; mediotergite broadly darkened medially; strong black mark between insertion points of front coxae and another smaller oblique mark on katepisternum; small dark marks also present immediately in front of base of wing, obliquely about prothoracic spiracle and on sutures of laterotergite. All setae whitish yellow, minute; notopleural setulae present but very small, scutellars apparently absent. Legs. Yellowish, distal tarsal segments somewhat darkened. C1~1.1X long as distance between base of C1 and C2. F1 hardly longer than C1, ~4.0X long as wide, strongly and evenly inflated, slightly constricted on proximal 0.2; femoral formula ~ 6–8/22–25/21–24/5–6, denticles black, rows converging apically; spines yellow except 1–2 near base black. T1 ~ 0.7X long as F1; evenly curved, ventral face shallowly concave, with 2 rows of ~20 sharply pointed spinose setae ventrally; ventroapical spur weakly developed; distinct ventroapical erect black spinose seta present. Mid and hind legs slender with only small setulae. Wing. Membrane darkened by greyish microtrichia on distal 0.9, paler about base of wing. Veins greyish black, R 4+5 darker still. Veins C and R 1 yellowish at extreme base. Marginal setulae dark. R 2+3 almost linear, joining C ~0.8–0.9X distance between ends of R 1 and R 4. Length of C between ends of R 2+3 and R 4 about as long as R 4. R 4 slightly S-shaped. R 5 ~ 2.0X long as R 4, almost linear. R 4+5 fork distal to M 1+2 fork by ~ 1.3X length of R 4. M 1 almost linear, slightly convergent with R 5, becoming parallel at extreme tip. Cell bm+dm short, ending at tip of R 1. Halter yellowish white. Abdomen. Black dorsally, paler ventrally, tergites 1 and 2 yellowish. Terminalia. Black with dark setae. Cercus (Figs 1–2) short, not extending beyond tip of epandrium; rather broad in lateral view with short bluntly pointed dorsoapical process and longer ventroapical process; outer surface with fine black hairs. Epandrium long and narrow with fine black hairs (Fig. 1). Hypandrium small, strongly arched, bare. Phallus with pair of apical hook-like processes. Female: similar to male but scutum with median stripe somewhat narrower anteriorly and abdomen yellowish ventrally. Tip of abdomen truncate, terminalia rather short, hardly extensible and not at all ovipositor-like.</p><p>Type material. HOLOTYPE ♂: THAILAND, Mae Hong Son Province, river near <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=98.12089&amp;materialsCitation.latitude=19.47797" title="Search Plazi for locations around (long 98.12089/lat 19.47797)">Susa</a> waterfall, 306 m, 19.47797°N, 98.12089°E, 3.iv.2019, netted A. R. Plant (QSBG) . PARATYPES: 2♀, same data as holotype (QSBG); 1♂, 3♀, Mae Hong Son Province, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=98.12421&amp;materialsCitation.latitude=19.47457" title="Search Plazi for locations around (long 98.12421/lat 19.47457)">Susa</a> waterfall, tufa cliff at confluence of tufa str. and river, 305 m, 19.47457°N, 98.12421°E, 3.iv.2019, netted A. R . Plant (QSBG) .</p><p>Etymology. The specific epithet is a concatenation of the Latin alia ex meaning ‘different from’ and striata in reference to this species not being identical with H. striata Yang &amp; Yang. It is used as a noun in apposition.</p><p>Remarks. The markings on the scutum of this species somewhat resemble those of H. acutata Grootaert, Yang &amp; Saigusa (in which the width and intensity of the median dark stripe varies greatly) or even H. systoechon Plant (in which the dark scutal marking is usually very broad with diffuse edges) from which it may readily be separated by the male terminalia. A generally similarly broad cercus with bluntly pointed dorsoapical and ventroapical processes suggests close affinities with two extralimital species, H. oratoria (Fallén) (Holarctic distribution), and H. striata Yang &amp; Yang from the eastern Palaearctic (China, Beijing). The latter species also has an elongate epandrium. Hemerodromia aliaexstriata sp. nov. is known from a calcareous river in the extreme northwest of Thailand (Fig. 12).</p></div>	https://treatment.plazi.org/id/03C7D003BB135D2DD0F804A641ABFDED	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Plant, Adrian R.	Plant, Adrian R. (2020): New species of Hemerodromia Meigen (Diptera: Empididae: Hemerodromiinae) associated with limestone karstic waters in Thailand. Zootaxa 4758 (3): 549-560, DOI: 10.11646/zootaxa.4758.3.7
03C7D003BB125D2AD0F806F241ADFB70.text	03C7D003BB125D2AD0F806F241ADFB70.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hemerodromia deprimatura Plant 2020	<div><p>Hemerodromia deprimatura sp. nov.</p><p>(Figs 3–4, 13)</p><p>Diagnosis. A dirty yellow species with a triangular brownish marking behind eye on lower occiput. Male terminalia with inner margin of cercus evenly concave in dorsal view, tip of surstylus L-shaped and a narrow process emerging beyond tip of surstylus (possibly lower surstylus) with very narrow shaft and small black subcircular process apically.</p><p>Description. Male: body length 2.1 mm; wing length 1.7 mm. Head. Slightly elongate. 1.4X long as deep, upper occiput making smooth curve where meeting vertex, hardly bulging. Dirty yellow with pale dusting, lower occiput with brown mark behind eye becoming broader ventrally (appearing triangular in lateral view). Antenna and mouthparts whitish. One pair of minute reclinate ocellar setulae; 3–4 pairs of vertical setulae positioned in linear series parallel with upper eye margin, clearly distinguished from other fine hairs; postocular and lower occipital setulae minute.Antenna with postpedicel ~2.5X long as wide, stylus of similar length. Thorax. Dirty whitish yellow with greyish dust; darker on posterior margins of scutellum and mediotergite; blackish about anterior spiracle, ‘scutoscutellar eye’ and ventrally between insertion points of front coxa. All setae yellowish, minute, but notopleural distinct and pair of weak scutellars. Legs. Dirty whitish yellow, apical tarsomeres of all legs very slightly darker. C1 ~1.2–1.3X longer than distance between base of C1 and C2; C1, C2 and C3 with all setulae minute. F1 ~1.0–1.1X long as C1; F1 4–5X long as wide, very slightly constricted on inner face at 0.2 from base. F1 with two parallel rows of 17–21 minute black denticles ventrally, either side of which is linear series of less numerous and inconspicuous fine yellow bristles, noticeably stronger around 0.2 from base. T1 ~0.8X long as F1, slightly curved, ventral face shallowly concave bearing two rows of minute, sharply pointed, blackish spinose setulae; ventroapical spur weakly developed; distinct ventroapical erect black spinose seta present. Wing. Membrane clear but darkened on distal 0.9 by blackish microtrichia. Veins greyish on distal 0.9, paler on proximal 0.1. R 2+3 linear, joining C ~ 0.7X distance between ends of R 1 and R 4. Length of C between ends of R 2+3 and R 4 ~1.7–1.8X long as R 4. R 4 linear, not at all Sshaped, angle with R 5 at extreme base ~ 70°. R 5 ~2.2X long as R 4. R 4+5 fork distal to M 1+2 fork by ~1.5X length of R 4. R 5 and M 1 almost linear, distinctly convergent distally. Cell bm+dm rather long, ending just beyond end of R 1. Halter dirty white. Abdomen. Tergites blackish but tergite 1 pale with narrow transverse black fascia at middle, tergite 7 and all sternites more uniformly pale. All setulae small and dark, longer on posterior margin of 7 th segment. Terminalia dirty yellowish. Cercus (Figs 3–4) elongate, extending slightly beyond tip of epandrium, slightly inflated apically in lateral view, some fine hairs dorsally and on inner face slightly stronger dorsoapically; inner margin distinctly evenly concave in dorsal view. Epandrium (Fig. 4) bluntly pointed apically, covered with fine tomentum and a few strong bristles apically. Surstylus present, extending distinctly beyond tip of epandrium, rather broad, Lshaped apically with broad dorsal process; narrow process emerging beyond tip of surstylus (possibly lower lobe of surstylus) with very narrow shaft and small black subcircular process apically (apparently flat on inner face, more convex on outer). Hypandrium (Fig. 4) large, pointed apically, covered with fine tomentum and only a few distinct bristles. Female: similar to male but abdomen with terminalia elongate, somewhat ovipositor-like.</p><p>Type material. HOLOTYPE ♂: THAILAND, Phang Nga Province, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=98.5414&amp;materialsCitation.latitude=8.51063" title="Search Plazi for locations around (long 98.5414/lat 8.51063)">Sa Nang Namora</a> waterfall, tufa stream, 59 m, 8.51063°N, 98.54140°E, 16.i.2019, netted A. R. Plant (QSBG) . PARATYPES: 2♂, 2♀, same data as holotype (QSBG) .</p><p>Etymology. The specific epithet is derived from the Latin deprimatur, meaning ‘perfectly poised’ in reference to the apparent hunting posture adopted by adult Hemerodromia when searching for prey.</p><p>Remarks. This species is very similar to and clearly closely related to H. anisoserrata Plant and H. ocellata Plant from Thailand and H. serrata Saigusa and Yang from China (Henan). The new species may be distinguished by the triangular brown marking on the lower occiput and the form of the male terminalia in which the cercus is evenly curved on the inner face (irregular in anisoserrata / ocellata / serrata), the elongate bluntly pointed epandrium (apically broad in anisoserrata, short and broad in ocellata, apically narrower in serrata) and the complex shapes of the two elongate processes of the upper and lower surstylus (very different shapes in anisoserrata / ocellata / serrata). Hemerodromia deprimatura sp. nov. is only known from a calcareous tufa stream in southern Thailand (Fig. 13).</p></div>	https://treatment.plazi.org/id/03C7D003BB125D2AD0F806F241ADFB70	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Plant, Adrian R.	Plant, Adrian R. (2020): New species of Hemerodromia Meigen (Diptera: Empididae: Hemerodromiinae) associated with limestone karstic waters in Thailand. Zootaxa 4758 (3): 549-560, DOI: 10.11646/zootaxa.4758.3.7
03C7D003BB155D28D0F8011745A1FECD.text	03C7D003BB155D28D0F8011745A1FECD.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hemerodromia oretenebraea Plant 2020	<div><p>Hemerodromia oretenebraea sp. nov.</p><p>(Figs 5–7, 14)</p><p>Diagnosis. A blackish species with head and thorax rather quadrate. The anterior of the wing is covered with dark microtrichia, usually contrasting strongly with the posterior part. The front femur is deep black, contrasting strikingly with otherwise pale yellowish legs.</p><p>Description. Male: body length 2.8 mm; wing length 1.8 mm. Head. Rather short, only slightly longer than deep, rather quadrate (especially in dorsal view); upper occiput vertical in lateral view, upper part slightly but distinctly bulging where meeting vertex. Black, dusted greyish; antenna blackish; mouthparts pale yellow, proboscis with darker tip. One pair reclinate ocellar setulae; 4–5 pairs small indistinct vertical setulae, about as strong as ocellars; pair of minute frontals behind base of antenna. Antenna with postpedicel &lt;1.5X long as wide (viewed laterally); stylus about as long as postpedicel; scape with small fine dorsal setae. Thorax. Distinctly quadrate anteriorly; anterior margin of scutum vertical in lateral view making distinct but smooth right angle with disc (not gradually curving). Ground colour deep black, dusted greyish. All setae black, minute apart from one small notopleural. Scutellum with one subapical pair of very small convergent setae. Legs. Pale yellowish with apical tarsomeres somewhat darker, F1 and front trochanter contrastingly black. C1 about as long as distance between C1 and C2, a more or less linear series of minute setulae along length of limb dorsally. F1 hardly longer than C1, distinctly inflated, ~ 4–5X long as wide, not constricted on proximal 0.3. F1with two closely spaced rows of ~ 15–21 minute black denticles, either side of which is linear series of ~ 5–6 less numerous, rather hair-like yellowish or brownish spines, noticeable stronger and more bristle-like about 0.2 from base.T1 ~ 0.7X long as F1, sublinear, ventral face only slightly concave; with one row of ~12 sharply pointed spinose setae ventrally; ventroapical spur weakly developed; distinct ventroapical erect black spinose seta present. Mid and hind legs slender with only small setulae. Wing. Membrane distinctly darkened with blackish microtrichia, especially anteriorly of M in distal part of wing beyond radial fork; cell bm+dm, base and posterior part of wing paler. Veins greyish black, R 2+3, R 4+5, R 4 and R 5 distinctly darker and somewhat thickened. Marginal setulae greyish black. R 2+3 linear, joining C ~ 0.7–0.8X distance between ends of R 1 and R 4. Length of C between ends of R 2+3 and R 4 ~ 1.0–1.1X long as R 4. R 4 slightly S-shaped, angle with R 5 at extreme base ~ 65–70°. R 5 ~ 1.7–2.0X long as R 4, almost linear, almost parallel with M 1 but divergent apically. Cell bm+dm rather long, ending distinctly beyond end of R 1. Halter greyish white. Abdomen. Blackish, all setulae inconspicuous, very short, dark, a few longer setulae on sternites 7 and 8. Terminalia. Black with dark setae. Moderately small and compact. Cercus (Figs 5–6) elongate, extending beyond tip of epandrium, narrow in lateral view, inner margin distinctly evenly concave in dorsal view. Epandrium (Fig. 5) rather ovate with some fine setae and patch of microscopic pile distally. Surstylus apparently more or less fused with tip of epandrium, bearing two distinct spines (Fig. 7). Hypandrium (Fig. 5) short, compact, distinctly arched. Apex of phallus bifid in posterior view. Female: similar to male but abdomen with shorter setulae and terminalia elongate, ovipositor-like.</p><p>Type material. HOLOTYPE ♂: THAILAND, Loei Province, Hin Pha Ngam, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=101.74813&amp;materialsCitation.latitude=17.06639" title="Search Plazi for locations around (long 101.74813/lat 17.06639)">Pang Din</a> waterfall, tufa stream, 628 m, 17.06639°N, 101.74813°E, 5.iii.2019, netted A. R. Plant (QSBG) . PARATYPES: 4♀ same data as holo- type (QSBG); 1♀, Phang Nga Province, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=98.5414&amp;materialsCitation.latitude=8.51063" title="Search Plazi for locations around (long 98.5414/lat 8.51063)">Sa Nang Namora</a> waterfall, tufa stream, 59 m, 8.51063°N, 98.54140°E, 16.i.2019, netted A. R . Plant (QSBG) ; 7♂, Lampang Province, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=99.6628&amp;materialsCitation.latitude=19.3164" title="Search Plazi for locations around (long 99.6628/lat 19.3164)">Wang Kaew</a> waterfall, 568 m, 19.3164°N, 99.6628°E, 14.i.2020, netted A. R . Plant (QSBG) ; 1♂, Phrae Province, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=100.4495&amp;materialsCitation.latitude=18.4424" title="Search Plazi for locations around (long 100.4495/lat 18.4424)">Huai Rong</a> tufa waterfall, 450 m, 18.4424°N, 100.4495°E, 18.i.2020, netted A. R . Plant (QSBG) .</p><p>Additional material. 2♂, 1♀, Chiangrai Province, Doi Luang National Park, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=99.6949&amp;materialsCitation.latitude=19.4422" title="Search Plazi for locations around (long 99.6949/lat 19.4422)">Pu Kaeng</a> waterfall, 540 m, 19.4422°N, 99.6949°E, 15.i.2020, netted A. R . Plant (QSBG) ; 2♂, 4♀, Lampang Province, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=99.8164&amp;materialsCitation.latitude=18.7457" title="Search Plazi for locations around (long 99.8164/lat 18.7457)">Mae Kae</a> waterfall, 537 m, 18.7457°N, 99.8164°E, 16.i.2020, netted A. R . Plant (QSBG) .</p><p>Etymology. The specific epithet is derived from the Latin ore, meaning ‘edge’ and tenebrae meaning ‘darkness’ in reference to the darker membrane on the anterior part of the wing.</p><p>Remarks. The quadrate, head and thorax and relatively simple and small male terminalia with fused surstylus bearing two spines suggest that this species is related to H. conspecta Plant, also from Thailand. However, it may readily be distinguished by the bulging occiput, vein C being all dark (basally yellow in H. conspecta), wing distinctly darkened anteriorly and having F1 black (yellow in H. conspecta) contrasting strongly with the rest of the legs as well as shape of the cercus and surstylus of the male terminalia. In some examples the wing is more gener- ally darkened although still darker anteriorly. Although darkened front femora are found in several other species of Hemerodromia, the front coxa is seldom clear yellow as in the new species where the contrast between the front femur and coxa is striking, and is a rare example of leg colour being diagnostically useful in the genus. Hemerodromia oretenebraea sp. nov. has a disjunct distribution occurring at several sites in the north and one in the south of the country (Fig. 14).</p></div>	https://treatment.plazi.org/id/03C7D003BB155D28D0F8011745A1FECD	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Plant, Adrian R.	Plant, Adrian R. (2020): New species of Hemerodromia Meigen (Diptera: Empididae: Hemerodromiinae) associated with limestone karstic waters in Thailand. Zootaxa 4758 (3): 549-560, DOI: 10.11646/zootaxa.4758.3.7
03C7D003BB175D28D0F805D24120F8FD.text	03C7D003BB175D28D0F805D24120F8FD.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hemerodromia pairoti Plant 2020	<div><p>Hemerodromia pairoti sp. nov.</p><p>(Figs 8–9, 15)</p><p>Diagnosis. A yellowish species obscurely darkened on scutum medially and on scutellum and mediotergite. Best recognised by characters of the male terminalia, with cercus dilated apically and surstylus apically L-shaped.</p><p>Description. Male: body length 2.4 mm; wing length 2.0 mm. Head. Slightly elongate. 1.4X long as deep, upper occiput making smooth curve where meeting vertex, hardly bulging. Black with paler dusting. Antenna and mouthparts whitish. One pair of minute reclinate ocellar setulae; ~ 3 pairs of smaller verticals. Antenna with postpedicel ~ 2.5–3.0X long as wide, stylus of similar length. Thorax. Yellow; scutum with vaguely darkened broad median stripe, at centre of which is much narrower and somewhat darker median stripe; scutellum and mediotergite brownish; small dark marks ventrally between insertion points of front coxae, obliquely about anterior spiracle, immediately in front of base of wing and in sutures of laterotergite. All setae yellowish, minute; notopleural very small; scutellum with pair of minute hairs on disc. Legs. Whitish yellow, distal tarsal segments hardly darker. C1 ~1.2X longer than distance between base of C1 and C2. C1, C2 and C3 all with minute setulae. F1 ~1.1X long as C1, ~ 5X long as wide, slightly constricted on ventral face 0.2 from base; with two parallel rows of ~16 minute black denticles ventrally, either side of which is linear series of ~5–6 inconspicuous fine yellow bristles, noticeably stronger around 0.2 from base. T1 ~0.75X long as F1, slightly curved, ventral face shallowly concave bearing two rows of ~ 20 minute, sharply pointed, blackish spinose setulae; ventroapical spur weakly developed; distinct ventroapical erect black spinose seta present. Wing. Membrane darkened by greyish microtrichia on distal 0.9, paler at base. Veins greyish, paler at extreme base. R 2+3 linear, joining C ~ 0.7–0.8X distance between ends of R 1 and R 4. Length of C between ends of R 2+3 and R 4 ~1.2–1.4X long as R 4. R 4 linear, hardly S-shaped, angle with R 5 at extreme base ~ 75°. R 5 ~2.2X long as R 4. R 4+5 fork distal to M 1+2 fork by ~1.5X length of R 4. R 5 and M 1 almost linear, slightly but distinctly convergent distally. Cell bm+dm ending just beyond end of R 1. Halter yellowish white. Abdomen. Black, paler basally, especially on sternites 1–3. All setulae small and dark, becoming slightly longer on posterior margins of distal segments. Terminalia black. Cercus (Figs 8–9) elongate, extending slightly beyond tip of epandrium, conspicuously expanded dorsoapically (Fig. 8); with numerous distinct setae. Epandrium (Fig. 8) bluntly pointed apically, with some moderately strong bristles apically. Surstylus present, extending distinctly beyond tip of epandrium, broadly L-shaped distally. Hypandrium pointed apically, with fine tomentum and few distinct bristles. Phallus not examined. Female: similar to male but scutum with broad median stripe more clearly defined. T1 with ventroapical setae rather longer. Abdomen with sternites 2–5 more distinctly yellowish; terminalia not elongate, not ovipositor-like.</p><p>Type material. HOLOTYPE ♂: THAILAND, Mae Hong Son Province, Ban Muang Phaeum, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=98.30049&amp;materialsCitation.latitude=19.5854" title="Search Plazi for locations around (long 98.30049/lat 19.5854)">Phaeum River</a>, 649 m, 19.5854°N, 98.30049°E, 4.iv.2019, netted A. R. Plant (QSBG) . PARATYPES: 1♀, same data as holotype .</p><p>Etymology. The specific epithet honours Pairot Pramual in recognition of consistent support for my explorations of Thailand’s Empididae fauna.</p><p>Remarks. The yellowish thorax with variably dark median stripe on the scutum recalls H. systoechon or H. acutata. However the thoracic markings of these species (and very probably of this species) are very variable and identification must rely on examination of the male terminalia. The male cercus is greatly dilated apically resembling that of H. songsee Plant, which has a very distinctive black and yellow pattern on the thorax, quite unlike the new species. A generally similar L-shaped apex of the surstylus occurs in other southeast Asian species (e.g., H. attenuata, H. furcata, H. yunnanensis, H. digitata, H. oretenebraea sp. nov. and H. samoha sp. nov.). Hemerodromia pairoti sp. nov. is known only from a calcareous river in the extreme northwest of Thailand (Fig. 15).</p></div>	https://treatment.plazi.org/id/03C7D003BB175D28D0F805D24120F8FD	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Plant, Adrian R.	Plant, Adrian R. (2020): New species of Hemerodromia Meigen (Diptera: Empididae: Hemerodromiinae) associated with limestone karstic waters in Thailand. Zootaxa 4758 (3): 549-560, DOI: 10.11646/zootaxa.4758.3.7
03C7D003BB165D26D0F8005F44E4FC26.text	03C7D003BB165D26D0F8005F44E4FC26.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hemerodromia samoha Plant 2020	<div><p>Hemerodromia samoha sp. nov.</p><p>(Figs 10–11, 16)</p><p>Diagnosis. A species with brownish or yellowish thorax with wing membrane and veins mostly dark. The cercus is elongate and crenulated dorsally with numerous long bristles and the upper lobe of the surstylus is distinctly Lshaped.</p><p>Description. Male: body length 2.2 mm; wing length 1.7 mm. Head. Rather elongate. 1.6–1.8X long as deep; upper occiput rather linear in lateral view making smooth curve where meeting vertex, not bulging. Black dusted greyish, antenna and mouthparts pale whitish yellow. One pair of minute reclinate ocellar setulae; 3–4 pairs of vertical setulae. Antenna with postpedicel ~2.5–3.0X long as wide, stylus of slightly shorter. Thorax. Yellowish with scutum posteriorly, scutellum and mediotergite brownish; dark markings ventrally between insertion points of front coxae, obliquely about anterior spiracle, immediately in front of base of wing and in sutures of laterotergite. All setae yellowish, minute, but notopleural distinct, scutellars apparently absent. Legs. Whitish yellow, distal tarsomeres of all legs hardly darker. C1 ~1.3–1.4X longer than distance between base of C1 and C2; C1, C2 and C3 with all setulae minute. F1 ~1.0–1.1X long as C1; 4–5X long as wide, slightly constricted on inner face at 0.25 from base. F1 ventrally with two rows of ~5–6 yellow spines, inconspicuous except 1–2 near base stronger; between which are two rows of ~23 (anteroventral series) and ~18 (posteroventral series) minute black denticles. T1 ~0.7X long as F1, slightly curved, ventral face shallowly concave bearing two rows of ~20 minute, sharply pointed, blackish spinose setulae; ventroapical spur weakly developed; distinct ventroapical erect black spinose seta present. Wing. Membrane darkened by blackish microtrichia on distal 0.9; proximal 0.1 contrastingly yellowish white. Veins blackish on distal 0.9 except about base of R 1 and R 4; pale on proximal 0.1. R 2+3 linear, joining C ~ 0.7X distance between ends of R 1 and R 4. Length of C between ends of R 2+3 and R 4 ~1.7X long as R 4. R 4 hardly S-shaped, angle with R 5 at extreme base ~ 75–80°. R 5 long, ~3.0X long as R 4. R 4+5 fork distal to M 1+2 fork by ~1.5X length of R 4. R 5 and M 1 almost linear, distinctly convergent distally. Cell bm+dm ending just beyond end of R 1. Halter yellowish white. Abdomen. Yellow but tergites 2–7 blackish. Setae mostly small and inconspicuous. Terminalia blackish. Cercus (Figs 10–11) elongate, extending beyond tip of epandrium, apically expanded and crenulate with numerous conspicuous bristles on crenulations apically. Epandrium (Fig.10) bluntly pointed apically, some distinct bristles distally. Two elongate processes emerging beyond tip of epandrium; upper process (upper lobe of surstylus?) large, L-shaped; lower process (lower lobe of surstylus?) with small black leaf-like appendage apically. Hypandrium strongly arched, lacking distinct setae. Phallus not examined. Female: similar to male but abdomen with sternite 7 black, terminalia elongate, somewhat ovipositor-like.</p><p>Type material. HOLOTYPE ♂: THAILAND, Satun Province, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=99.90927&amp;materialsCitation.latitude=7.09092" title="Search Plazi for locations around (long 99.90927/lat 7.09092)">Wang Sai Thong</a> waterfall, 92 m, 7.09092°N, 99.909274°E, 13.v.2019, netted A. R. Plant (QSBG) . PARATYPES: 1♀, same data as holotype (QSBG); 2♂, 1♀, Nakhon Si Thammarat Province, Nan Sawan waterfall, 331 m, 7.89195°N, 99.78899°E, 11.v.2019, netted A. R . Plant (QSBG).</p><p>Etymology. The specific epithet honours Abdullah Samoh, fellow student of Empidoidea.</p><p>Remarks. The colour of the thorax of this species varies greatly from predominantly yellow with dark markings, to predominantly brownish and the species is probably best recognised by its distinctive male terminalia. The upper elongate process (here interpreted as the upper lobe of surstylus following Plant 2015) is distinctly L-shaped recalling particularly H. furcata Grootaert, Yang &amp; Yang and H. yunnanensis (both widespread in Thailand) and H. digitata Grootaert, Yang &amp; Saigusa (a species known from China, Yunnan) although many Asian species have apically inflated structures that despite being variously shaped, are probably analogous, and may be characteristic of a species-group within Hemerodromia (Plant 2015) . Leaf-like appendages on the tip of the lower lobe of the surstylus are found in several regional species including H. anisoserrata Plant, H. betalutea Plant, H. fusca Yang &amp;Yang, H. ocellata Plant and H. yunnanensis. The general pattern of colouration of the thorax and wing and crenulated dorsal process on the cercus of H. samoha sp. nov. recall H. namtokhinpoon Plant, a stenotopic tufa species only known from Loei province. The combination of cercus shape and form of the surstylus should enable identification of this species. Hemerodromia samoha sp. nov. is known from two tufa waterfall localities in the south of Thailand (Fig. 16).</p></div>	https://treatment.plazi.org/id/03C7D003BB165D26D0F8005F44E4FC26	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Plant, Adrian R.	Plant, Adrian R. (2020): New species of Hemerodromia Meigen (Diptera: Empididae: Hemerodromiinae) associated with limestone karstic waters in Thailand. Zootaxa 4758 (3): 549-560, DOI: 10.11646/zootaxa.4758.3.7
