identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
8DED010F0388039F833595CD4F29E972.text	8DED010F0388039F833595CD4F29E972.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Liogluta Thomson 1858	<div><p>Taxon classification Animalia Coleoptera Staphylinidae</p><p>Liogluta Thomson, 1858</p><p>Liogluta Thomson, 1858: 35. Type species Homalota umbonata Erichson, 1839, by monotypy. As valid genus: Lohse (1974); Lohse et al. (1990).</p><p>Anepsiota Casey, 1894: 321; Casey (1906: 335); as Atheta (Anepsiota): Casey (1910: 12), Fenyes (1920: 203). Synonymized by Bernhauer and Scheerpeltz (1926: 656); Moore and Legner (1975: 350).</p><p>Athetota Casey, 1906: 334. Synonymized with Atheta (Anepsiota) by Fenyes (1920: 203); as synonym of Atheta (Liogluta): Bernhauer and Scheerpeltz (1926: 656), Moore and Legner (1975: 350).</p><p>Hypnota Mulsant &amp; Rey, 1873: 591. Synonymized with Atheta (Liogluta) by Fenyes (1920: 203); Bernhauer and Scheerpeltz (1926: 656); Moore and Legner (1975: 350).</p><p>Diagnosis.</p><p>Body length ranging from 2.8-5.4 mm, body narrowly subparallel (Figs 1, 8, 16, 23, 30, 35, 44, 50, 58, 65, 72, 79, 86, 90), moderately flattened; elytra and abdomen wider than head and pronotum; uniformly dark brown or reddish-brown with head and posterior abdomen dark brown to almost black; integument of forebody with strong meshed microsculpture; surface of elytra often granulose (Figs 50, 51); head with incomplete postocular carinae, postocular region of head long and in most species longer than diameter of eye; glossae split into two lobes; antennae long with articles VI-X subquadrate, slightly transverse or rarely slightly elongate; pronotum with hypomera fully visible in lateral view; pubescence of pronotum directed posteriad on midline of disc and posterolaterad to laterad on sides; mesocoxae narrowly separated, metasternal process short and acute; legs long, three basal articles of metatarsi highly elongate in most species; tarsal formula 5-5-5. Male. Apical margin of male tergite VIII often with broad, variably-shaped rectangular projection, with edge entire or bearing crenulation or variably shaped structures, with two lateral teeth and sometimes with additional median tooth; integument of disc often with broadly distributed asperate punctation near apex (Figs 3, 11, 18, 25, 38, 46, 53, 60, 67, 74, 81, 88, 92); median lobe of aedeagus simple with apical part variably shaped in lateral view (Figs 2, 9, 17, 24, 36, 45, 52, 59, 66, 73, 80, 87, 91). Female. Spermatheca with capsule narrowly club-shaped or tubular, with apical invagination moderate to deep, stem long, sinuate, looped or twisted posteriorly (Figs 7, 15, 22, 29, 34, 42, 43, 57, 64, 71, 78, 85, 96); sternite VIII with apical margin rounded or medially emarginate, antecostal suture arcuate, or slightly to distinctly sinuate (Figs 6, 14, 21, 28, 33, 41, 49, 56, 63, 70, 77, 84, 89, 95).</p><p>Distribution.</p><p>The genus is holarctic in distribution (Smetana 2004). Seevers (1978) mentioned a few species from Africa and Jamaica but these records need verification.</p><p>Key to Canadian and Alaskan species of Liogluta</p></div>	https://treatment.plazi.org/id/8DED010F0388039F833595CD4F29E972	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Klimaszewski, Jan;Webster, Reginald P.;Langor, David W.;Sikes, Derek;Bourdon, Caroline;Godin, Benoit;Ernst, Crystal	Klimaszewski, Jan, Webster, Reginald P., Langor, David W., Sikes, Derek, Bourdon, Caroline, Godin, Benoit, Ernst, Crystal (2016): A review of Canadian and Alaskan species of the genus Liogluta Thomson, and descriptions of three new species (Coleoptera, Staphylinidae, Aleocharinae). ZooKeys 273: 217-256, DOI: http://dx.doi.org/10.3897/zookeys.573.7878, URL: http://dx.doi.org/10.3897/zookeys.573.7878
3D5339B5E610D73EBD5C18424D565CBA.text	3D5339B5E610D73EBD5C18424D565CBA.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Liogluta terminalis (Casey 1906) Casey 1906	<div><p>Taxon classification Animalia Coleoptera Staphylinidae</p><p>Liogluta terminalis (Casey, 1906) Figs 1-7</p><p>Anepsiota terminalis Casey, 1906: 339. As Atheta (Liogluta): Bernhauer and Scheerpeltz 1926: 658 (as syn. of Atheta renominata). Holotype (female): Canada, British Columbia, Glenora, Wickham; terminalis Casey; Type USNM 39472; Casey bequest 1925; Liogluta terminalis (Casey) V.I. Gusarov 1998; cf. Liogluta aloconoides (USNM). Examined.</p><p>Atheta (Liogluta) renominata Bernhauer &amp; Scheerpeltz, 1926: 658 (nom. nov. for Anepsiota terminalis Casey, 1906 in Atheta, not Atheta terminalis Gravenhorst, 1806 and Gyllenhal, 1810; synonymized by Seevers 1978).</p><p>Liogluta (Anepsiota) aloconotoides Lohse, in Lohse et al. 1990: 165. New synonymy. Holotype (male): Canada, Labrador, L’Anse au Loup, 9.VIII.1972, J.M. Campbell (CNC). Paratypes: Canada, Labrador, Red Bay, 8.VIII.1972, J.M. Campbell (5, sex undetermined, CNC).</p><p>New locality data.</p><p>USA: Montana: Flathead Co., Glacier National Park, N Fork Flathead area, 1988, Red Bench Fire study; N Mud Lake, 3520 feet, lodgepole unburned T34N, R21W, Sec 1, 7.VI.1990, pitfall trap, M.A. Ivie (1 ♀, LFC). New Hampshire, Coos Co., Hwy. 16, 5-6 km S Gorham, 9.IX.1987, A. Smetana (1 ♂, 2 ♀, CNC).</p><p>Diagnosis.</p><p>This species may be distinguished by the following combination of characters: body subparallel, entirely reddish-brown or with head and posterior abdomen chestnut brown (Fig. 1); length 3.9-4.5 mm; integument of forebody with meshed microsculpture, moderately glossy; head about one-quarter narrower than maximum width of pronotum; pronotum transverse, narrower at base and widest in apical third; elytra at suture about as long as pronotum; basal three articles of metatarsus elongate, first longest, second about as long as third, fourth shorter than either of preceding articles; apical margin of male tergite VIII with broad, short, truncate projection with rounded lateral angles, apical margin smooth or slightly crenulate (Fig. 3); female tergite VIII with apical margin broadly, just visibly emarginate (Fig. 6); genital structures as illustrated (Figs 2, 7).</p><p>Natural history.</p><p>Klimaszewski et al. (2011) reported this species (as Liogluta aloconotoides) from various forest types and on coastal limestone barrens in Newfoundland. Specimens from New Brunswick were collected from dung in a coastal red spruce forest, by treading sedges along a small lake margin, from a Lindgren funnel trap deployed in a rich Appalachian hardwood forest with some conifers, and from a pitfall trap (Webster et al. 2012). In Alberta, adults were reared from well-decayed white spruce logs (Klimaszewski et al. 2015). Elsewhere, adults were captured in various forest types including a recently burned forest. The type specimens of Liogluta aloconotoides were captured in August (Lohse et al. 1990). Klimaszewski et al. (2015) reported adults from July to October.</p><p>Distribution.</p><p>Recorded from LB, NB, NF, NS, ON, QC, AB, YT, and BC (Casey 1906, Lohse et al. 1990, Klimaszewski et al. 2008, Majka and Klimaszewski 2008, Klimaszewski et al. 2011, Webster et al. 2012, Bousquet et al. 2013, Klimaszewski et al. 2015), and newly in USA from MT and NH.</p><p>Comments.</p><p>We have examined the female holotype of Liogluta terminalis (Casey) from Glenora, British Columbia, and compared it with the specimens of Liogluta aloconotoides Lohse east of the Rocky Mountains. We found no external or genital differences between the holotype of Liogluta terminalis and the other female specimens identified as Liogluta aloconotoides and therefore we consider Liogluta aloconotoides as a new synonym of Liogluta terminalis .</p></div>	https://treatment.plazi.org/id/3D5339B5E610D73EBD5C18424D565CBA	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Klimaszewski, Jan;Webster, Reginald P.;Langor, David W.;Sikes, Derek;Bourdon, Caroline;Godin, Benoit;Ernst, Crystal	Klimaszewski, Jan, Webster, Reginald P., Langor, David W., Sikes, Derek, Bourdon, Caroline, Godin, Benoit, Ernst, Crystal (2016): A review of Canadian and Alaskan species of the genus Liogluta Thomson, and descriptions of three new species (Coleoptera, Staphylinidae, Aleocharinae). ZooKeys 273: 217-256, DOI: http://dx.doi.org/10.3897/zookeys.573.7878, URL: http://dx.doi.org/10.3897/zookeys.573.7878
266DA326B15F0AA4AE8B5868A99D4526.text	266DA326B15F0AA4AE8B5868A99D4526.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Liogluta trapezicollis Lohse 1990	<div><p>Taxon classification Animalia Coleoptera Staphylinidae</p><p>Liogluta trapezicollis Lohse, 1990 Figs 8-15</p><p>Liogluta (Anepsiota) trapezicollis Lohse, in Lohse et al. 1990: 165. Holotype (male): Canada, Yukon Territory, Dempster Hwy., Mi. 60, 3500 ft., 19.VII.1978, J.M. Campbell and A. Smetana (CNC). Not examined.</p><p>New locality data.</p><p>Summarized for 146 specimens captured at 45 collection events from 6 major regions of Southeast Alaska, see http://dx.doi.org/10.7299/X79023ZM for the full data. USA: Alaska: Baranof Island (11 specimens, UAM), Chichagof Island (84 specimens, UAM), Dall Island (2 specimens, UAM), Haines, Flower Mountain (3 specimens, UAM), Hawthorne Peak (45 specimens, UAM), South Chilkat Peninsula (1 specimen, UAM). Excel file with locality data is available from LFC.</p><p>Diagnosis.</p><p>This species may be distinguished by the following combination of characters: body subparallel, slender, dark brown to black with pronotum brown and paler than head, elytra yellowish or reddish-brown (Fig. 8); length 3.8-4.4 mm; integument of forebody with meshed microsculpture moderately pronounced, surface moderately glossy; head about one-quarter narrower than maximum width of pronotum; pronotum transverse, narrower at base and widest at middle (width of pronotum variable, some specimens have pronotum markedly narrower than base of elytra and some have pronotum nearly as wide as base of elytra); elytra at suture slightly shorter than pronotum; basal three articles of metatarsus elongate, subequal in length and each slightly longer than fourth article; male tergite VIII with apical margin truncate, bordered by two short lateral teeth, variably sculptured and ranging from smooth to crenulate, or denticulate along margin (Fig. 11); genital structures as illustrated (Figs 9-15).</p><p>Natural history.</p><p>The holotype was collected in July (Lohse et al. 1990). The Alaskan specimens were collected in July only from alpine zones between 453 and 1071 m elevation, none were collected in lowland forests. Habitats include alpine flood meadows, under rocks, herbaceous heath with Luetkea, Cassiopes, and Lupinus, low rocky tundra with Dryas, meadow heath with Phyllodoce, Senecio, and Luetkea, shrubby krummholz with Elliottia and Tsuga, wet meadows with Carex, Petasites, Senecio and Ranunculus .</p><p>Distribution.</p><p>Canada: BC, YT (Klimaszewski et al. 2012). USA: AK (Lohse et al. 1990).</p><p>Comments.</p><p>We were not able to compare types of Liogluta trapezicollis Lohse with the specimens we examined, and our determinations are based on the published description by Lohse in Lohse et al. (1990). The types of Liogluta trapezicollis housed in the Canadian National Collection of Insects were borrowed several years ago by V. Gusarov (Oslo, Norway) and our persistent efforts to have these specimens returned to Canada have failed.</p><p>Five specimens in UAM were successfully DNA barcoded (UAM GUID, BOLD Process ID): UAM:Ento:145576, UAMIC2696-15; UAM:Ento:145623, UAMIC2740-15; UAM:Ento:152467, UAMIC2750-15; UAM:Ento:232527, UAMIC2677-15; UAM:Ento:232696, UAMIC2753-15. The DNA sequences for these specimens are all very similar (maximum distance of 0.32%, nearest neighbour of 3.13%) and fall within the same BIN (Barcode Index Number) (Ratnasingham and Hebert 2013), BOLD:ACU9772, which is not shared by any other species.</p></div>	https://treatment.plazi.org/id/266DA326B15F0AA4AE8B5868A99D4526	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Klimaszewski, Jan;Webster, Reginald P.;Langor, David W.;Sikes, Derek;Bourdon, Caroline;Godin, Benoit;Ernst, Crystal	Klimaszewski, Jan, Webster, Reginald P., Langor, David W., Sikes, Derek, Bourdon, Caroline, Godin, Benoit, Ernst, Crystal (2016): A review of Canadian and Alaskan species of the genus Liogluta Thomson, and descriptions of three new species (Coleoptera, Staphylinidae, Aleocharinae). ZooKeys 273: 217-256, DOI: http://dx.doi.org/10.3897/zookeys.573.7878, URL: http://dx.doi.org/10.3897/zookeys.573.7878
0ADA3B0D9C850125AC2A4DF09AFF5A37.text	0ADA3B0D9C850125AC2A4DF09AFF5A37.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Liogluta quadricollis (Casey 1894) Casey 1894	<div><p>Taxon classification Animalia Coleoptera Staphylinidae</p><p>Liogluta quadricollis (Casey, 1894) Figs 16-22</p><p>Anepsiota quadricollis Casey, 1894: 330. As Atheta (Liogluta): Bernhauer and Scheerpeltz 1926: 658. Holotype (female). Canada, British Columbia, Vancouver Island, Anepsiota quadricollis; Type USNM 39471 (USNM). Examined.</p><p>New locality data.</p><p>Canada: British Columbia: Hwy 5 at Juliet Creek, 25.IX.1994, Lot 2 [in forest under rocks in poplar stand], B.F. &amp; J.L. Carr, J. &amp; B. Carr Coll., bequested to CNC August 2000 (1 ♂, CNC).</p><p>Diagnosis.</p><p>This species may be distinguished by the following combination of characters: body subparallel, slender, bicoloured, pronotum orange and remainder of body dark brown to reddish-brown; length 4.1-4.3 mm (Fig. 16); integument of forebody with weak meshed microsculpture, surface highly glossy; head slightly narrower than pronotum; pronotum subquadrate; antennae enlarged and black to brown; elytra about as wide as pronotum and at suture about as long as pronotum; basal two articles of metatarsus distinctly elongate, subequal in length, each longer than third article. New description of male. Apical margin of tergite VIII with broad, obtusely triangular projection in almost middle half with rounded lateral angles (Fig. 18); sternite VIII elongate, rounded apically, with antecostal suture arcuate, well separated from basal margin (Fig. 19); median lobe of aedeagus with tubus short and sinuate, its subapical part narrowly elongate in lateral view, internal sac structures distinct (Fig. 17). Female. Tergite VIII with apical margin obtusely angulate, broadly rounded at middle (Fig. 20); sternite VIII shallowly, broadly emarginated medially (Fig. 21); spermatheca with a short club-shaped capsule and short apical invagination, stem long and highly sinuate (Fig. 22).</p><p>Natural history.</p><p>The holotype and the other BC specimen were collected in September, the Carrs collected a male under a rock in a poplar stand.</p><p>Distribution.</p><p>Vancouver Island, British Columbia (Casey 1894).</p></div>	https://treatment.plazi.org/id/0ADA3B0D9C850125AC2A4DF09AFF5A37	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Klimaszewski, Jan;Webster, Reginald P.;Langor, David W.;Sikes, Derek;Bourdon, Caroline;Godin, Benoit;Ernst, Crystal	Klimaszewski, Jan, Webster, Reginald P., Langor, David W., Sikes, Derek, Bourdon, Caroline, Godin, Benoit, Ernst, Crystal (2016): A review of Canadian and Alaskan species of the genus Liogluta Thomson, and descriptions of three new species (Coleoptera, Staphylinidae, Aleocharinae). ZooKeys 273: 217-256, DOI: http://dx.doi.org/10.3897/zookeys.573.7878, URL: http://dx.doi.org/10.3897/zookeys.573.7878
8DEA14C899858ECCEDC53E28C6657B30.text	8DEA14C899858ECCEDC53E28C6657B30.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Liogluta wickhami (Casey 1894) Casey 1894	<div><p>Taxon classification Animalia Coleoptera Staphylinidae</p><p>Liogluta wickhami (Casey, 1894) Figs 23-29</p><p>Anepsiota wickhami Casey, 1894: 331. As Atheta (Liogluta): Bernhauer and Scheerpeltz 1926: 656. Holotype (female): Canada, British Columbia; Stickeen River Canyon; Anepsiota wickhami; Type USNM 39474, Casey bequest 1925 (USNM). Examined.</p><p>New locality data.</p><p>Canada: British Columbia: Mi. 56 Haines Hwy., Three Guardsmen Pass, 4.VII.1968, 3200 feet, J.M. Campbell and A. Smetana (1 ♂, CNC).</p><p>Diagnosis.</p><p>This species may be distinguished by the following combination of characters: body broadly subparallel (Fig. 23); pronotum, elytra, legs and basal antennal article reddish-brown, head and abdomen chestnut brown (Fig. 23); length 4.0-4.2 mm; integument of forebody with meshed microsculpture, moderately glossy; head about one-third narrower than maximum width of pronotum; pronotum more or less evenly arcuate laterally (Fig. 23); elytra at suture about as long as pronotum; basal three articles of metatarsus missing in holotype (Fig. 23). New description of male. Apical margin of tergite VIII with very broad, short, subtruncate projection with rounded lateral angles, with apical margin faintly crenulate (Fig. 25); sternite VIII broadly rounded apically (Fig. 26); median lobe of aedeagus with tubus slightly arcuate ventrally, with apex narrow and rounded (Fig. 24). Female. Tergite VIII truncate apically (Fig. 27); sternite VIII very slightly broadly emarginate apically (Fig. 28); spermatheca with tubular capsule and deep apical invagination, stem thin, long, and highly sinuate (Fig. 29).</p><p>This species is similar to Liogluta terminalis but has dark brown antennae, head and pronotum (antennae, head, and pronotum are uniformly reddish-brown or only slightly darker than remaining parts of the body in Liogluta terminalis). Spermatheca is differently shaped in each species; Liogluta wickhami has smaller and differently shaped capsule with a deep apical invagination and has a shorter and differently looped posterior stem (Fig. 29). Liogluta wickhami is also very similar to Liogluta vasta but can be distinguished by the shape of pronotum which has evenly arcuate sides and is broadest at middle (Fig. 23), while it is trapezoidal in shape and is broadest in apical third in the latter species (Fig. 30).</p><p>Natural history.</p><p>Unknown.</p><p>Distribution.</p><p>The female holotype was captured in the Stickeen River Valley of British Columbia (Casey 1894), and one male was found in Three Guardsmen Pass, British Columbia.</p></div>	https://treatment.plazi.org/id/8DEA14C899858ECCEDC53E28C6657B30	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Klimaszewski, Jan;Webster, Reginald P.;Langor, David W.;Sikes, Derek;Bourdon, Caroline;Godin, Benoit;Ernst, Crystal	Klimaszewski, Jan, Webster, Reginald P., Langor, David W., Sikes, Derek, Bourdon, Caroline, Godin, Benoit, Ernst, Crystal (2016): A review of Canadian and Alaskan species of the genus Liogluta Thomson, and descriptions of three new species (Coleoptera, Staphylinidae, Aleocharinae). ZooKeys 273: 217-256, DOI: http://dx.doi.org/10.3897/zookeys.573.7878, URL: http://dx.doi.org/10.3897/zookeys.573.7878
311DED817411DB63D88AB88042395100.text	311DED817411DB63D88AB88042395100.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Liogluta vasta (Mӓklin 1853) M ӓklin 1853	<div><p>Taxon classification Animalia Coleoptera Staphylinidae</p><p>Liogluta vasta (Mӓklin, 1853) Figs 30-34</p><p>Homalota vasta Mӓklin, 1853: 183. As Atheta (Liogluta): Moore and Legner 1975: 376; as Liogluta: Lohse and Smetana 1985: 297, Gusarov 2003a: 39. Lectotype (female): USA, Alaska: Sitka; Holmberg; Mus. Zool. H:fors, Spec. Typ. No. 2251, Homalota vasta Mӓkl.; Mus. Zool. Helsinki, Loan No. C98-138; Paralectotypus Homalota vasta Mӓklin, Lohse des. 1985, Gusarov rev. 2000; http://id.luomus.fi/GAC. 16963, UNITED STATES Alaska, Sitka, 57.1483N, 135.23W, Holmberg leg. Examined. We have added a new lectotype label (see discussion below) [there was no original label designated by Lohse]. Paralectotype (female): USA, Alaska: Sitka; Holmberg; Homalota vasta m. Sitkcha [Sitka]; Mus. Zool. Typ. No. 2250, Homalota vasta Mӓkl.; typus; Mus. Zool. Helsinki, Loan No. C 14527; Mus. Zool. Helsinki, Loan No. C 98; Lectotypus Homalota vasta Mӓklin, Lohse des. 1985, Gusarov rev. 2000; http://id.luomus.fi/GAC. 16962, UNITED STATES Alaska, Sitka, 57.1483N, 135.23W, Holmberg leg. Examined. We have added a new Paralectotype label (see discussion below) [there was no original label designated by Lohse].</p><p>Diagnosis</p><p>(based on female lectotype, male unknown). This species is very similar to Liogluta wickhami, but in Liogluta vasta the forebody is less reddish, and the pronotum is more trapezoidal and narrowest at base (Fig. 30); the apical margin of tergite VIII is broadly arcuate with the antecostal suture very narrowly separated from the basal margin (Fig. 32); the apical margin of sternite VIII is slightly, very broadly emarginate with the antecostal suture highly sinuate and well separated from the basal margin (Fig. 33).</p><p>The female lectotype is missing the spermatheca. The spermatheca of the Yukon specimen in CNC cited by Lohse and Smetana (1985), tentatively identified as belonging to this species, is illustrated in Fig. 34 (after Lohse and Smetana 1985). Males and more females from the type locality are needed to clearly define this species, which is here tentatively listed as a valid species. When more specimens of Liogluta vasta become available for study and the morphological variation is known, we will be able to understand the relationship of this species to other nearctic Liogluta species. Liogluta vasta is also similar to some specimens (with broad pronotum) of Liogluta trapezicollis . A DNA comparison between Liogluta vasta and other Liogluta species is needed to clarify its identity and relationships.</p><p>Distribution.</p><p>Canada: YT?. USA: AK.</p><p>Discussion.</p><p>The original type material of Homalota vasta Mӓklin, 1853 (ZMH) consists of two female specimens representing two different species in two genera, Atheta (as Boreophilia in Lohse and Smetana 1985) and Liogluta . Lohse and Smetana (1985) designated the female specimen belonging to Liogluta as the lectotype of Homalota vasta Mӓklin. However, the label data published by Lohse and Smetana (1985) for the Liogluta specimen corresponded to the Atheta specimen (see also discussion in Gusarov 2003b, who also mislabelled the specimens). We consider Lohse and Smetana’s lectotype designation as valid regardless of the obvious mistake of publishing the wrong label data; therefore, the name vasta is affiliated with Liogluta . The female paralectotype belongs to Atheta keeni Casey, 1910.</p><p>It is noteworthy that despite years of intensive collections made primarily between 2008-2013 in southeast Alaskan lowland forests and alpine zones, including in and around Sitka, which have resulted in 22,029 specimens of Staphylinidae (http://arctos.database.museum/saved/SE-AK-Staphylinidae), no specimens of Liogluta vasta were found.</p></div>	https://treatment.plazi.org/id/311DED817411DB63D88AB88042395100	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Klimaszewski, Jan;Webster, Reginald P.;Langor, David W.;Sikes, Derek;Bourdon, Caroline;Godin, Benoit;Ernst, Crystal	Klimaszewski, Jan, Webster, Reginald P., Langor, David W., Sikes, Derek, Bourdon, Caroline, Godin, Benoit, Ernst, Crystal (2016): A review of Canadian and Alaskan species of the genus Liogluta Thomson, and descriptions of three new species (Coleoptera, Staphylinidae, Aleocharinae). ZooKeys 273: 217-256, DOI: http://dx.doi.org/10.3897/zookeys.573.7878, URL: http://dx.doi.org/10.3897/zookeys.573.7878
5831279A239410B9DEEB5CE923955057.text	5831279A239410B9DEEB5CE923955057.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Liogluta nigropolita (Bernhauer 1907) Bernhauer 1907	<div><p>Taxon classification Animalia Coleoptera Staphylinidae</p><p>Liogluta nigropolita (Bernhauer, 1907) Figs 35-43</p><p>Atheta nigropolita Bernhauer, 1907: 390. As Liogluta: Lohse and Smetana 1985: 286.</p><p>Liogluta nigropolita Syntype (male): USA, New Hampshire, Mt. Washington (FMNH).</p><p>New locality data.</p><p>CANADA: Quebec: Gt. Whale Riv., 5.IX.1949, J.R. Vockeroth (1 sex undetermined, CNC). Northwest Territories: Yellowknife, 62.50714°N, 113.39443°W, 236 m, mesic habitat replicate #2, Yellow Pan Trap #2, 15-18.VI.2011, col. NBP Field Party (1 ♀, LFC).</p><p>Diagnosis.</p><p>This species may be distinguished by the following combination of characters: body elongate with elytra and abdomen broad, moderately robust, dark brown to black with tarsi and tibiae often reddish-brown, elytra sometimes with reddish tinge (Figs 35); length 3.5-4.0 mm; integument of forebody with moderately pronounced meshed microsculpture, surface highly glossy; head about one-eighth narrower than maximum width of pronotum (Fig. 35); pronotum transverse, about evenly wide in posterior half, then distinctly narrowing apicad, forming round apical angles (Fig. 35); elytra at suture slightly longer than pronotum (Fig. 35); basal four articles of metatarsus about the same length, each shorter than fifth article. Male. Tergite VIII with apical margin broadly arcuate, without teeth or crenulations (Fig. 38); sternite VIII rounded apically (Fig. 39); median lobe of aedeagus with tubus broadly arcuate ventrally and with apex narrow and pointed in lateral view (Fig. 36); tubus broad and triangular apically in dorsal view (Fig. 37). Female. Tergite VIII broadly arcuate apically (Fig. 40); sternite VIII evenly rounded apically, with antecostal suture distinctly sinuate (Fig. 41); spermatheca with spherical capsule with invagination deep and narrow, stem S-shaped, gradually becoming very narrow posteriad (Figs 42, 43).</p><p>Natural history.</p><p>Adults occur in moss and leaf litter (Lohse et al. 1990).</p><p>Distribution.</p><p>Canada: LB, NF, NT, NU, QC, YT. USA: NH (Lohse et al. 1990; Klimaszewski et al. 2012; Bousquet et al. 2013).</p><p>Comments.</p><p>This species is probably transcontinental in northern Canada.</p></div>	https://treatment.plazi.org/id/5831279A239410B9DEEB5CE923955057	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Klimaszewski, Jan;Webster, Reginald P.;Langor, David W.;Sikes, Derek;Bourdon, Caroline;Godin, Benoit;Ernst, Crystal	Klimaszewski, Jan, Webster, Reginald P., Langor, David W., Sikes, Derek, Bourdon, Caroline, Godin, Benoit, Ernst, Crystal (2016): A review of Canadian and Alaskan species of the genus Liogluta Thomson, and descriptions of three new species (Coleoptera, Staphylinidae, Aleocharinae). ZooKeys 273: 217-256, DOI: http://dx.doi.org/10.3897/zookeys.573.7878, URL: http://dx.doi.org/10.3897/zookeys.573.7878
138BA9E24E5EB5220EB7FBBE7401E816.text	138BA9E24E5EB5220EB7FBBE7401E816.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Liogluta nitens (Mӓklin 1852) M ӓklin 1852	<div><p>Taxon classification Animalia Coleoptera Staphylinidae</p><p>Liogluta nitens (Mӓklin, 1852) Figs 44-49</p><p>Homalota nitens Mӓklin, 1852: 307. As Liogluta: Lohse and Smetana 1985: 288.</p><p>Liogluta nitens (male): USA, Alaska: Sitcha [Sitka], Holmberg, coll Mӓklin; Homalota nitens Mkln., Sitka pr. Hlm. Berg (ZMH). Designated by Lohse and Smetana (1985).</p><p>Liogluta apposita (Casey, 1911). Synonymized by Gusarov 2003b [type locality BC: Metlakatla].</p><p>Liogluta insolens (Casey, 1910). Synonymized by Gusarov 2003b [type locality BC: Queen Charlotte Islands: Massett].</p><p>Liogluta resplendens (Casey, 1910). Synonymized by Gusarov 2003b [type locality: BC: Queen Charlotte Islands].</p><p>New locality data.</p><p>CANADA: Alberta: 28 km NW Hinton, 0.5 km S of Rock Lake Road, 53.524°N, 117.957°W, Ecosite Surrogacy Study, Ecoregion: UF, Ecosite C2, Stand C205, pitfall trap 2, 5, 14.V-4.VI.2004, J. Hammond et al. (1 ♂, 9 ♀, NoFC); same data except: 30.VII-13.VIII.2004, pitfall trap 5 (3 ♂, 2 ♀, NoFC); 23.1 km NW Hinton, W.A. Switzer Prov. Pk., 53.560°N, 117.808°W, Ecosite Surrogacy Study, Ecoregion: UF, Ecosite F2, Stand F214, pitfall trap 5, 31.VII-13.VIII.2004, J. Hammond et al. (1 ♂, NoFC); same data except: Stand F216, pitfall trap 4 (1 ♀, NoFC); 32 km NW Hinton, 0.5 km E Wild Hay Campgr., 53.529°N, 117.946°W, Ecosite Surrogacy Study, Ecoregion: UF, Ecosite F2, Stand F216, pitfall trap 3, 16-30.VII.2004, J. Hammond et al. (1 ♂, 1 ♀, NoFC); same data except: pitfall trap 4, 30.VII-13.VIII.2004 (1 ♂, NoFC); 57 km N Hinton, 1.5 km W of J. Wright Rd., 53.921°N, 117.617°W, Ecosite F1, Stand F101, pitfall trap 2, 28.VII-11.VIII.204, J. Hammond et al. (2 ♀, NoFC); 62 km N Hinton, 5 km W of J. Wright Rd., 53.969°N, 117.668°W, Stand F105, 30.VI-14.VII.2004, J. Hammond et al. (1 ♀, NoFC); same data except: 53.921°N, 117.663°W, Stand F202, pitfall trap 1, 28.VII-11.VIII.2004, J. Hammond et al. (1 ♂, NoFC).</p><p>Diagnosis.</p><p>This species may be distinguished by the following combination of characters: body narrowly elongate, robust, dark-brown to black with elytra, tarsi and tibiae often yellowish- or reddish-brown (Fig. 44); length 2.8-3.3 mm; integument of forebody with moderately pronounced meshed microsculpture, surface highly glossy (Fig. 44); head about one-eighth narrower than maximum width of pronotum (Fig. 44); pronotum transverse, about evenly wide in basal half and then distinctly narrowing anteriad (Fig. 44); elytra at suture about as long as pronotum (Fig. 44); basal four articles of metatarsus about same length, each shorter than fifth article. Male. Tergite VIII with short subrectangular projection on more than half width of apical margin, with rounded lateral angles, apical margin smooth or micro-crenulate (Fig. 46); sternite VIII parabolically rounded apically (Fig. 47); median lobe of aedeagus with tubus almost straight in lateral view, with apex moderately narrow, rounded (Fig. 45). Female. Tergite VIII truncate apically (Fig. 48); sternite VIII with apical margin evenly rounded, antecostal suture slightly sinuate (Fig. 49). Spermatheca unknown.</p><p>Natural history.</p><p>Adults were captured using pitfall traps in Carmanah Valley, Vancouver Island, from June to September, with the peak catch in June (Klimaszewski and Winchester 2002). They were found mainly in the interior and transition zones of a Sitka spruce forest (Klimaszewski and Winchester 2002). Several adults were collected from moss at the edge of an old road in the Queen Charlotte Islands, British Columbia. In Alberta, adults were collected in pitfall traps in various forest types in the Upper Cordilleran Ecoregion. Adults in Alaska were collected in a wide variety of habitats spanning lowland forests to alpine zones: alpine meadow litter, lowland forest clearcuts, floodplain meadows with Athyrium, Caltha, and Rubus, under rocks, in krummholz alpine habitats of Tsuga mertensiana, near bear dung in alpine habitats, old growth temperate rain coniferous forests, alpine heath with Empetrum, and Vaccinium, subalpine habitats with Salix, and Veratrum .</p><p>Distribution.</p><p>Canada: AB, BC, YT. USA: AK, OR, WA (Mӓklin 1852, Bernhauer 1907, Hatch 1957, Moore and Legner 1975, Lohse and Smetana 1985, Klimaszewski and Winchester 2002).</p><p>Comments .</p><p>There is considerable variation in length and width of elytra in specimens from Vancouver Island, Oregon (having broader and longer elytra), and those with narrow and shorter elytra from the Queen Charlotte Islands, Alberta, and Alaska. The genitalic features were the same in those of the typical form with the longer and broader elytra, and those with narrower and shorter elytra. Therefore, we consider this as intraspecific variation. Additional studies, including DNA comparison, are needed to reveal the relationship between these two morphotypes. Two UAM Alaskan specimens (UAM:Ento:152502, UAM:Ento:232546) were DNA barcoded (UAMIC2665-15, UAMIC2701-15) and they cluster closely with two specimens of this species DNA barcoded from Alberta, Canada.</p></div>	https://treatment.plazi.org/id/138BA9E24E5EB5220EB7FBBE7401E816	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Klimaszewski, Jan;Webster, Reginald P.;Langor, David W.;Sikes, Derek;Bourdon, Caroline;Godin, Benoit;Ernst, Crystal	Klimaszewski, Jan, Webster, Reginald P., Langor, David W., Sikes, Derek, Bourdon, Caroline, Godin, Benoit, Ernst, Crystal (2016): A review of Canadian and Alaskan species of the genus Liogluta Thomson, and descriptions of three new species (Coleoptera, Staphylinidae, Aleocharinae). ZooKeys 273: 217-256, DOI: http://dx.doi.org/10.3897/zookeys.573.7878, URL: http://dx.doi.org/10.3897/zookeys.573.7878
C35069FEB50F14003F74EE37EA1D0CBF.text	C35069FEB50F14003F74EE37EA1D0CBF.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Liogluta granulosa Lohse 1990	<div><p>Taxon classification Animalia Coleoptera Staphylinidae</p><p>Liogluta granulosa Lohse, 1990 Figs 50-57</p><p>Liogluta (Liogluta) granulosa Lohse, in Lohse et al. 1990: 164. Holotype (male): USA, Alaska, King Salmon, Naknek R. Alaska, 6.VII.1952, W.R. Mason, No. 20313 (CNC). Examined.</p><p>New locality data.</p><p>CANADA: Yukon Territory: location EMAN Plot, Cadet Camp, EP-Yukon, 15.X.2001 (1 ♀, NoFC); Tombstone Mts., 64.60560°, 138.36413°, Rep. 1, mesic, yellow pan trap, 21-24.VI.2011, NBP Field Party (1 ♀, LFC).</p><p>USA: Alaska: Quinhagak site G, 3 m elevation, 59.71035°, 161.89102°, dry tundra, between Rubus sp. hummocks, pitfall, 18-26, VIII.2014, V. Forbes (1 ♂, LFC) [submitted for barcoding]; Naknek, 58.73973°N - 157.0636°W, 2-5 m elev., creekside/ocean beach confluence, under boards and driftwood 10.VI.2007, D.S. Sikes. UAM:Ento:29798 (1 ♂, UAM) [DNA barcoded: http://arctos.database.museum/guid/UAM:Ento:29798].</p><p>Diagnosis.</p><p>This species may be distinguished by the following combination of characters: body broadly subparallel, dark brown, with elytra, tarsi and tibiae often reddish-brown (Fig. 50) (one specimen from northern Yukon was entirely black); length 2.8-3.3 mm; integument of forebody with moderately pronounced meshed microsculpture; head about one-eighth narrower than maximum width of pronotum (Fig. 50); pronotum transverse, about evenly wide in basal one-third of its length, then strongly broadest at apical one-third and gradually narrowed apically (Fig. 50); elytra at suture about as long as pronotum, its surface coarsely granulose (Fig. 50); basal two articles of metatarsus about the same length, each shorter than fifth article. Male. Apical margin of tergite VIII with short, very obtusely angular projection in medial two-thirds with rounded lateral angles, margin of projection smooth or micro-denticulate (Fig. 53); apical margin of sternite VIII broadly parabolic (Fig. 54); median lobe of aedeagus with tubus broadly arched, bent ventrad, apex narrow and rounded (Fig. 52). Female. Apical margin of tergite VIII truncate in middle one-third (Fig. 55); apical margin of sternite VIII arcuate, antecostal suture distinctly sinuate (Fig. 56); spermatheca highly sinuate as illustrated (Fig. 57).</p><p>Natural history.</p><p>Adults were captured in June, July, August, and October. One Alaskan specimen was captured in tundra between Rubus species and another at a creekside/ocean beach confluence, under boards and driftwood.</p><p>Distribution.</p><p>Canada: YT. USA: AK (Lohse et al. 1990, Klimaszewski et al. 2008, Klimaszewski et al. 2012).</p><p>Comments.</p><p>Only a few specimens of this species are known. Its distribution is nordic and the habitat is unknown. One specimen (UAM:Ento:29798) in UAM was DNA barcoded (UAMIC2693-15), the first and only for this species so far.</p></div>	https://treatment.plazi.org/id/C35069FEB50F14003F74EE37EA1D0CBF	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Klimaszewski, Jan;Webster, Reginald P.;Langor, David W.;Sikes, Derek;Bourdon, Caroline;Godin, Benoit;Ernst, Crystal	Klimaszewski, Jan, Webster, Reginald P., Langor, David W., Sikes, Derek, Bourdon, Caroline, Godin, Benoit, Ernst, Crystal (2016): A review of Canadian and Alaskan species of the genus Liogluta Thomson, and descriptions of three new species (Coleoptera, Staphylinidae, Aleocharinae). ZooKeys 273: 217-256, DOI: http://dx.doi.org/10.3897/zookeys.573.7878, URL: http://dx.doi.org/10.3897/zookeys.573.7878
433C71A6DB31979836398DDE00E2E691.text	433C71A6DB31979836398DDE00E2E691.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Liogluta microgranulosa Klimaszewski & Webster	<div><p>Taxon classification Animalia Coleoptera Staphylinidae</p><p>Liogluta microgranulosa Klimaszewski &amp; Webster sp. n. Figs 58-64</p><p>Holotype</p><p>(male). Canada, New Brunswick, Restigouche Co., Jacquet River Gorge P.N.A., 47.7361°N, 66.0778°W, 16.VIII.2010, R.P. Webster // beaver dam, among sticks and debris near an overflow area of dam (near flowing water) (LFC). Paratypes: same data as holotype (1 ♂, 3 ♀, LFC; 2 ♂, 7 ♀, NBM; 4 ♂, 5 ♀, 1 sex undetermined, RWC); Jacquet River Gorge P.N.A., 47.7357°N, 66.0774°W, 24.VII.2008, R.P. Webster// Margin of pond, among leaves and sedges near pond margin (1♀, LFC). York Co., Fredericton, 45.9361°N, 66.6747°W, 17.VIII.2009, R.P. Webster // Beaver dam, outer margin under overhanging sticks near water (1 ♀, RWC).</p><p>Etymology.</p><p>Microgranulosa is a Latin adjective meaning microgranulate, in reference to the minute sculpture on the elytra of this species.</p><p>Description.</p><p>This species may be distinguished by the following combination of characters: body narrowly subparallel; head, apical articles of antennae, and posterior part of abdomen black, elytra brownish and mottled with black, remaining parts reddish-brown (Fig. 58); length 4.6-5.1 mm; integument of forebody with moderately pronounced meshed microsculpture, surface moderately glossy (Fig. 58); head about one-quarter narrower than maximum width of pronotum (Fig. 58); pronotum transverse, about evenly wide in basal half of its length, then strongly narrowed apically (Fig. 58); elytra at suture about as long as pronotum, surface finely and densely microgranulose; basal three articles of metatarsus about equally elongate, each longer than fourth article. Male. Apical margin of tergite VIII with very broad, very obtusely angular projection, with obtuse lateral angles and small tooth medially, margin often micro-crenulate (Fig. 60); sternite VIII rounded apically (Fig. 61); median lobe of aedeagus with tubus distinctly arched ventrad in apical half, apical part narrow (Fig. 59). Female. Tergite VIII with apical margin obtusely angulate (Fig. 62); sternite VIII with apical margin slightly emarginate medially (Fig. 63); spermatheca with stem long, sinuate, spiral posteriorly, capsule club-shaped with apical invagination deep and narrow (Fig. 64).</p><p>Distribution.</p><p>Canada: Known only from New Brunswick, Canada.</p><p>Natural history.</p><p>Nearly all adults from New Brunswick were collected from American beaver ( Castor canadensis Kuhl) dams. Most were collected from among sticks and debris near an overflow area of the dam, another from under overhanging sticks on the outer margin of the dam. One individual was collected from among leaves and sedges near a pond margin. Specimens were collected in July and August.</p></div>	https://treatment.plazi.org/id/433C71A6DB31979836398DDE00E2E691	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Klimaszewski, Jan;Webster, Reginald P.;Langor, David W.;Sikes, Derek;Bourdon, Caroline;Godin, Benoit;Ernst, Crystal	Klimaszewski, Jan, Webster, Reginald P., Langor, David W., Sikes, Derek, Bourdon, Caroline, Godin, Benoit, Ernst, Crystal (2016): A review of Canadian and Alaskan species of the genus Liogluta Thomson, and descriptions of three new species (Coleoptera, Staphylinidae, Aleocharinae). ZooKeys 273: 217-256, DOI: http://dx.doi.org/10.3897/zookeys.573.7878, URL: http://dx.doi.org/10.3897/zookeys.573.7878
C5ADE7F59540220AD9B86B749FFCC754.text	C5ADE7F59540220AD9B86B749FFCC754.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Liogluta castoris Klimaszewski & Webster	<div><p>Taxon classification Animalia Coleoptera Staphylinidae</p><p>Liogluta castoris Klimaszewski &amp; Webster sp. n. Figs 65-71</p><p>Holotype</p><p>(male). Canada, New Brunswick, York Co., Charters Settlement, 45.8395°N, 66.7391°W, 21.IV.2010, R.P. Webster coll. //Mixed forest opening, collected with net during evening flight between 16:30 and 19:00 h (LFC). Paratypes. York Co., same data as holotype except (2 ♀, RWC); same data as holotype except: 17.VI.2005 // mixed forest in flight (1 ♂, LFC) [barcoded BIO]; same data as holotype except 23.IV.2008 // Mixed forest, in flight, collected with net between 15:00 and 18:00 h (1 ♂, RWC); same data as holotype except 5.IV.2010 // Mixed forest opening, collected with net during evening flight between 16:30 and 19:00 h (1 ♂, RWC); Charters Settlement, 45.8456°N, 66.7267°W, 5.V.2010, 16.V.2010, beaver dam, among sticks and debris near overflow area of dam, near flowing water (1 ♀, LFC; 2 ♂, 1 ♀, RWC); Charters Settlement, 45.8331°N, 66.7279°W, 20.V.2010, among sticks and debris near overflow area of dam, near flowing water (1 ♂, RWC). Saint John Co., ca 2 km NE of Maces Bay, 45.1161°N, 66.4560°W, 8.V.2006, R.P. Webster, eastern white cedar swamp, in sphagnum and litter near brook (1 ♀, RWC). Nova Scotia: Cape Breton H.N.P., North Mtn., 15.VIII.1983, J.E.H. &amp; R.J. Martin (1 ♂, CNC); Cape Breton H.N.P., Lone Shieling, PG729861, 19.VI.1983, Y. Bousquet, interception trap (1 ♂, CNC); Cape Breton H.N.P., Lone Shieling, PG729861, 3.VI.1983, H. Goulet, Pans, Malaise (1 ♂, CNC). Québec: Gatineau Pk., near Mud Lake, 24.X.1967, A. Smetana (2 ♂, CNC).</p><p>Etymology.</p><p>Castoris is a Latin adjective derived from the name of the American beaver ( Castor canadensis Kuhl), in reference to beaver dams where some of the type specimens were captured.</p><p>Description.</p><p>Body length 4.6-5.4 mm, subparallel (Fig. 65); head and at least apical part of abdomen dark brown with pronotum, elytra, basal articles of antennae and legs yellowish to reddish-brown; integument moderately glossy, more so on posterior abdomen; forebody with minute and sparse punctation and sparse pubescence (Fig. 65); elytra with minute micro-granulation; head rounded and narrowed posteriorly, with large eyes, each about as long as postocular area in dorsal view (Fig. 65); antennae with articles V-X subquadrate to slightly elongate (Fig. 65); pronotum slightly transverse, broadly rounded laterally, slightly wider than head and narrower than elytra, pubescence directed latero-posteriad from midline of disc (Fig. 65); elytra transverse, at suture as long as pronotum, slightly longer laterally, with pubescence directed posteriad (Fig. 65); abdomen subparallel for most of its length, about as wide as elytra (Fig. 65). Male. Aedeagus with bulbus narrowly oval, median lobe with apical half of tubus slightly arched ventrad, apical part moderately broad in lateral view (Fig. 66); internal sac with few pronounced structures/membrane folds (Fig. 66); apical margin of tergite VIII with very broad truncate projection with obtuse lateral angles, with margin smooth or minutely crenulate (Fig. 67); apical margin of sternite VIII rounded (Fig. 68). Female. Tergite VIII with apical margin broadly rounded (Fig. 69); sternite VIII scarcely emarginate apically, antecostal suture distinctly sinuate, well separated from basal margin (Fig. 70); spermatheca with stem long, sinuate, twisted posteriorly, capsule tubular, with apical invagination narrow, short (Fig. 71).</p><p>Natural history.</p><p>In New Brunswick, adults were collected using an aerial (butterfly) net in a mixed forest opening during evening flights (between 15:00 and 19:00 h) during April and May. A number of individuals were collected from among sticks and debris near the overflow area of a beaver dam during May. One individual was sifted from sphagnum and litter near a brook in an eastern white cedar swamp in May. In Nova Scotia, specimens were captured in flight interception, pan, and Malaise traps during the months of June and August. The single specimen from Ontario was captured in October.</p><p>Distribution .</p><p>Canada: Known from NB, NS, QC.</p><p>Comments.</p><p>This species is similar to Liogluta microgranulosa but in Liogluta castoris the pronotum and elytra are more elongate and more reddish-brown (Fig. 65); the median lobe of the aedeagus has the apical part of the tubus broader and shorter in lateral view (Fig. 66); male tergite VIII is truncate and not at all angulate medially (Fig. 67); the spermatheca has a longer stem (Fig. 71); and female sternite VIII has an apical emargination which is much less noticeable and the antecostal suture is more distinctly sinuate (Fig. 70).</p></div>	https://treatment.plazi.org/id/C5ADE7F59540220AD9B86B749FFCC754	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Klimaszewski, Jan;Webster, Reginald P.;Langor, David W.;Sikes, Derek;Bourdon, Caroline;Godin, Benoit;Ernst, Crystal	Klimaszewski, Jan, Webster, Reginald P., Langor, David W., Sikes, Derek, Bourdon, Caroline, Godin, Benoit, Ernst, Crystal (2016): A review of Canadian and Alaskan species of the genus Liogluta Thomson, and descriptions of three new species (Coleoptera, Staphylinidae, Aleocharinae). ZooKeys 273: 217-256, DOI: http://dx.doi.org/10.3897/zookeys.573.7878, URL: http://dx.doi.org/10.3897/zookeys.573.7878
92BCA175D3BA8F17C2C06CA3377214AB.text	92BCA175D3BA8F17C2C06CA3377214AB.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Liogluta pseudocastoris Klimaszewski & Webster	<div><p>Taxon classification Animalia Coleoptera Staphylinidae</p><p>Liogluta pseudocastoris Klimaszewski &amp; Webster sp. n. Figs 72-78</p><p>Holotype</p><p>(male). Canada, New Brunswick, York Co., Charters Settlement, 45.8456°N, 66.7267°W, 10.VI.2010, R.P. Webster, coll., beaver dam among sticks and debris near an overflow area of dam, near flowing water (LFC). Paratypes. same data as holotype: (2 ♂, 1 ♀, RWC): same data as holotype except 16.V.2010 (2 ♀, RWC); New Brunswick: York Co., Charters Settlement, 45.8395°N, 66.7391°W, 3.V.2012, R.P. Webster, mixed forest opening, during evening flight between 16:30 and 19:00 h (1 ♂, LFC [barcoded BIO]; 1 ♀, RWC).</p><p>Etymology.</p><p>Pseudocastoris is the Latin prefix pseudo-, false, added to the species name castoris, reflecting the close similarity of the two species.</p><p>Description.</p><p>Body length 3.9-4.4 mm, subparallel; dark brown with irregularly shaped lighter areas on pronotum in some individuals, head and abdomen dark brown, antennae dark, and legs yellowish; integument moderately glossy, more so on posterior portion of abdomen; forebody with minute and sparse punctation and sparse pubescence (Fig. 72); elytra with micro-granulation (Fig. 72); head rounded and narrowed posteriorly, eyes large, each about as long as postocular area in dorsal view (Fig. 72); antennae with articles V-X subquadrate to slightly transverse (Fig. 72); pronotum transverse, broadly rounded laterally, slightly wider than head and narrower than elytra, pubescence directed latero-posteriad from midline of disc (Fig. 72); elytra transverse, at suture about as long as pronotum, slightly longer laterally, with pubescence directed posteriad; abdomen subparallel for most of its length, about as wide as elytra (Fig. 72). Male. Tergite VIII broadly rounded apically, margin smooth (Fig. 74); apical margin of sternite VIII broadly parabolic (Fig. 75); median lobe of aedeagus with bulbus narrowly oval, tubus almost straight with apical part narrowly rounded in lateral view (Fig. 73); internal sac without distinct sclerites but with some vaguely-shaped structures (Fig. 73). Female. Tergite VIII broadly rounded apically (Fig. 76); sternite VIII a little less broadly rounded apically, antecostal suture slightly sinuate, moderately separated from basal margin (Fig. 77); spermatheca with capsule club-shaped, [invagination not perceptible], stem sinuate, about equally narrow throughout with only posterior part enlarged but not twisted (Fig. 78).</p><p>Natural history.</p><p>Most individuals were collected from among sticks and debris near an overflow area of a beaver dam during May and June. Others were collected using an aerial (butterfly) net in a mixed forest opening during an evening flight (between 16:30 and 19:00 h) during May.</p><p>Distribution.</p><p>Known only from New Brunswick, Canada.</p><p>Comments.</p><p>This species is closely related to Liogluta castoris and Liogluta microgranulosa but in Liogluta pseudocastoris the body is darker, particularly the pronotum, the pronotum is strongly narrowed basally with more angular posterior angles (Fig. 72); the shape of the median lobe of the aedeagus is different in lateral view, with the apical part narrower and very slightly arched ventrad (Fig. 73); the apical margin of male tergite VIII is evenly rounded (Fig. 74); the apical margin of female sternite VIII is not emarginate, with the antecostal suture only slightly sinuate (Fig. 77), and the shape of the spermatheca is different, with the posterior part of the stem enlarged but not twisted (Fig. 78).</p></div>	https://treatment.plazi.org/id/92BCA175D3BA8F17C2C06CA3377214AB	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Klimaszewski, Jan;Webster, Reginald P.;Langor, David W.;Sikes, Derek;Bourdon, Caroline;Godin, Benoit;Ernst, Crystal	Klimaszewski, Jan, Webster, Reginald P., Langor, David W., Sikes, Derek, Bourdon, Caroline, Godin, Benoit, Ernst, Crystal (2016): A review of Canadian and Alaskan species of the genus Liogluta Thomson, and descriptions of three new species (Coleoptera, Staphylinidae, Aleocharinae). ZooKeys 273: 217-256, DOI: http://dx.doi.org/10.3897/zookeys.573.7878, URL: http://dx.doi.org/10.3897/zookeys.573.7878
BCCAA8F4D45A6EF52B4934AC960581B3.text	BCCAA8F4D45A6EF52B4934AC960581B3.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Liogluta intermedia Klimaszewski & Langor	<div><p>Taxon classification Animalia Coleoptera Staphylinidae</p><p>Liogluta intermedia Klimaszewski &amp; Langor Figs 79-85</p><p>Liogluta intermedia Klimaszewski &amp; Langor, 2011: 168. Holotype (female): Canada, Newfoundland, Baie Verte Pen., 10 km SE Pumbly Cove, 49.68°N, 56.62°W, 3.X.2006, Site D, ex pitfall trap in riparian forest// NL Dept. Env. &amp; Conserv., Riparian Biodiversity Study, Site D Trap C5, (LFC).</p><p>New locality data.</p><p>CANADA: Newfoundland: Notre Dame Jct. Prov. Pk., 49.116°N, 55.079°W, pitfall trap, conifer forest, 27.VIII.2011, col. L. Pollett (1 ♂, LFC); same data except: 20.VIII.2011 (1 ♂, LFC); 13.IX.2011 (1 ♂, 1 ♀, 1 sex undetermined, LFC). S-W Labrador: 40 km W Churchill Falls, Rt. 500, km 229, 53.373°N, 64.309°W, 12-26.VIII.2001, S. &amp; J. Peck, carrion trap, elevation 550 m, Spruce-moss forest (1 ♀, LFC). Nova Scotia: Cape Breton H.N.P., Lone Shieling, 60 m, PG730860, 15.IX.1984, J.M. Campbell &amp; A. Davies, sifting litter and moss (1 ♂, 3 ♀, CNC); Cape Breton H.N.P., 5 m, S. Ingonish Harbour, PG963674, 12.IX.1984, J.M. Campbell &amp; A. Davies, tread flooded Carex and grasses (1 ♀, CNC); Hants Co., Upper Rawadon, 21.VII.2009, J. Renkema, highbush blueberry field R3T5C (1 sex undetermined, LFC); same data except: 25.VI.2009, highbush blueberry field R2T4A (1 ♀, LFC). Québec: Scotstown, 15.V.2006, 2.X.2006, 9.X.2006, 22.X.2006, 23.X.2006, C. Levesque (4 ♂, 2 ♀, LFC; Mt. Orford Pk., 20.IX.-11.X.1972, Dondale &amp; Redner (1 ♀, CNC); Venice, 45.45°N, 73.08°W, 19.IX.-11.X.1972, Dondale &amp; Redner (1 ♂, CNC). Ontario: Moosonee, 51.24622°N, 80.67281°W, 17-20.VI.2010, NBP field party M1MP111 (1 ♀, LFC). USA: New Hampshire: Coos Co., 8 mi S Gorham Pinkham Notch, 2000 feet, 11.IX.1987, J.M. Campbell &amp; A. Davies, sifting Alnus litter and Sphagnum near pond (1 ♂, CNC).</p><p>Diagnosis .</p><p>This species may be distinguished by the following combination of characters: length 4.2-4.5 mm; body dark reddish-brown, with head dark brown, and legs and at least basal three antennal articles reddish-yellow; integument glossy; pronotum with dense punctation and pubescence; elytra with dense punctation and pubescence with very fine micro-granulation (Fig. 79); head subquadrate, slightly narrower than pronotum, large eyes, each about as long as postocular region in dorsal view (Fig. 79); pronotum subquadrate, widest at apical third (Fig. 79); elytra subparallel, as wide as pronotum and at suture about as long as pronotum (Fig. 79); abdomen subparallel, about as wide as elytra (Fig. 79); Male. Apical margin of tergite VIII with broad, moderate projection in middle three-fifth, with apical margin crenulate (Fig. 81); apical margin of sternite VIII broadly parabolic (Fig. 82); median lobe of aedeagus with tubus short and straight, apical part narrowly rounded in lateral view (Fig. 80). Female. Tergite VIII rounded apically (Fig. 83); apical margin of sternite VIII with broad, shallow median emargination (Fig. 84); spermatheca short, S-shaped, capsule short, club-shaped, stem broad, sinuate, slightly twisted posteriorly (Fig. 85).</p><p>This species may be distinguished from Liogluta castoris, Liogluta pseudocastoris, Liogluta microgranulosa, and Liogluta atriventris by the following combination of characters: antennae, pronotum and elytra reddish-yellow (Fig. 79); pronotum subquadrate (Fig. 79); shape of median lobe of aedeagus different in lateral view (Fig. 80); male tergite VIII with projection crenulate along apical margin (Fig. 81), spermatheca short, S-shaped with broad stem (Fig. 85).</p><p>Natural history.</p><p>Adults were collected in a conifer forest using pitfall traps, in a spruce-moss forest using carrion-baited traps, and in a highbush blueberry field. Others were collected by sifting litter and moss, sifting Alnus litter and Sphagnum moss near a pond, and treading flooded Carex and grasses. The flight period is from May to October.</p><p>Distribution.</p><p>Canada: LB, NF, NS, QC, ON. USA: NH.</p></div>	https://treatment.plazi.org/id/BCCAA8F4D45A6EF52B4934AC960581B3	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Klimaszewski, Jan;Webster, Reginald P.;Langor, David W.;Sikes, Derek;Bourdon, Caroline;Godin, Benoit;Ernst, Crystal	Klimaszewski, Jan, Webster, Reginald P., Langor, David W., Sikes, Derek, Bourdon, Caroline, Godin, Benoit, Ernst, Crystal (2016): A review of Canadian and Alaskan species of the genus Liogluta Thomson, and descriptions of three new species (Coleoptera, Staphylinidae, Aleocharinae). ZooKeys 273: 217-256, DOI: http://dx.doi.org/10.3897/zookeys.573.7878, URL: http://dx.doi.org/10.3897/zookeys.573.7878
6DD0D1D54E08A36EC1325D17F1D7174B.text	6DD0D1D54E08A36EC1325D17F1D7174B.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Liogluta atriventris (Casey 1906) Casey 1906	<div><p>Taxon classification Animalia Coleoptera Staphylinidae</p><p>Liogluta atriventris (Casey, 1906) Figs 86-89</p><p>Athetota atriventris Casey, 1906: 336. As Atheta (Liogluta): Bernhauer and Scheerpeltz 1926: 656; Moore and Legner 1975: 355. Lectotype (male): Canada, Victoria, Vancouver Island; atriventris Casey; Type USNM 39475; H.F. Wickham, Casey Bequest 1925 (USNM). Present designation.</p><p>Diagnosis</p><p>(based on male lectotype). This species may be distinguished by the following combination of characters: small body size, length 2.8 mm; head and abdomen dark brown, pronotum, elytra and legs reddish-yellow (Fig. 86); integument glossy with weak meshy microsculpture; pronotum and elytra with moderately dense punctation and pubescence, elytra with very fine micro-granulation (Fig. 86); head subquadrate, slightly narrower than pronotum; large eyes, each about as long as postocular region in dorsal view (Fig. 86); antennae of the holotype are partially damaged and cannot be completely described, but fifth and sixth articles suggest that missing funicle articles are subquadrate; pronotum slightly transverse, widest near the middle (Fig. 86); elytra wider and slightly longer than pronotum (Fig. 86); abdomen subparallel, about as wide as elytra (Fig. 86). Male. Apical margin of tergite VIII with broad, truncate projection in middle two-thirds bounded laterally by small tooth-like processes, apical margin crenulate (Fig. 88); apical margin of sternite VIII evenly broadly parabolic from base (Fig. 89); median lobe of aedeagus with tubus bent slightly ventrad at middle, apical part relatively broadly rounded in lateral view (Fig. 87). Female. Unknown.</p><p>Liogluta atriventris may be distinguished from the other species of the granulosa group by the following combination of characters: body size small, length 2.8 mm; elytra slightly longer than pronotum (Fig. 86); pronotum glossy with weak microsculpture (Fig. 86); shape of median lobe of aedeagus different in lateral view (Fig. 87), and projection on apical margin of male tergite VIII crenulate, with tooth-like processes laterally (Fig. 88).</p><p>Natural history.</p><p>Unknown.</p><p>Distribution.</p><p>Known only from Vancouver Island, British Columbia.</p><p>Comments.</p><p>This species is known only from one damaged male specimen. More specimens, including females, are needed for study to confirm the status of this species.</p></div>	https://treatment.plazi.org/id/6DD0D1D54E08A36EC1325D17F1D7174B	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Klimaszewski, Jan;Webster, Reginald P.;Langor, David W.;Sikes, Derek;Bourdon, Caroline;Godin, Benoit;Ernst, Crystal	Klimaszewski, Jan, Webster, Reginald P., Langor, David W., Sikes, Derek, Bourdon, Caroline, Godin, Benoit, Ernst, Crystal (2016): A review of Canadian and Alaskan species of the genus Liogluta Thomson, and descriptions of three new species (Coleoptera, Staphylinidae, Aleocharinae). ZooKeys 273: 217-256, DOI: http://dx.doi.org/10.3897/zookeys.573.7878, URL: http://dx.doi.org/10.3897/zookeys.573.7878
971751F84EFBE744D0ABB39CB45E4689.text	971751F84EFBE744D0ABB39CB45E4689.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Liogluta gigantea Klimaszewski & Langor 2011	<div><p>Taxon classification Animalia Coleoptera Staphylinidae</p><p>Liogluta gigantea Klimaszewski &amp; Langor, 2011 Figs 90-96</p><p>Liogluta gigantea Klimaszewski &amp; Langor, in Klimaszewski et al. 2011: 167. Holotype (female): Canada, Newfoundland, Labrador, 75 km SW Goose Bay, Tr. 500, 53° 02.6 N, 61° 16.6 W, 13-26.VIII.2001, S. and J. Peck// Carrion trap, elevation 100 m, spruce-lichen forest, 2001-44 (LFC).</p><p>New locality data.</p><p>CANADA: Québec: 4 mi W Masham, near Mud Lake, 24.X.1967, J.M. Campbell &amp; A. Smetana, Berlese sample ex lining of deserted beaver lodge (1 sex undetermined, CNC); Gatineau Park, near Mud Lake, 24.X.1967, A. Smetana (4 sex undetermined, CNC); Ontario: Rondeau Pr. Pk., Marsh Trail, 4.VI.1985, A. Davies &amp; J.M. Campbell (1 ♂, CNC); Lake Superior Pr. Pk., Sand Riv., 6.VI.1973, J.M. Campbell &amp; R. Parry (1 sex undetermined, CNC).</p><p>Diagnosis.</p><p>This species may be distinguished by: body length 4.2-5.0 mm, robust, broad, dark brown, with pronotum, elytra (except for scutellar region), and legs reddish-brown; forebody moderately glossy, with fine and dense punctation, short pubescence and meshed microsculpture (Fig. 90); head subquadrate, slightly narrower than pronotum, large eyes, each as long as postocular region in dorsal view (Fig. 90); antennae thin, all articles elongate to subquadrate (Fig. 90); pronotum transverse, widest at apical third (Fig. 90); elytra wider than pronotum, at suture as long as or slightly longer than pronotum, with posterior margin almost rectangular (Fig. 90); abdomen broad and flattened (Fig. 90). Male (new description). Apical margin of tergite VIII rounded with broad crenulations and small rounded process at middle (Fig. 92); apical margin of sternite VIII parabolic (Fig. 93); median lobe of aedeagus short and stout, with tubus arched slightly ventrad near apex, apical part narrow in lateral view (Fig. 91). Female. Apical margin of tergite VIII rounded-triangular (Fig. 94); apical margin of sternite VIII broadly truncate, with row of microsetae (Fig. 95); spermatheca vaguely S-shaped, capsule club-shaped, stem sinuate and twisted posteriorly (Fig. 96).</p><p>Distribution.</p><p>Canada: NF, QC, ON.</p><p>Natural history.</p><p>Adults were collected in June, August, and October, in carrion-baited pitfall traps in spruce forests, and from a Berlese funnel extraction of the interior of a deserted beaver lodge.</p></div>	https://treatment.plazi.org/id/971751F84EFBE744D0ABB39CB45E4689	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Pensoft via Plazi	Klimaszewski, Jan;Webster, Reginald P.;Langor, David W.;Sikes, Derek;Bourdon, Caroline;Godin, Benoit;Ernst, Crystal	Klimaszewski, Jan, Webster, Reginald P., Langor, David W., Sikes, Derek, Bourdon, Caroline, Godin, Benoit, Ernst, Crystal (2016): A review of Canadian and Alaskan species of the genus Liogluta Thomson, and descriptions of three new species (Coleoptera, Staphylinidae, Aleocharinae). ZooKeys 273: 217-256, DOI: http://dx.doi.org/10.3897/zookeys.573.7878, URL: http://dx.doi.org/10.3897/zookeys.573.7878
