taxonID	type	description	language	source
8DED010F0388039F833595CD4F29E972.taxon	diagnosis	Diagnosis. Body length ranging from 2.8 - 5.4 mm, body narrowly subparallel (Figs 1, 8, 16, 23, 30, 35, 44, 50, 58, 65, 72, 79, 86, 90), moderately flattened; elytra and abdomen wider than head and pronotum; uniformly dark brown or reddish-brown with head and posterior abdomen dark brown to almost black; integument of forebody with strong meshed microsculpture; surface of elytra often granulose (Figs 50, 51); head with incomplete postocular carinae, postocular region of head long and in most species longer than diameter of eye; glossae split into two lobes; antennae long with articles VI-X subquadrate, slightly transverse or rarely slightly elongate; pronotum with hypomera fully visible in lateral view; pubescence of pronotum directed posteriad on midline of disc and posterolaterad to laterad on sides; mesocoxae narrowly separated, metasternal process short and acute; legs long, three basal articles of metatarsi highly elongate in most species; tarsal formula 5 - 5 - 5. Male. Apical margin of male tergite VIII often with broad, variably-shaped rectangular projection, with edge entire or bearing crenulation or variably shaped structures, with two lateral teeth and sometimes with additional median tooth; integument of disc often with broadly distributed asperate punctation near apex (Figs 3, 11, 18, 25, 38, 46, 53, 60, 67, 74, 81, 88, 92); median lobe of aedeagus simple with apical part variably shaped in lateral view (Figs 2, 9, 17, 24, 36, 45, 52, 59, 66, 73, 80, 87, 91). Female. Spermatheca with capsule narrowly club-shaped or tubular, with apical invagination moderate to deep, stem long, sinuate, looped or twisted posteriorly (Figs 7, 15, 22, 29, 34, 42, 43, 57, 64, 71, 78, 85, 96); sternite VIII with apical margin rounded or medially emarginate, antecostal suture arcuate, or slightly to distinctly sinuate (Figs 6, 14, 21, 28, 33, 41, 49, 56, 63, 70, 77, 84, 89, 95).	en	Klimaszewski, Jan, Webster, Reginald P., Langor, David W., Sikes, Derek, Bourdon, Caroline, Godin, Benoit, Ernst, Crystal (2016): A review of Canadian and Alaskan species of the genus Liogluta Thomson, and descriptions of three new species (Coleoptera, Staphylinidae, Aleocharinae). ZooKeys 273: 217-256, DOI: http://dx.doi.org/10.3897/zookeys.573.7878, URL: http://dx.doi.org/10.3897/zookeys.573.7878
8DED010F0388039F833595CD4F29E972.taxon	distribution	Distribution. The genus is holarctic in distribution (Smetana 2004). Seevers (1978) mentioned a few species from Africa and Jamaica but these records need verification.	en	Klimaszewski, Jan, Webster, Reginald P., Langor, David W., Sikes, Derek, Bourdon, Caroline, Godin, Benoit, Ernst, Crystal (2016): A review of Canadian and Alaskan species of the genus Liogluta Thomson, and descriptions of three new species (Coleoptera, Staphylinidae, Aleocharinae). ZooKeys 273: 217-256, DOI: http://dx.doi.org/10.3897/zookeys.573.7878, URL: http://dx.doi.org/10.3897/zookeys.573.7878
3D5339B5E610D73EBD5C18424D565CBA.taxon	diagnosis	Diagnosis. This species may be distinguished by the following combination of characters: body subparallel, entirely reddish-brown or with head and posterior abdomen chestnut brown (Fig. 1); length 3.9 - 4.5 mm; integument of forebody with meshed microsculpture, moderately glossy; head about one-quarter narrower than maximum width of pronotum; pronotum transverse, narrower at base and widest in apical third; elytra at suture about as long as pronotum; basal three articles of metatarsus elongate, first longest, second about as long as third, fourth shorter than either of preceding articles; apical margin of male tergite VIII with broad, short, truncate projection with rounded lateral angles, apical margin smooth or slightly crenulate (Fig. 3); female tergite VIII with apical margin broadly, just visibly emarginate (Fig. 6); genital structures as illustrated (Figs 2, 7).	en	Klimaszewski, Jan, Webster, Reginald P., Langor, David W., Sikes, Derek, Bourdon, Caroline, Godin, Benoit, Ernst, Crystal (2016): A review of Canadian and Alaskan species of the genus Liogluta Thomson, and descriptions of three new species (Coleoptera, Staphylinidae, Aleocharinae). ZooKeys 273: 217-256, DOI: http://dx.doi.org/10.3897/zookeys.573.7878, URL: http://dx.doi.org/10.3897/zookeys.573.7878
3D5339B5E610D73EBD5C18424D565CBA.taxon	distribution	Distribution. Recorded from LB, NB, NF, NS, ON, QC, AB, YT, and BC (Casey 1906, Lohse et al. 1990, Klimaszewski et al. 2008, Majka and Klimaszewski 2008, Klimaszewski et al. 2011, Webster et al. 2012, Bousquet et al. 2013, Klimaszewski et al. 2015), and newly in USA from MT and NH.	en	Klimaszewski, Jan, Webster, Reginald P., Langor, David W., Sikes, Derek, Bourdon, Caroline, Godin, Benoit, Ernst, Crystal (2016): A review of Canadian and Alaskan species of the genus Liogluta Thomson, and descriptions of three new species (Coleoptera, Staphylinidae, Aleocharinae). ZooKeys 273: 217-256, DOI: http://dx.doi.org/10.3897/zookeys.573.7878, URL: http://dx.doi.org/10.3897/zookeys.573.7878
266DA326B15F0AA4AE8B5868A99D4526.taxon	diagnosis	Diagnosis. This species may be distinguished by the following combination of characters: body subparallel, slender, dark brown to black with pronotum brown and paler than head, elytra yellowish or reddish-brown (Fig. 8); length 3.8 - 4.4 mm; integument of forebody with meshed microsculpture moderately pronounced, surface moderately glossy; head about one-quarter narrower than maximum width of pronotum; pronotum transverse, narrower at base and widest at middle (width of pronotum variable, some specimens have pronotum markedly narrower than base of elytra and some have pronotum nearly as wide as base of elytra); elytra at suture slightly shorter than pronotum; basal three articles of metatarsus elongate, subequal in length and each slightly longer than fourth article; male tergite VIII with apical margin truncate, bordered by two short lateral teeth, variably sculptured and ranging from smooth to crenulate, or denticulate along margin (Fig. 11); genital structures as illustrated (Figs 9 - 15).	en	Klimaszewski, Jan, Webster, Reginald P., Langor, David W., Sikes, Derek, Bourdon, Caroline, Godin, Benoit, Ernst, Crystal (2016): A review of Canadian and Alaskan species of the genus Liogluta Thomson, and descriptions of three new species (Coleoptera, Staphylinidae, Aleocharinae). ZooKeys 273: 217-256, DOI: http://dx.doi.org/10.3897/zookeys.573.7878, URL: http://dx.doi.org/10.3897/zookeys.573.7878
266DA326B15F0AA4AE8B5868A99D4526.taxon	distribution	Distribution. Canada: BC, YT (Klimaszewski et al. 2012). USA: AK (Lohse et al. 1990).	en	Klimaszewski, Jan, Webster, Reginald P., Langor, David W., Sikes, Derek, Bourdon, Caroline, Godin, Benoit, Ernst, Crystal (2016): A review of Canadian and Alaskan species of the genus Liogluta Thomson, and descriptions of three new species (Coleoptera, Staphylinidae, Aleocharinae). ZooKeys 273: 217-256, DOI: http://dx.doi.org/10.3897/zookeys.573.7878, URL: http://dx.doi.org/10.3897/zookeys.573.7878
0ADA3B0D9C850125AC2A4DF09AFF5A37.taxon	diagnosis	Diagnosis. This species may be distinguished by the following combination of characters: body subparallel, slender, bicoloured, pronotum orange and remainder of body dark brown to reddish-brown; length 4.1 - 4.3 mm (Fig. 16); integument of forebody with weak meshed microsculpture, surface highly glossy; head slightly narrower than pronotum; pronotum subquadrate; antennae enlarged and black to brown; elytra about as wide as pronotum and at suture about as long as pronotum; basal two articles of metatarsus distinctly elongate, subequal in length, each longer than third article. New description of male. Apical margin of tergite VIII with broad, obtusely triangular projection in almost middle half with rounded lateral angles (Fig. 18); sternite VIII elongate, rounded apically, with antecostal suture arcuate, well separated from basal margin (Fig. 19); median lobe of aedeagus with tubus short and sinuate, its subapical part narrowly elongate in lateral view, internal sac structures distinct (Fig. 17). Female. Tergite VIII with apical margin obtusely angulate, broadly rounded at middle (Fig. 20); sternite VIII shallowly, broadly emarginated medially (Fig. 21); spermatheca with a short club-shaped capsule and short apical invagination, stem long and highly sinuate (Fig. 22).	en	Klimaszewski, Jan, Webster, Reginald P., Langor, David W., Sikes, Derek, Bourdon, Caroline, Godin, Benoit, Ernst, Crystal (2016): A review of Canadian and Alaskan species of the genus Liogluta Thomson, and descriptions of three new species (Coleoptera, Staphylinidae, Aleocharinae). ZooKeys 273: 217-256, DOI: http://dx.doi.org/10.3897/zookeys.573.7878, URL: http://dx.doi.org/10.3897/zookeys.573.7878
0ADA3B0D9C850125AC2A4DF09AFF5A37.taxon	distribution	Distribution. Vancouver Island, British Columbia (Casey 1894).	en	Klimaszewski, Jan, Webster, Reginald P., Langor, David W., Sikes, Derek, Bourdon, Caroline, Godin, Benoit, Ernst, Crystal (2016): A review of Canadian and Alaskan species of the genus Liogluta Thomson, and descriptions of three new species (Coleoptera, Staphylinidae, Aleocharinae). ZooKeys 273: 217-256, DOI: http://dx.doi.org/10.3897/zookeys.573.7878, URL: http://dx.doi.org/10.3897/zookeys.573.7878
8DEA14C899858ECCEDC53E28C6657B30.taxon	diagnosis	Diagnosis. This species may be distinguished by the following combination of characters: body broadly subparallel (Fig. 23); pronotum, elytra, legs and basal antennal article reddish-brown, head and abdomen chestnut brown (Fig. 23); length 4.0 - 4.2 mm; integument of forebody with meshed microsculpture, moderately glossy; head about one-third narrower than maximum width of pronotum; pronotum more or less evenly arcuate laterally (Fig. 23); elytra at suture about as long as pronotum; basal three articles of metatarsus missing in holotype (Fig. 23). New description of male. Apical margin of tergite VIII with very broad, short, subtruncate projection with rounded lateral angles, with apical margin faintly crenulate (Fig. 25); sternite VIII broadly rounded apically (Fig. 26); median lobe of aedeagus with tubus slightly arcuate ventrally, with apex narrow and rounded (Fig. 24). Female. Tergite VIII truncate apically (Fig. 27); sternite VIII very slightly broadly emarginate apically (Fig. 28); spermatheca with tubular capsule and deep apical invagination, stem thin, long, and highly sinuate (Fig. 29). This species is similar to Liogluta terminalis but has dark brown antennae, head and pronotum (antennae, head, and pronotum are uniformly reddish-brown or only slightly darker than remaining parts of the body in Liogluta terminalis). Spermatheca is differently shaped in each species; Liogluta wickhami has smaller and differently shaped capsule with a deep apical invagination and has a shorter and differently looped posterior stem (Fig. 29). Liogluta wickhami is also very similar to Liogluta vasta but can be distinguished by the shape of pronotum which has evenly arcuate sides and is broadest at middle (Fig. 23), while it is trapezoidal in shape and is broadest in apical third in the latter species (Fig. 30).	en	Klimaszewski, Jan, Webster, Reginald P., Langor, David W., Sikes, Derek, Bourdon, Caroline, Godin, Benoit, Ernst, Crystal (2016): A review of Canadian and Alaskan species of the genus Liogluta Thomson, and descriptions of three new species (Coleoptera, Staphylinidae, Aleocharinae). ZooKeys 273: 217-256, DOI: http://dx.doi.org/10.3897/zookeys.573.7878, URL: http://dx.doi.org/10.3897/zookeys.573.7878
8DEA14C899858ECCEDC53E28C6657B30.taxon	distribution	Distribution. The female holotype was captured in the Stickeen River Valley of British Columbia (Casey 1894), and one male was found in Three Guardsmen Pass, British Columbia.	en	Klimaszewski, Jan, Webster, Reginald P., Langor, David W., Sikes, Derek, Bourdon, Caroline, Godin, Benoit, Ernst, Crystal (2016): A review of Canadian and Alaskan species of the genus Liogluta Thomson, and descriptions of three new species (Coleoptera, Staphylinidae, Aleocharinae). ZooKeys 273: 217-256, DOI: http://dx.doi.org/10.3897/zookeys.573.7878, URL: http://dx.doi.org/10.3897/zookeys.573.7878
311DED817411DB63D88AB88042395100.taxon	diagnosis	Diagnosis (based on female lectotype, male unknown). This species is very similar to Liogluta wickhami, but in Liogluta vasta the forebody is less reddish, and the pronotum is more trapezoidal and narrowest at base (Fig. 30); the apical margin of tergite VIII is broadly arcuate with the antecostal suture very narrowly separated from the basal margin (Fig. 32); the apical margin of sternite VIII is slightly, very broadly emarginate with the antecostal suture highly sinuate and well separated from the basal margin (Fig. 33). The female lectotype is missing the spermatheca. The spermatheca of the Yukon specimen in CNC cited by Lohse and Smetana (1985), tentatively identified as belonging to this species, is illustrated in Fig. 34 (after Lohse and Smetana 1985). Males and more females from the type locality are needed to clearly define this species, which is here tentatively listed as a valid species. When more specimens of Liogluta vasta become available for study and the morphological variation is known, we will be able to understand the relationship of this species to other nearctic Liogluta species. Liogluta vasta is also similar to some specimens (with broad pronotum) of Liogluta trapezicollis. A DNA comparison between Liogluta vasta and other Liogluta species is needed to clarify its identity and relationships.	en	Klimaszewski, Jan, Webster, Reginald P., Langor, David W., Sikes, Derek, Bourdon, Caroline, Godin, Benoit, Ernst, Crystal (2016): A review of Canadian and Alaskan species of the genus Liogluta Thomson, and descriptions of three new species (Coleoptera, Staphylinidae, Aleocharinae). ZooKeys 273: 217-256, DOI: http://dx.doi.org/10.3897/zookeys.573.7878, URL: http://dx.doi.org/10.3897/zookeys.573.7878
311DED817411DB63D88AB88042395100.taxon	distribution	Distribution. Canada: YT?. USA: AK.	en	Klimaszewski, Jan, Webster, Reginald P., Langor, David W., Sikes, Derek, Bourdon, Caroline, Godin, Benoit, Ernst, Crystal (2016): A review of Canadian and Alaskan species of the genus Liogluta Thomson, and descriptions of three new species (Coleoptera, Staphylinidae, Aleocharinae). ZooKeys 273: 217-256, DOI: http://dx.doi.org/10.3897/zookeys.573.7878, URL: http://dx.doi.org/10.3897/zookeys.573.7878
311DED817411DB63D88AB88042395100.taxon	discussion	Discussion. The original type material of Homalota vasta Mӓklin, 1853 (ZMH) consists of two female specimens representing two different species in two genera, Atheta (as Boreophilia in Lohse and Smetana 1985) and Liogluta. Lohse and Smetana (1985) designated the female specimen belonging to Liogluta as the lectotype of Homalota vasta Mӓklin. However, the label data published by Lohse and Smetana (1985) for the Liogluta specimen corresponded to the Atheta specimen (see also discussion in Gusarov 2003 b, who also mislabelled the specimens). We consider Lohse and Smetana's lectotype designation as valid regardless of the obvious mistake of publishing the wrong label data; therefore, the name vasta is affiliated with Liogluta. The female paralectotype belongs to Atheta keeni Casey, 1910. It is noteworthy that despite years of intensive collections made primarily between 2008 - 2013 in southeast Alaskan lowland forests and alpine zones, including in and around Sitka, which have resulted in 22,029 specimens of Staphylinidae (http: // arctos. database. museum / saved / SE-AK-Staphylinidae), no specimens of Liogluta vasta were found.	en	Klimaszewski, Jan, Webster, Reginald P., Langor, David W., Sikes, Derek, Bourdon, Caroline, Godin, Benoit, Ernst, Crystal (2016): A review of Canadian and Alaskan species of the genus Liogluta Thomson, and descriptions of three new species (Coleoptera, Staphylinidae, Aleocharinae). ZooKeys 273: 217-256, DOI: http://dx.doi.org/10.3897/zookeys.573.7878, URL: http://dx.doi.org/10.3897/zookeys.573.7878
5831279A239410B9DEEB5CE923955057.taxon	diagnosis	Diagnosis. This species may be distinguished by the following combination of characters: body elongate with elytra and abdomen broad, moderately robust, dark brown to black with tarsi and tibiae often reddish-brown, elytra sometimes with reddish tinge (Figs 35); length 3.5 - 4.0 mm; integument of forebody with moderately pronounced meshed microsculpture, surface highly glossy; head about one-eighth narrower than maximum width of pronotum (Fig. 35); pronotum transverse, about evenly wide in posterior half, then distinctly narrowing apicad, forming round apical angles (Fig. 35); elytra at suture slightly longer than pronotum (Fig. 35); basal four articles of metatarsus about the same length, each shorter than fifth article. Male. Tergite VIII with apical margin broadly arcuate, without teeth or crenulations (Fig. 38); sternite VIII rounded apically (Fig. 39); median lobe of aedeagus with tubus broadly arcuate ventrally and with apex narrow and pointed in lateral view (Fig. 36); tubus broad and triangular apically in dorsal view (Fig. 37). Female. Tergite VIII broadly arcuate apically (Fig. 40); sternite VIII evenly rounded apically, with antecostal suture distinctly sinuate (Fig. 41); spermatheca with spherical capsule with invagination deep and narrow, stem S-shaped, gradually becoming very narrow posteriad (Figs 42, 43).	en	Klimaszewski, Jan, Webster, Reginald P., Langor, David W., Sikes, Derek, Bourdon, Caroline, Godin, Benoit, Ernst, Crystal (2016): A review of Canadian and Alaskan species of the genus Liogluta Thomson, and descriptions of three new species (Coleoptera, Staphylinidae, Aleocharinae). ZooKeys 273: 217-256, DOI: http://dx.doi.org/10.3897/zookeys.573.7878, URL: http://dx.doi.org/10.3897/zookeys.573.7878
5831279A239410B9DEEB5CE923955057.taxon	distribution	Distribution. Canada: LB, NF, NT, NU, QC, YT. USA: NH (Lohse et al. 1990; Klimaszewski et al. 2012; Bousquet et al. 2013).	en	Klimaszewski, Jan, Webster, Reginald P., Langor, David W., Sikes, Derek, Bourdon, Caroline, Godin, Benoit, Ernst, Crystal (2016): A review of Canadian and Alaskan species of the genus Liogluta Thomson, and descriptions of three new species (Coleoptera, Staphylinidae, Aleocharinae). ZooKeys 273: 217-256, DOI: http://dx.doi.org/10.3897/zookeys.573.7878, URL: http://dx.doi.org/10.3897/zookeys.573.7878
138BA9E24E5EB5220EB7FBBE7401E816.taxon	diagnosis	Diagnosis. This species may be distinguished by the following combination of characters: body narrowly elongate, robust, dark-brown to black with elytra, tarsi and tibiae often yellowish- or reddish-brown (Fig. 44); length 2.8 - 3.3 mm; integument of forebody with moderately pronounced meshed microsculpture, surface highly glossy (Fig. 44); head about one-eighth narrower than maximum width of pronotum (Fig. 44); pronotum transverse, about evenly wide in basal half and then distinctly narrowing anteriad (Fig. 44); elytra at suture about as long as pronotum (Fig. 44); basal four articles of metatarsus about same length, each shorter than fifth article. Male. Tergite VIII with short subrectangular projection on more than half width of apical margin, with rounded lateral angles, apical margin smooth or micro-crenulate (Fig. 46); sternite VIII parabolically rounded apically (Fig. 47); median lobe of aedeagus with tubus almost straight in lateral view, with apex moderately narrow, rounded (Fig. 45). Female. Tergite VIII truncate apically (Fig. 48); sternite VIII with apical margin evenly rounded, antecostal suture slightly sinuate (Fig. 49). Spermatheca unknown.	en	Klimaszewski, Jan, Webster, Reginald P., Langor, David W., Sikes, Derek, Bourdon, Caroline, Godin, Benoit, Ernst, Crystal (2016): A review of Canadian and Alaskan species of the genus Liogluta Thomson, and descriptions of three new species (Coleoptera, Staphylinidae, Aleocharinae). ZooKeys 273: 217-256, DOI: http://dx.doi.org/10.3897/zookeys.573.7878, URL: http://dx.doi.org/10.3897/zookeys.573.7878
138BA9E24E5EB5220EB7FBBE7401E816.taxon	distribution	Distribution. Canada: AB, BC, YT. USA: AK, OR, WA (Mӓklin 1852, Bernhauer 1907, Hatch 1957, Moore and Legner 1975, Lohse and Smetana 1985, Klimaszewski and Winchester 2002).	en	Klimaszewski, Jan, Webster, Reginald P., Langor, David W., Sikes, Derek, Bourdon, Caroline, Godin, Benoit, Ernst, Crystal (2016): A review of Canadian and Alaskan species of the genus Liogluta Thomson, and descriptions of three new species (Coleoptera, Staphylinidae, Aleocharinae). ZooKeys 273: 217-256, DOI: http://dx.doi.org/10.3897/zookeys.573.7878, URL: http://dx.doi.org/10.3897/zookeys.573.7878
C35069FEB50F14003F74EE37EA1D0CBF.taxon	diagnosis	Diagnosis. This species may be distinguished by the following combination of characters: body broadly subparallel, dark brown, with elytra, tarsi and tibiae often reddish-brown (Fig. 50) (one specimen from northern Yukon was entirely black); length 2.8 - 3.3 mm; integument of forebody with moderately pronounced meshed microsculpture; head about one-eighth narrower than maximum width of pronotum (Fig. 50); pronotum transverse, about evenly wide in basal one-third of its length, then strongly broadest at apical one-third and gradually narrowed apically (Fig. 50); elytra at suture about as long as pronotum, its surface coarsely granulose (Fig. 50); basal two articles of metatarsus about the same length, each shorter than fifth article. Male. Apical margin of tergite VIII with short, very obtusely angular projection in medial two-thirds with rounded lateral angles, margin of projection smooth or micro-denticulate (Fig. 53); apical margin of sternite VIII broadly parabolic (Fig. 54); median lobe of aedeagus with tubus broadly arched, bent ventrad, apex narrow and rounded (Fig. 52). Female. Apical margin of tergite VIII truncate in middle one-third (Fig. 55); apical margin of sternite VIII arcuate, antecostal suture distinctly sinuate (Fig. 56); spermatheca highly sinuate as illustrated (Fig. 57).	en	Klimaszewski, Jan, Webster, Reginald P., Langor, David W., Sikes, Derek, Bourdon, Caroline, Godin, Benoit, Ernst, Crystal (2016): A review of Canadian and Alaskan species of the genus Liogluta Thomson, and descriptions of three new species (Coleoptera, Staphylinidae, Aleocharinae). ZooKeys 273: 217-256, DOI: http://dx.doi.org/10.3897/zookeys.573.7878, URL: http://dx.doi.org/10.3897/zookeys.573.7878
C35069FEB50F14003F74EE37EA1D0CBF.taxon	distribution	Distribution. Canada: YT. USA: AK (Lohse et al. 1990, Klimaszewski et al. 2008, Klimaszewski et al. 2012).	en	Klimaszewski, Jan, Webster, Reginald P., Langor, David W., Sikes, Derek, Bourdon, Caroline, Godin, Benoit, Ernst, Crystal (2016): A review of Canadian and Alaskan species of the genus Liogluta Thomson, and descriptions of three new species (Coleoptera, Staphylinidae, Aleocharinae). ZooKeys 273: 217-256, DOI: http://dx.doi.org/10.3897/zookeys.573.7878, URL: http://dx.doi.org/10.3897/zookeys.573.7878
433C71A6DB31979836398DDE00E2E691.taxon	etymology	Etymology. Microgranulosa is a Latin adjective meaning microgranulate, in reference to the minute sculpture on the elytra of this species.	en	Klimaszewski, Jan, Webster, Reginald P., Langor, David W., Sikes, Derek, Bourdon, Caroline, Godin, Benoit, Ernst, Crystal (2016): A review of Canadian and Alaskan species of the genus Liogluta Thomson, and descriptions of three new species (Coleoptera, Staphylinidae, Aleocharinae). ZooKeys 273: 217-256, DOI: http://dx.doi.org/10.3897/zookeys.573.7878, URL: http://dx.doi.org/10.3897/zookeys.573.7878
433C71A6DB31979836398DDE00E2E691.taxon	description	Description. This species may be distinguished by the following combination of characters: body narrowly subparallel; head, apical articles of antennae, and posterior part of abdomen black, elytra brownish and mottled with black, remaining parts reddish-brown (Fig. 58); length 4.6 - 5.1 mm; integument of forebody with moderately pronounced meshed microsculpture, surface moderately glossy (Fig. 58); head about one-quarter narrower than maximum width of pronotum (Fig. 58); pronotum transverse, about evenly wide in basal half of its length, then strongly narrowed apically (Fig. 58); elytra at suture about as long as pronotum, surface finely and densely microgranulose; basal three articles of metatarsus about equally elongate, each longer than fourth article. Male. Apical margin of tergite VIII with very broad, very obtusely angular projection, with obtuse lateral angles and small tooth medially, margin often micro-crenulate (Fig. 60); sternite VIII rounded apically (Fig. 61); median lobe of aedeagus with tubus distinctly arched ventrad in apical half, apical part narrow (Fig. 59). Female. Tergite VIII with apical margin obtusely angulate (Fig. 62); sternite VIII with apical margin slightly emarginate medially (Fig. 63); spermatheca with stem long, sinuate, spiral posteriorly, capsule club-shaped with apical invagination deep and narrow (Fig. 64).	en	Klimaszewski, Jan, Webster, Reginald P., Langor, David W., Sikes, Derek, Bourdon, Caroline, Godin, Benoit, Ernst, Crystal (2016): A review of Canadian and Alaskan species of the genus Liogluta Thomson, and descriptions of three new species (Coleoptera, Staphylinidae, Aleocharinae). ZooKeys 273: 217-256, DOI: http://dx.doi.org/10.3897/zookeys.573.7878, URL: http://dx.doi.org/10.3897/zookeys.573.7878
433C71A6DB31979836398DDE00E2E691.taxon	distribution	Distribution. Canada: Known only from New Brunswick, Canada.	en	Klimaszewski, Jan, Webster, Reginald P., Langor, David W., Sikes, Derek, Bourdon, Caroline, Godin, Benoit, Ernst, Crystal (2016): A review of Canadian and Alaskan species of the genus Liogluta Thomson, and descriptions of three new species (Coleoptera, Staphylinidae, Aleocharinae). ZooKeys 273: 217-256, DOI: http://dx.doi.org/10.3897/zookeys.573.7878, URL: http://dx.doi.org/10.3897/zookeys.573.7878
C5ADE7F59540220AD9B86B749FFCC754.taxon	etymology	Etymology. Castoris is a Latin adjective derived from the name of the American beaver (Castor canadensis Kuhl), in reference to beaver dams where some of the type specimens were captured.	en	Klimaszewski, Jan, Webster, Reginald P., Langor, David W., Sikes, Derek, Bourdon, Caroline, Godin, Benoit, Ernst, Crystal (2016): A review of Canadian and Alaskan species of the genus Liogluta Thomson, and descriptions of three new species (Coleoptera, Staphylinidae, Aleocharinae). ZooKeys 273: 217-256, DOI: http://dx.doi.org/10.3897/zookeys.573.7878, URL: http://dx.doi.org/10.3897/zookeys.573.7878
C5ADE7F59540220AD9B86B749FFCC754.taxon	description	Description. Body length 4.6 - 5.4 mm, subparallel (Fig. 65); head and at least apical part of abdomen dark brown with pronotum, elytra, basal articles of antennae and legs yellowish to reddish-brown; integument moderately glossy, more so on posterior abdomen; forebody with minute and sparse punctation and sparse pubescence (Fig. 65); elytra with minute micro-granulation; head rounded and narrowed posteriorly, with large eyes, each about as long as postocular area in dorsal view (Fig. 65); antennae with articles V-X subquadrate to slightly elongate (Fig. 65); pronotum slightly transverse, broadly rounded laterally, slightly wider than head and narrower than elytra, pubescence directed latero-posteriad from midline of disc (Fig. 65); elytra transverse, at suture as long as pronotum, slightly longer laterally, with pubescence directed posteriad (Fig. 65); abdomen subparallel for most of its length, about as wide as elytra (Fig. 65). Male. Aedeagus with bulbus narrowly oval, median lobe with apical half of tubus slightly arched ventrad, apical part moderately broad in lateral view (Fig. 66); internal sac with few pronounced structures / membrane folds (Fig. 66); apical margin of tergite VIII with very broad truncate projection with obtuse lateral angles, with margin smooth or minutely crenulate (Fig. 67); apical margin of sternite VIII rounded (Fig. 68). Female. Tergite VIII with apical margin broadly rounded (Fig. 69); sternite VIII scarcely emarginate apically, antecostal suture distinctly sinuate, well separated from basal margin (Fig. 70); spermatheca with stem long, sinuate, twisted posteriorly, capsule tubular, with apical invagination narrow, short (Fig. 71).	en	Klimaszewski, Jan, Webster, Reginald P., Langor, David W., Sikes, Derek, Bourdon, Caroline, Godin, Benoit, Ernst, Crystal (2016): A review of Canadian and Alaskan species of the genus Liogluta Thomson, and descriptions of three new species (Coleoptera, Staphylinidae, Aleocharinae). ZooKeys 273: 217-256, DOI: http://dx.doi.org/10.3897/zookeys.573.7878, URL: http://dx.doi.org/10.3897/zookeys.573.7878
C5ADE7F59540220AD9B86B749FFCC754.taxon	distribution	Distribution. Canada: Known from NB, NS, QC.	en	Klimaszewski, Jan, Webster, Reginald P., Langor, David W., Sikes, Derek, Bourdon, Caroline, Godin, Benoit, Ernst, Crystal (2016): A review of Canadian and Alaskan species of the genus Liogluta Thomson, and descriptions of three new species (Coleoptera, Staphylinidae, Aleocharinae). ZooKeys 273: 217-256, DOI: http://dx.doi.org/10.3897/zookeys.573.7878, URL: http://dx.doi.org/10.3897/zookeys.573.7878
92BCA175D3BA8F17C2C06CA3377214AB.taxon	etymology	Etymology. Pseudocastoris is the Latin prefix pseudo-, false, added to the species name castoris, reflecting the close similarity of the two species.	en	Klimaszewski, Jan, Webster, Reginald P., Langor, David W., Sikes, Derek, Bourdon, Caroline, Godin, Benoit, Ernst, Crystal (2016): A review of Canadian and Alaskan species of the genus Liogluta Thomson, and descriptions of three new species (Coleoptera, Staphylinidae, Aleocharinae). ZooKeys 273: 217-256, DOI: http://dx.doi.org/10.3897/zookeys.573.7878, URL: http://dx.doi.org/10.3897/zookeys.573.7878
92BCA175D3BA8F17C2C06CA3377214AB.taxon	description	Description. Body length 3.9 - 4.4 mm, subparallel; dark brown with irregularly shaped lighter areas on pronotum in some individuals, head and abdomen dark brown, antennae dark, and legs yellowish; integument moderately glossy, more so on posterior portion of abdomen; forebody with minute and sparse punctation and sparse pubescence (Fig. 72); elytra with micro-granulation (Fig. 72); head rounded and narrowed posteriorly, eyes large, each about as long as postocular area in dorsal view (Fig. 72); antennae with articles V-X subquadrate to slightly transverse (Fig. 72); pronotum transverse, broadly rounded laterally, slightly wider than head and narrower than elytra, pubescence directed latero-posteriad from midline of disc (Fig. 72); elytra transverse, at suture about as long as pronotum, slightly longer laterally, with pubescence directed posteriad; abdomen subparallel for most of its length, about as wide as elytra (Fig. 72). Male. Tergite VIII broadly rounded apically, margin smooth (Fig. 74); apical margin of sternite VIII broadly parabolic (Fig. 75); median lobe of aedeagus with bulbus narrowly oval, tubus almost straight with apical part narrowly rounded in lateral view (Fig. 73); internal sac without distinct sclerites but with some vaguely-shaped structures (Fig. 73). Female. Tergite VIII broadly rounded apically (Fig. 76); sternite VIII a little less broadly rounded apically, antecostal suture slightly sinuate, moderately separated from basal margin (Fig. 77); spermatheca with capsule club-shaped, [invagination not perceptible], stem sinuate, about equally narrow throughout with only posterior part enlarged but not twisted (Fig. 78).	en	Klimaszewski, Jan, Webster, Reginald P., Langor, David W., Sikes, Derek, Bourdon, Caroline, Godin, Benoit, Ernst, Crystal (2016): A review of Canadian and Alaskan species of the genus Liogluta Thomson, and descriptions of three new species (Coleoptera, Staphylinidae, Aleocharinae). ZooKeys 273: 217-256, DOI: http://dx.doi.org/10.3897/zookeys.573.7878, URL: http://dx.doi.org/10.3897/zookeys.573.7878
92BCA175D3BA8F17C2C06CA3377214AB.taxon	distribution	Distribution. Known only from New Brunswick, Canada.	en	Klimaszewski, Jan, Webster, Reginald P., Langor, David W., Sikes, Derek, Bourdon, Caroline, Godin, Benoit, Ernst, Crystal (2016): A review of Canadian and Alaskan species of the genus Liogluta Thomson, and descriptions of three new species (Coleoptera, Staphylinidae, Aleocharinae). ZooKeys 273: 217-256, DOI: http://dx.doi.org/10.3897/zookeys.573.7878, URL: http://dx.doi.org/10.3897/zookeys.573.7878
BCCAA8F4D45A6EF52B4934AC960581B3.taxon	diagnosis	Diagnosis. This species may be distinguished by the following combination of characters: length 4.2 - 4.5 mm; body dark reddish-brown, with head dark brown, and legs and at least basal three antennal articles reddish-yellow; integument glossy; pronotum with dense punctation and pubescence; elytra with dense punctation and pubescence with very fine micro-granulation (Fig. 79); head subquadrate, slightly narrower than pronotum, large eyes, each about as long as postocular region in dorsal view (Fig. 79); pronotum subquadrate, widest at apical third (Fig. 79); elytra subparallel, as wide as pronotum and at suture about as long as pronotum (Fig. 79); abdomen subparallel, about as wide as elytra (Fig. 79); Male. Apical margin of tergite VIII with broad, moderate projection in middle three-fifth, with apical margin crenulate (Fig. 81); apical margin of sternite VIII broadly parabolic (Fig. 82); median lobe of aedeagus with tubus short and straight, apical part narrowly rounded in lateral view (Fig. 80). Female. Tergite VIII rounded apically (Fig. 83); apical margin of sternite VIII with broad, shallow median emargination (Fig. 84); spermatheca short, S-shaped, capsule short, club-shaped, stem broad, sinuate, slightly twisted posteriorly (Fig. 85). This species may be distinguished from Liogluta castoris, Liogluta pseudocastoris, Liogluta microgranulosa, and Liogluta atriventris by the following combination of characters: antennae, pronotum and elytra reddish-yellow (Fig. 79); pronotum subquadrate (Fig. 79); shape of median lobe of aedeagus different in lateral view (Fig. 80); male tergite VIII with projection crenulate along apical margin (Fig. 81), spermatheca short, S-shaped with broad stem (Fig. 85).	en	Klimaszewski, Jan, Webster, Reginald P., Langor, David W., Sikes, Derek, Bourdon, Caroline, Godin, Benoit, Ernst, Crystal (2016): A review of Canadian and Alaskan species of the genus Liogluta Thomson, and descriptions of three new species (Coleoptera, Staphylinidae, Aleocharinae). ZooKeys 273: 217-256, DOI: http://dx.doi.org/10.3897/zookeys.573.7878, URL: http://dx.doi.org/10.3897/zookeys.573.7878
BCCAA8F4D45A6EF52B4934AC960581B3.taxon	distribution	Distribution. Canada: LB, NF, NS, QC, ON. USA: NH.	en	Klimaszewski, Jan, Webster, Reginald P., Langor, David W., Sikes, Derek, Bourdon, Caroline, Godin, Benoit, Ernst, Crystal (2016): A review of Canadian and Alaskan species of the genus Liogluta Thomson, and descriptions of three new species (Coleoptera, Staphylinidae, Aleocharinae). ZooKeys 273: 217-256, DOI: http://dx.doi.org/10.3897/zookeys.573.7878, URL: http://dx.doi.org/10.3897/zookeys.573.7878
6DD0D1D54E08A36EC1325D17F1D7174B.taxon	diagnosis	Diagnosis (based on male lectotype). This species may be distinguished by the following combination of characters: small body size, length 2.8 mm; head and abdomen dark brown, pronotum, elytra and legs reddish-yellow (Fig. 86); integument glossy with weak meshy microsculpture; pronotum and elytra with moderately dense punctation and pubescence, elytra with very fine micro-granulation (Fig. 86); head subquadrate, slightly narrower than pronotum; large eyes, each about as long as postocular region in dorsal view (Fig. 86); antennae of the holotype are partially damaged and cannot be completely described, but fifth and sixth articles suggest that missing funicle articles are subquadrate; pronotum slightly transverse, widest near the middle (Fig. 86); elytra wider and slightly longer than pronotum (Fig. 86); abdomen subparallel, about as wide as elytra (Fig. 86). Male. Apical margin of tergite VIII with broad, truncate projection in middle two-thirds bounded laterally by small tooth-like processes, apical margin crenulate (Fig. 88); apical margin of sternite VIII evenly broadly parabolic from base (Fig. 89); median lobe of aedeagus with tubus bent slightly ventrad at middle, apical part relatively broadly rounded in lateral view (Fig. 87). Female. Unknown. Liogluta atriventris may be distinguished from the other species of the granulosa group by the following combination of characters: body size small, length 2.8 mm; elytra slightly longer than pronotum (Fig. 86); pronotum glossy with weak microsculpture (Fig. 86); shape of median lobe of aedeagus different in lateral view (Fig. 87), and projection on apical margin of male tergite VIII crenulate, with tooth-like processes laterally (Fig. 88).	en	Klimaszewski, Jan, Webster, Reginald P., Langor, David W., Sikes, Derek, Bourdon, Caroline, Godin, Benoit, Ernst, Crystal (2016): A review of Canadian and Alaskan species of the genus Liogluta Thomson, and descriptions of three new species (Coleoptera, Staphylinidae, Aleocharinae). ZooKeys 273: 217-256, DOI: http://dx.doi.org/10.3897/zookeys.573.7878, URL: http://dx.doi.org/10.3897/zookeys.573.7878
6DD0D1D54E08A36EC1325D17F1D7174B.taxon	distribution	Distribution. Known only from Vancouver Island, British Columbia.	en	Klimaszewski, Jan, Webster, Reginald P., Langor, David W., Sikes, Derek, Bourdon, Caroline, Godin, Benoit, Ernst, Crystal (2016): A review of Canadian and Alaskan species of the genus Liogluta Thomson, and descriptions of three new species (Coleoptera, Staphylinidae, Aleocharinae). ZooKeys 273: 217-256, DOI: http://dx.doi.org/10.3897/zookeys.573.7878, URL: http://dx.doi.org/10.3897/zookeys.573.7878
971751F84EFBE744D0ABB39CB45E4689.taxon	diagnosis	Diagnosis. This species may be distinguished by: body length 4.2 - 5.0 mm, robust, broad, dark brown, with pronotum, elytra (except for scutellar region), and legs reddish-brown; forebody moderately glossy, with fine and dense punctation, short pubescence and meshed microsculpture (Fig. 90); head subquadrate, slightly narrower than pronotum, large eyes, each as long as postocular region in dorsal view (Fig. 90); antennae thin, all articles elongate to subquadrate (Fig. 90); pronotum transverse, widest at apical third (Fig. 90); elytra wider than pronotum, at suture as long as or slightly longer than pronotum, with posterior margin almost rectangular (Fig. 90); abdomen broad and flattened (Fig. 90). Male (new description). Apical margin of tergite VIII rounded with broad crenulations and small rounded process at middle (Fig. 92); apical margin of sternite VIII parabolic (Fig. 93); median lobe of aedeagus short and stout, with tubus arched slightly ventrad near apex, apical part narrow in lateral view (Fig. 91). Female. Apical margin of tergite VIII rounded-triangular (Fig. 94); apical margin of sternite VIII broadly truncate, with row of microsetae (Fig. 95); spermatheca vaguely S-shaped, capsule club-shaped, stem sinuate and twisted posteriorly (Fig. 96).	en	Klimaszewski, Jan, Webster, Reginald P., Langor, David W., Sikes, Derek, Bourdon, Caroline, Godin, Benoit, Ernst, Crystal (2016): A review of Canadian and Alaskan species of the genus Liogluta Thomson, and descriptions of three new species (Coleoptera, Staphylinidae, Aleocharinae). ZooKeys 273: 217-256, DOI: http://dx.doi.org/10.3897/zookeys.573.7878, URL: http://dx.doi.org/10.3897/zookeys.573.7878
971751F84EFBE744D0ABB39CB45E4689.taxon	distribution	Distribution. Canada: NF, QC, ON.	en	Klimaszewski, Jan, Webster, Reginald P., Langor, David W., Sikes, Derek, Bourdon, Caroline, Godin, Benoit, Ernst, Crystal (2016): A review of Canadian and Alaskan species of the genus Liogluta Thomson, and descriptions of three new species (Coleoptera, Staphylinidae, Aleocharinae). ZooKeys 273: 217-256, DOI: http://dx.doi.org/10.3897/zookeys.573.7878, URL: http://dx.doi.org/10.3897/zookeys.573.7878
