taxonID	type	description	language	source
03C587B7FF85697AC5B9C250FB9AFC34.taxon	materials_examined	Type species: Kumatoeides aulacia sp. nov. The new genus currently includes nine species of small sized (2.23 – 3.20 mm), winged species, glabrous on dorsum and of orange to brownish coloration, with or most typically without greenish metallic luster, and antennae, legs and mouth appendages uniformly pale. All the species have bulging, nearly entire eyes, and very short genae, with slender antennae, reaching slightly beyond middle of elytra in males and shorter in females, with pedicel short and slightly globose in males, shorter than third antennomere, fifth antennomere longest of basal six antennomeres, and antennomeres 7 – 11 feebly incrassate, with seventh longest of the antennal club (except for eleventh antennomere). Punctation generally strong and dense (K. aulacia group) or moderate and sparser (K. tarsalis group) on frons and pronotum. Pronotum transverse, anterior border of hypomera straight, and elytra as wide or imperceptibly wider at base than base of pronotum, with humeri marked, and generally with weak transverse depression behind humeri; elytra striate-punctate, with eight regular striae in their mid section, an additional subhumeral, curved stria connected to margin of elytron before middle, and short scutellar striae (missing in K. anomala sp. nov.). Tibiae dilated at apex and mesotibiae slightly shorter than mesofemora, feebly curved, with first pro- and mesotarsomeres of males strongly dilated and all claws appendiculate. Pygidium with median longitudinal groove, as an important trait to place the new genus among the Eumolpini (e. g., Gómez-Zurita et al. 2005). The genus is tentatively subdivided in two species groups, as named above, based on the characters used in the first couplet of the dichotomic key.	en	Gómez-Zurita, Jesús (2018): Description of Kumatoeides gen. nov. (Coleoptera: Chrysomelidae, Eumolpinae) from New Caledonia. Zootaxa 4521 (1): 89-115, DOI: 10.11646/zootaxa.4521.1.4
03C587B7FF85697AC5B9C250FB9AFC34.taxon	etymology	Derivatio nominis: The genus name is the Latin transliteration of the Greek adjective κυµατoειδής (fem.), meaning corrugated, i. e. surface bent into alternate folds and ridges, in reference to the regular striae on the elytra of these animals, an infrequent trait in Eumolpini, and particularly the New Caledonian Eumolpini.	en	Gómez-Zurita, Jesús (2018): Description of Kumatoeides gen. nov. (Coleoptera: Chrysomelidae, Eumolpinae) from New Caledonia. Zootaxa 4521 (1): 89-115, DOI: 10.11646/zootaxa.4521.1.4
03C587B7FF85697AC5B9C250FB9AFC34.taxon	distribution	Distribution: All known species of Kumatoeides gen. nov. have ranges in Grande Terre within the broad terrestrial ecoregion defined as New Caledonia rain forests in Olson et al. (2001). No species is known for the time being from the New Caledonia dry forests ecoregion or from the Loyalty Islands.	en	Gómez-Zurita, Jesús (2018): Description of Kumatoeides gen. nov. (Coleoptera: Chrysomelidae, Eumolpinae) from New Caledonia. Zootaxa 4521 (1): 89-115, DOI: 10.11646/zootaxa.4521.1.4
03C587B7FF83697CC5B9C5FAFCB3FC11.taxon	description	(Figs 1 f, 2 a, 3 a, 4 a, 5 e)	en	Gómez-Zurita, Jesús (2018): Description of Kumatoeides gen. nov. (Coleoptera: Chrysomelidae, Eumolpinae) from New Caledonia. Zootaxa 4521 (1): 89-115, DOI: 10.11646/zootaxa.4521.1.4
03C587B7FF83697CC5B9C5FAFCB3FC11.taxon	materials_examined	Holotype: Male, New Caledonia (N), Mt. Mandjélia, rainforest, - 20.40439 164.51798, 550 – 600 m, 19. xi. 2008, beating, leg. M. Wanat, HOLOTYPE Kumatoeides anomala sp. nov. J. Gómez-Zurita det. [red label] (MNHN). Paratype: Female, New Caledonia (N), - 20.40439 164.51798, 550 – 600 m, Mt Mandjélia, beating, rainforest, 19. xi. 2008, leg. M. Wanat PARATYPE Kumatoeides anomala sp. nov. J. Gómez-Zurita det. [red label] (MNHW). Other material examined: (1) one female, New Caledonia (N), - 20.40007 164.52788, Mt Mandjélia, 700 – 780 m, montane rainforest, 20. xi. 2008, leg. M. Wanat, Kumatoeides anomala m. J. Gómez-Zurita det. 2018 (MNHW); (2) two females, New Caledonia (N), - 20.40007 164.52788, Mt Mandjélia, 700 – 780 m, montane rainforest, 20. xi. 2008, leg. M. Wanat, Kumatoeides anomala m. J. Gómez-Zurita det. 2018 (MNHW); (3) one male, New Caledonia (N), 21 ° 09 ' S 165 ° 19 ' E, Aoupinié, 420 – 530 m, road to sawmill, 7. ii. 2004, leg. M. Wanat, Kumatoeides anomala m. J. Gómez-Zurita det. 2018 (MNHW). Head, pronotum, scutellum, ventral thoracic parts and trochanters dark brown, with slight bronze metallic luster on head, pronotum and scutellum; elytra hazelnut brown with suture, punctures and inner border of epipleura dark brown; labrum, palpi, antennae and legs yellowish orange (Fig. 1 f). Length: 2.77 mm; width: 1.39 mm. Head hypognathous at rest; frons and vertex regularly convex, microsculptured, sulcate medially and with scattered strong, deep punctation; sulcus above eyes imprecise; glabrous except for long seta above eye. Clypeus nearly as long as wide at apex, microsculptured, punctate. Labrum transverse, smooth, weakly emarginate at apex. Apical maxillary palpomere spindle-shaped, obliquely cut shortly at apex. Eyes large, bulging, entire. Antennae slender, reaching beyond middle of elytra; scape nearly cylindrical, slightly bent posteriorly, twice as long as thick, strongly narrowed at base; pedicel cup-shaped, half as long as scape; antennomeres 3 – 10 club-shaped, 3 – 6 glossy with sparse long setae, 7 – 11 slightly incrassate, finely granulose, clothed with denser, shorter hairs; antennomeres 3, 4 and 6 subequal, longer than pedicel; antennomeres 5 and 8 – 10 subequal, longer than sixth and shorter than seventh; eleventh antennomere longest, pointed, obliquely cut at apex internally. Pronotum transverse (1.44 x as wide at base than long at middle), convex with stronger slopes at sides and toward anterior angles; anterior border nearly straight, slightly produced forward medially, unmargined, narrower than posterior border, weakly bisinuate, with very short median lobe before scutellum, finely margined; anterior angles obtuse, feebly callous, with setigerous pore anteriorly at middle; posterior angles obtuse, very slightly salient laterally; sides feebly sinuate before base, widest after middle, with explanate margin; surface glabrous, shiny, very finely microsculptured, slightly irregular with transverse depression at anterior half, and with dense, strong deep roundish punctures larger than distance separating them, sometimes confluent. Hypomera smooth, very finely shagreened, with few scattered punctures near basal border; anterior border very feebly convex. Prosternum very narrow before procoxae, with anterior border concave and finely margined; prosternal process relatively wide, about 0.8 x as wide as transverse diameter of procoxae, slightly widened at apex to enclose procoxae posteriorly, finely rugose-punctate at both ends, glossy and punctate at middle, with disheveled long, fine translucent hairs. Mesoventral process as wide as prosternal process, shortly expanded before convex apical border, glossy, with scattered fine punctures and very short fine hairs. Metaventrite slightly longer than mesoventrite, smooth on disc, with fine transverse impressions near basal border and deep narrow median notch at apical border, sides finely microreticulate with sparse punctures and short fine translucent hairs. Scutellum arched, as long as wide at base, round at apex, finely microreticulate. Elytra long, as wide at base as base of pronotum, gently curved at sides, broadest at middle and regularly curved at apex; elytra with very weak transverse impression at basal 1 / 3; humeri strongly marked, unpunctured; eight regular striae on apical half of each elytron, with only striae 6 – 8 reaching humeral area more or less regularly; striae on apical half of elytra impressed with longitudinally cojoined slit-like small punctures or dot-like, deeper punctures in outer rows; punctation in basal half of elytra, including regular outer striae deep, strong, dense and generally confused; punctures of scutellar stria confused with surrounding punctation; short subhumeral, curved stria of large, deep puntures joining premarginal stria before middle of elytron; intervals convex in regularly striated part of elytra; intervals very finely microreticulate. Femora strongly widened medially; anterior and hind tibiae nearly as long as their corresponding femur and mesotibiae slightly shorter; tarsi much shorter than corresponding tibia, with first pro- and mesotarsomeres expanded; claws appendiculate. Abdominal ventrites finely shagreened with sparse fine punctures and short appressed translucent hairs; last visible ventrite weakly emarginate at apex, with small curcular median impression before apical border. Penis (Fig. 2 a) narrow, slender, strongly curved ventrally at base and weakly bent toward apex; more or less parallel-sided except at level of operculum in dorso-ventral view, slightly broadened before converging to apex; dorsal flap of operculum short triangular; apical end of penis expanded as subrectangular tooth with blunt angles. Females (Fig. 4 a): The females of K. anomala sp. nov. lack modified tarsomeres in pro- and mesotarsi, have more convex elytra with raised costae in their lateral declivities, and the punctures on pronotum and in basal area of elytra are stronger and tighter; moreover, at least their head, pronotum, scutellum, suture and all ventral surfaces (as in the paratype) and also the elytra (in all other females examined) are dark metallic bronze. Spermatheca (Fig. 5 e) weakly sigmoidal; cornu bulbous, shorter than wide at base, connected to nodulus, more than twice as long as cornu, narrowed at middle, at level with insertion of spermathecal gland, wider at distal half compared to bulbous basal half, with spermathecal duct inserted at base, opposite to cornu orientation.	en	Gómez-Zurita, Jesús (2018): Description of Kumatoeides gen. nov. (Coleoptera: Chrysomelidae, Eumolpinae) from New Caledonia. Zootaxa 4521 (1): 89-115, DOI: 10.11646/zootaxa.4521.1.4
03C587B7FF83697CC5B9C5FAFCB3FC11.taxon	diagnosis	Diagnosis: The defining trait for this species, absent in all other known species of the genus is the lack of ordered punctation at the base of elytra. Apart from this trait, obvious both in males and females, it can be distinguished from K. metallica sp. nov., the most similar species, and also found in Mt. Mandjélia, by the lack of intense green metallic shine on dorsal surfaces in males.	en	Gómez-Zurita, Jesús (2018): Description of Kumatoeides gen. nov. (Coleoptera: Chrysomelidae, Eumolpinae) from New Caledonia. Zootaxa 4521 (1): 89-115, DOI: 10.11646/zootaxa.4521.1.4
03C587B7FF83697CC5B9C5FAFCB3FC11.taxon	etymology	Derivatio nominis: The name derives from the Greek adjective (fem.) όνώµαλoς, meaning anomalous, unusual, in reference to the scrambling of punctures at the base of elytra, which deviates from the norm for the genus.	en	Gómez-Zurita, Jesús (2018): Description of Kumatoeides gen. nov. (Coleoptera: Chrysomelidae, Eumolpinae) from New Caledonia. Zootaxa 4521 (1): 89-115, DOI: 10.11646/zootaxa.4521.1.4
03C587B7FF83697CC5B9C5FAFCB3FC11.taxon	distribution	Distribution: The few known specimens of this species were found in two distant localities at mid elevations in the mountains of Mandjélia and L'Aoupinié (Fig. 3 a).	en	Gómez-Zurita, Jesús (2018): Description of Kumatoeides gen. nov. (Coleoptera: Chrysomelidae, Eumolpinae) from New Caledonia. Zootaxa 4521 (1): 89-115, DOI: 10.11646/zootaxa.4521.1.4
03C587B7FF826971C5B9C6F3FBC6FF45.taxon	description	(Figs 1 a, 2 b, 3 a, 4 b, 5 b)	en	Gómez-Zurita, Jesús (2018): Description of Kumatoeides gen. nov. (Coleoptera: Chrysomelidae, Eumolpinae) from New Caledonia. Zootaxa 4521 (1): 89-115, DOI: 10.11646/zootaxa.4521.1.4
03C587B7FF826971C5B9C6F3FBC6FF45.taxon	materials_examined	Holotype: Male, New Caledonia, Ponérihouen, Mt. Ounda, 700 m. 11. x. 1969, No. PC-U. 3, leg. Dr. J. Balogh, HOLOTYPE Kumatoeides aulacia sp. nov. J. Gómez-Zurita det. [red label] (HMNH). Paratypes: seven males and two females, New Caledonia, Ponérihouen, Mt. Ounda, 700 m. 11. x. 1969, No. PC-U. 3, leg. Dr. J. Balogh, (3 spp. with PARATYPE Kumatoeides aulacia sp. nov. J. Gómez-Zurita det. [red label]) (HMNH). Other material examined: (1) ten males, eleven females, New Caledonia (N), 20 ° 47 ' S 164 ° 53 ' E, 450 m, Thoven near Tiendanite forest [Hienghène County], 5. ii. 2004, leg. M. Wanat, Kumatoeides aulacia m. J. Gómez- Zurita det. 2018 (MNHW); (2) two females, New Caledonia (N), 21 ° 09 ' S 165 ° 19 ' E, Aoupinié, 530 m, sawmill, ad lucem, 6. ii. 2004, leg. M. Wanat, Kumatoeides aulacia m. J. Gómez-Zurita det. 2018 (MNHW); (3) seven males, one female, New Caledonia (N), 21 ° 09 ' S 165 ° 19 ' E, Aoupinié, 420 – 530 m, road to sawmill, 7. ii. 2004, leg. M. Wanat, Kumatoeides aulacia m. J. Gómez-Zurita det. 2018 (MNHW); (4) one female, New Caledonia (N), 21 ° 11 ' S 165 ° 16 ' E, Aoupinié, 950 – 1000 m, meteo st. — summit, 8. ii. 2004, leg. M. Wanat, Kumatoeides aulacia m. J. Gómez-Zurita det. 2018 (MNHW); (5) one female, New Caledonia (N), 20 ° 57.2 ' S 165 ° 17.5 ' E, Pic d'Amoa, 360 m, forest, 15. i. 2007, leg. M. Wanat & R. Dobosz, Kumatoeides aulacia m. J. Gómez-Zurita det. 2018 (MNHW); (6) one female, New Caledonia (N), 21 ° 11.0 ' S 165 ° 17.6 ' E, Aoupinié, 650 – 800 m, forest, 19. i. 2007, leg. M. Wanat, Kumatoeides aulacia m. J. Gómez-Zurita det. 2018 (MNHW); (7) one male, New Caledonia (N), 21 ° 11.0 ' S 165 ° 17.4 ' E, Aoupinié (gate), 900 – 950 m, 19. i. 2007, leg. M. Wanat, Kumatoeides aulacia m. J. Gómez-Zurita det. 2018 (MNHW); (8) thirteen males and eleven females, New Caledonia (N), - 20.95280 165.29135, Pic d'Amoa (Povila), 400 m, rainforest, 22. xi. 2008, leg. M. Wanat, Kumatoeides aulacia Gómez-Zurita, J. Gómez-Zurita det. 2018 (MNHW); (9) one male, New Caledonia (N), - 20.95659 165.29103, Pic d'Amoa (Povila), 450 m, 0.5 km end of road, 23. xi. 2008, leg. M. Wanat, Kumatoeides aulacia m. J. Gómez-Zurita det. 2018 (MNHW); (10) three females, New Caledonia (N), - 20.95659 165.29103, Pic d'Amoa (Povila), 450 m, road end, at light, 24. xi. 2008, leg. M. Wanat, Kumatoeides aulacia m. J. Gómez-Zurita det. 2018 (MNHW); (11) male, New Caledonia (N), - 21.14890 165.32348, Aoupinié (refuge), 400 m, at light, 25. xi. 2008, leg. M. Wanat, Kumatoeides aulacia m. J. Gómez-Zurita det. 2018 (MNHW); (12) male, New Caledonia (N), - 21.1609 165.3231, Aoupinié (sawmill), 530 m, 26. xi. 2008, leg. M. Wanat, Kumatoeides aulacia m. J. Gómez-Zurita det. 2018 (MNHW); (13) four males and two females, New Caledonia (N), - 21.14890 165.32348, Aoupinié (refuge), 400 m, beating rainforest, 29. xi. 2008, leg. M. Wanat, Kumatoeides aulacia m. J. Gómez-Zurita det. 2018 (MNHW); (14) one male and five females, New Caledonia (N), - 21.17539 165.30952, Aoupinié, 700 m, up of gravelpit, 27. xi. 2008, leg. M. Wanat, Kumatoeides aulacia m. J. Gómez-Zurita det. 2018 (MNHW); (15) three males, one female, New Caledonia (N), - 21.17947 165.30182, Aoupinié, 730 m, Gohapin road junction, 27. xi. 2008, leg. M. Wanat, Kumatoeides aulacia m. J. Gómez-Zurita det. 2018 (MNHW); (16) one male, one female, New Caledonia (N), - 21.17947 165.30182, Aoupinié, 730 m, Gohapin road junction, 27. xi. 2008, leg. M. Wanat, Kumatoeides aulacia m. J. Gómez-Zurita det. 2018 (IBE-JGZ); (17) one female, New Caledonia (N), - 21.18151 165.30048, Aoupinié, 790 – 830 m, 27. xi. 2008, leg. M. Wanat, Kumatoeides aulacia m. J. Gómez-Zurita det. 2018 (MNHW); (18) female, New Caledonia (N), - 21.14890 165.32348, Aoupinié (refuge), 400 m, at light, 28. xi. 2008, leg. M. Wanat, Kumatoeides aulacia m. J. Gómez-Zurita det. 2018 (MNHW); (19) one male and two females, New Caledonia (N), - 21.17947 165.30182, Aoupinié, 700 – 730 m, down Gohapin road junction, 28. xi. 2008, leg. M. Wanat, Kumatoeides aulacia m. J. Gómez- Zurita det. 2018 (IBE-JGZ); (20) eight males, twelve females, New Caledonia (N), - 21.14890 165.32348, Aoupinié (refuge), 400 m, beating rainforest, 29. xi. 2008, leg. M. Wanat, Kumatoeides aulacia m. J. Gómez-Zurita det. 2018 (MNHW); (21) two males, one female, New Caledonia (N), - 21.17539 165.30952, Aoupinié, 700 m, up of gravelpit, 29. xi. 2008, leg. M. Wanat, Kumatoeides aulacia m. J. Gómez-Zurita det. 2018 (MNHW); (22) three females, New Caledonia (N), - 21.17947 165.30182, Aoupinié, 700 – 750 m, up & down Gohapin road junction, 29. xi. 2008, leg. M. Wanat, Kumatoeides aulacia m. J. Gómez-Zurita det. 2018 (MNHW); (23) two males and two females, New Caledonia (N), - 20.9502 165.2932, Pic d'Amoa (Povila), rainforest, 400 – 450 m, 17. xi. 2010, leg. M. Wanat & R. Ruta, Kumatoeides aulacia m. J. Gómez-Zurita det. 2018 (MNHW); (24) two males, New Caledonia (N), - 20.9500 165.2921, Pic d'Amoa (Povila), rainforest, 450 m, night (lamp & beating), 17. xi. 2010, leg. M. Wanat & R. Ruta, Kumatoeides aulacia m. J. Gómez-Zurita det. 2018 (MNHW); (25) male, New Caledonia (N), - 20.9500 165.2921, Pic d'Amoa (Povila), rainforest, 450 m, night (lamp & log), 18. xi. 2010, leg. M. Wanat & R. Ruta, Kumatoeides aulacia m. J. Gómez-Zurita det. 2018 (MNHW); (26) 17 males, 12 females, New Caledonia (N), - 20.9510 165.2912, Pic d'Amoa (Povila), lower rainforest, 400 m, 18. xi. 2010, leg. M. Wanat & R. Ruta, Kumatoeides aulacia m. J. Gómez-Zurita det. 2018 (MNHW); (27) 1 male (IBE-JGZ- 3978), 1 female, New Caledonia (N), - 20.9510 165.2912, Pic d'Amoa (Povila), lower rainforest, 400 m, 18. xi. 2010, leg. M. Wanat & R. Ruta, Kumatoeides aulacia m. J. Gómez-Zurita det. 2018 (IBE-JGZ); (28) one female, New Caledonia (N), - 21.1739 165.3116, Aoupinié, 700 – 800 m, near Gohapin road junction, 20. xi. 2010, leg. M. Wanat & R. Ruta, Kumatoeides aulacia m. J. Gómez-Zurita det. 2018 (MNHW); (29) one female, New Caledonia (N), - 21.1814 165.2879, Aoupinié, 700 – 900 m, Gohapin road junction to gate, 20. xi. 2010, leg. R. Ruta & M. Wanat, Kumatoeides aulacia m. J. Gómez-Zurita det. 2018 (MNHW); (30) four females, New Caledonia (N), - 21.1814 165.2879, Aoupinié, 850 – 900 m, Gohapin road junction to gate, 20. xi. 2010, leg. M. Wanat & R. Ruta, Kumatoeides aulacia m. J. Gómez-Zurita det. 2018 (MNHW); (31) one male, New Caledonia (N), - 21.1739 165.3116, Aoupinié, 700 – 730 m, near Gohapin road junction, 21. xi. 2010, leg. M. Wanat & R. Ruta, Kumatoeides aulacia m. J. Gómez-Zurita det. 2018 (MNHW); (32) three males, one female, New Caledonia (N), - 21.1739 165.3116, Aoupinié, 700 – 750 m, Gohapin road junction down to gravelpit, 21. xi. 2010, leg. M. Wanat & R. Ruta, Kumatoeides aulacia m. J. Gómez-Zurita det. 2018 (MNHW); (33) two females, New Caledonia (N), - 21.1788 165.2786, Aoupinié, 950 m, meteo station to summit, roadside, 21. xi. 2010, leg. M. Wanat & R. Ruta, Kumatoeides aulacia m. J. Gómez-Zurita det. 2018 (MNHW); (34) one male and one female, New Caledonia (N), 21 ° 00.3 ' S 165 ° 14.9 ' E, Tchamba (Wâo Uni), refuge, 400 m, 15. i. 2007, night coll. (lamp & beating), leg. M. Wanat & R. Dobosz, Kumatoeides aulacia m. J. Gómez-Zurita det. 2018 (MNHW); (35) one female, New Caledonia (N), 21 ° 00.3 ' S 165 ° 14.9 ' E, Tchamba (Wâo Uni), refuge, 400 m, 16. i. 2007, night coll. (lamp & beating), leg. M. Wanat & R. Dobosz, Kumatoeides aulacia m. J. Gómez-Zurita det. 2018 (MNHW); (36) two females, New Caledonia (N), 21 ° 00.2 ' S 165 ° 14.6 ' E, Tchamba (Wâo Uni), 500 – 530 m, forest, old northern road, 16. i 2007, leg. M. Wanat, Kumatoeides aulacia m. J. Gómez-Zurita det. 2018 (MNHW). Body orange testaceous with irregular infuscate areas on pronotum and elytra (at base and along some intervals); peripheral rims of pronotum and elytra and striae on elytra slightly darkened; thoracic ventrites and trochanters darkened; basal antennomeres, palpi and legs more or less uniformly yellowish orange, paler than rest of body, and apical antennomeres slightly infuscate (Fig. 1 a). Length: 2.67 mm (range: 2.24 – 2.94 mm); width: 1.33 mm (range: 1.12 – 1.41 mm). Head hypognathous at rest; frons and vertex regularly convex, finely microsculptured, with obsolete median longitudinal furrow and rather uniformly, deeply punctured, except on frontoclypeal area, sulcate above eyes with sulcus weakly impressed but visible above supra-antennal swellings; mostly glabrous except for long seta at middle of supraocular furrow and sparse short, very fine appressed yellowish setae near eyes and on short genae. Clypeus as long as wide at apex, punctured with smaller punctation than frons. Labrum small, smooth, about 1.5 x as wide as long, emarginate at apex. Last maxillary palpomeres spindle-shaped, obliquely cut at apex. Eyes large, bulging, weakly emarginate at inner border. Antennae slender, reaching beyond middle of elytra; scape nearly cylindrical, slightly bent posteriorly, twice as long as thick, strongly narrowed at base; pedicel cup-shaped, half as long as scape; antennomeres 3 – 10 club-shaped, 3 – 6 glossy with sparse long setae, 7 – 11 slightly incrassate, finely granulose, clothed with dense, short hairs; third antennomere slightly longer and fourth as long as pedicel; fifth antennomere longer than third and sixth; seventh anntenomere longer than fifth; antennomeres 8 – 10 subequal, as long as fifth; eleventh antennomere longest, pointed, obliquely cut at apex internally. Pronotum transverse (1.38 x as wide at base than long at middle), convex with stronger declivity toward anterior angles and subtle transverse, discontinuous impression at anterior half; anterior border weakly convex, produced forward medially, unmargined, narrower than posterior border, weakly bisinuate, with very short median lobe before scutellum, finely margined; anterior angles obtuse, blunt, slightly produced forward, with trichobothria displaced toward anterior border bearing long anterior setae; posterior angles obtuse, weakly produced laterally, with trichobothria at angle; sides slightly bisinuate, widest in front of middle, with narrow explanate margin; surface glabrous, shiny, very finely microsculptured, slightly irregular, with dense, deep, slightly elongate punctures, as big as or larger than distance separating them. Hypomera smooth, very finely microreticulate, unpunctured; anterior border straight. Prosternum short, narrower before than behind procoxae; prosternal process about half as wide as transverse diameter of procoxa and expanded apically enclosing procoxae posteriorly, smooth, glossy, with fine disheveled yellowish hairs. Scutellum arched, as long as wide at base, blunt at apex, finely microreticulated. Elytra long, basally as wide as base of pronotum, gently curved at sides, widest at middle and regularly curved at apex; elytra with transverse impression at basal 1 / 3; humeri strongly marked, unpunctured; eight complete regular striae on each elytron with longitudinally conjoined slit-like small punctation or dot-like individual, large punctures in 5 – 6 outer striae basally; additional short scutellar stria reaching transversely depressed area and short subhumeral dotted stria joining marginal bead of punctures before middle of elytron; intervals weakly convex basally and on disc, more convex at apical and lateral declivities of elytra; intervals unpunctured, very finely microreticulate. Mesoventrite relatively long, about as wide as prosternal process, slightly spatulate. Metaventrite short, glossy on disc, with short posterior median incision, finely granulose at sides, with fine posteriorly recumbent translucent hairs. All femora subequal, unarmed, narrow basally and strongly inflated medially; tibiae weakly curved, slightly shorter than corresponding femur, gradually enlarged toward apex; tarsi much shorter than corresponding tibia, with first pro- and mesotarsomeres conspicuously dilated in males; claws appendiculate. Abdominal ventrites finely shagreened, with imperceptible punctures and fine translucent hairs; last ventrite shortly emarginate at apex. Penis (Fig. 2 b) slender, moderately and relatively regularly curved in lateral view, with apical area straight and acute; laterally compressed at middle in dorso-ventral view, slightly expanded at level with operculum and converging in regular curve toward apex; dorsal flap of operculum short, transverse; apical end of penis expanded as axehead, with apical border feebly curved and wider at both ends than at base of apical expansion. Females (Fig. 4 b): Females have first pro- and mesotarsomeres unmodified and their humeral interval is raised as a keel in basal half of elytra. Moreover, females often have their scutellum, the sutural interval, elytral intervals 4 and 5 on disc, sides and base of pronotum medially and apical antennomeres darkened. Spermatheca (Fig. 5 b) small, consisting of round bulbous cornu and curved, progressively narrowing nodulus slightly twisted at base and connected at angle to spermathecal duct.	en	Gómez-Zurita, Jesús (2018): Description of Kumatoeides gen. nov. (Coleoptera: Chrysomelidae, Eumolpinae) from New Caledonia. Zootaxa 4521 (1): 89-115, DOI: 10.11646/zootaxa.4521.1.4
03C587B7FF826971C5B9C6F3FBC6FF45.taxon	diagnosis	Diagnosis: The males of Kumatoeides aulacia sp. nov. could be confused with three other brownish Kumatoeides, including K. costata (Jolivet, Verma et Mille), K. megale sp. nov. and K. millei sp. nov. K. aulacia (and the latter two species) has the scutellar row of punctures in a more or less continuous stria, while the punctures in K. costata are individualized. K. megale and K. millei are larger than K. aulacia, with first pro- and mesotarsomeres proportionally longer and broader. The males of the four species can be distinguished by their distinctive penises and, in general, the females of K. aulacia can be recognized by their pattern of darkened areas, but also by the marked posthumeral costa on elytra. All these species, with slight transverse depression at basal third of elytra, can be distinguished from others in the genus by the lack of metallic shine, by the close, strong punctation of pronotum, and by the completely ordered punctation of elytra.	en	Gómez-Zurita, Jesús (2018): Description of Kumatoeides gen. nov. (Coleoptera: Chrysomelidae, Eumolpinae) from New Caledonia. Zootaxa 4521 (1): 89-115, DOI: 10.11646/zootaxa.4521.1.4
03C587B7FF826971C5B9C6F3FBC6FF45.taxon	etymology	Derivatio nominis: The name is a feminine adjective derived from the Greek noun αυλάκι, meaning groove or furrow, referring to the channeled sculpture of elytra.	en	Gómez-Zurita, Jesús (2018): Description of Kumatoeides gen. nov. (Coleoptera: Chrysomelidae, Eumolpinae) from New Caledonia. Zootaxa 4521 (1): 89-115, DOI: 10.11646/zootaxa.4521.1.4
03C587B7FF826971C5B9C6F3FBC6FF45.taxon	distribution	Distribution: The species is known from localities in the southern part of the Massif du Panié and central areas of the Central Range (Chaîne centrale), mainly in and around L'Aoupinié (Fig. 3 a).	en	Gómez-Zurita, Jesús (2018): Description of Kumatoeides gen. nov. (Coleoptera: Chrysomelidae, Eumolpinae) from New Caledonia. Zootaxa 4521 (1): 89-115, DOI: 10.11646/zootaxa.4521.1.4
03C587B7FF8F6973C5B9C230FE51FB81.taxon	description	(Figs 1 d, 2 c, 3 b, 4 c, 5 a) Jolivet et al. (2007 a) described a new species of Eumolpinae from New Caledonia which had the " elytra punctuate in lines in grooves " as a distinctive character and placed it in a new genus, Montrouzierella Jolivet, Verma et Mille, described in the same article. The explicit mention of the unusual grooves on the elytra, as well as other traits mentioned in the succinct description of M. costata Jolivet, Verma et Mille, including small size and heavy punctation on pronotum, hinted at the possibility of this species belonging to Kumatoeides gen. nov. instead. The original description also mentioned traits absent in any other known species of the genus, namely dense pilosity on pronotum and abdomen. However, the examination of the holotype of M. costata held in the collection of the Muséum national d'Histoire naturelle (MNHN, Paris, France), even if in poor shape (Fig. 1 d), showed that the latter characters are actually absent in the type and that it fits the description of Kumatoeides gen. nov. in every respect. A number of specimens very similar to this type and from the same geographic area in the small central counties of Farino, Moindou and Sarraméa, specifically from the valleys of the Fonwhary and Moindou rivers, were available for study in the collections of the author and the Museum of Natural History of the University of Wroclaw (MNHW, Wroclaw, Poland). Based on this new information, M. costata Jolivet, Verma et Mille is combined with the new genus proposed in this study as Kumatoeides costata (Jolivet, Verma et Mille) comb. nov., and redescribed below.	en	Gómez-Zurita, Jesús (2018): Description of Kumatoeides gen. nov. (Coleoptera: Chrysomelidae, Eumolpinae) from New Caledonia. Zootaxa 4521 (1): 89-115, DOI: 10.11646/zootaxa.4521.1.4
03C587B7FF8F6973C5B9C230FE51FB81.taxon	materials_examined	Holotype: Nouvelle-Calédonie, Farino / M. Barbou, Lat. S 21.63286 Long. E 165.70684, collecté sur: Divers arbres de forêt, le 23 / 09 / 04 [original publication gives 29. ix. 2004], par battage, par S. Cazères, Type, Montrouzierella costata P. Jolivet dét. 0 6, Muséum Paris 2014 coll. P. Jolivet, HOLOTYPE [red label], HOLOTYPE Montrouzierella costata Jolivet, Verma & Mille, 2007, MNHN EC 8125 (MNHN). Other material examined: (1) one male, New Caledonia, Province Sud, forest above Sarraméa, 2 – 5. xii. 2001, K. A. Johanson, T. Pape & B. Viklund leg., Kumatoeides costata (Jolivet et al.) J. Gómez-Zurita det. 2018 (IBE- JGZ); (2) one female, New Caledonia (S), - 21.6274 165.8647, Sarraméa, 150 – 350 m, trail to Plateau de Dogny, 8. xi. 2010, leg. M. Wanat & R. Ruta, Kumatoeides costata (Jolivet et al.) J. Gómez-Zurita det. 2018 (MNHW); (3) five males, four females, New Caledonia (S), - 21.6229, 165.8684, Sarraméa, 300 – 560 m, trail to Plateau de Dogny, 9. xi. 2010, leg. M. Wanat & R. Ruta, Kumatoeides costata (Jolivet et al.) J. Gómez-Zurita det. 2018 (MNHW); (4) one male, one female, New Caledonia (S), - 21.6229, 165.8684, Sarraméa, 300 – 560 m, trail to Plateau de Dogny, 9. xi. 2010, leg. M. Wanat & R. Ruta, Kumatoeides costata (Jolivet et al.) J. Gómez-Zurita det. 2018 (IBE-JGZ). Head, most of pronotum, broad margins of scutellum and most of ventral surfaces pale brown in males and rather uniformly dark brown in females, darker in ventral surfaces; elytra and anterior and posterior areas medially on pronotum orange brown in males, and elytra brown with darkened suture in females; labrum, antennae (five apical antennomeres darkened in females), mouth appendages and legs pale orange (Fig. 1 d). Length: 2.74 mm (range: 2.33 – 2.74 mm); width: 1.43 mm (range: 1.22 – 1.47 mm). Head hypognathous at rest; frons and vertex convex, finely microreticulate, with dense, relatively strong punctures, without trace of frontal suture and slightly depressed but not furrowed above eyes, with long seta dorsally in depressed area. Clypeus short, flat, markedly emarginate at apical border, finely microsculptured with fine sparse punctures basally. Labrum short, transverse, weakly emarginate apically, finely microreticulate, unpunctured. Last maxillary palpomere elongate, conical at apical half. Eyes large, bulging, weakly emarginate at inner border. Antennae slender, reaching beyond middle of elytra, feebly clavate from and beyond seventh antennomere; scape thick, cylindrical, slightly curved posteriorly; pedicel thickened at middle, half as long as scape; antennomeres 3 – 4 subequal, slightly longer than pedicel, feebly widened toward apex; fifth antennomere nearly 1.5 x as long as fourth, slightly enlarged apically; sixth antennomere slightly longer than fourth, feebly dilated toward apex; seventh antennomere longer than fifth, slightly over 1.5 x as long as fourth, more incrassate at apex and hairier than preceding antennomeres; antennomeres 8 – 11 finely granulose, rather densely covered with fine whitish pilosity; antennomeres 8 – 10 progressively longer, tenth antennomere as long as seventh, slightly incrassate; eleventh antennomere longer than seventh, feebly enlarged toward apical third, acuminate at apex. Pronotum transverse (1.60 x as wide at base as long at middle), laterally convex, strongly slopping toward sides and anterior angles; anterior border straight, narrowly margined at sides, near angles, unmargined medially; sides weakly curved, widest at middle and finely margined; posterior border convex, finely margined; angles obtuse, anterior angles with margin incrassate and large trichobothria at anterior border; surface glabrous, finely microreticulate, with dense and strong punctation, finer on disc. Hypomera finely microreticulate, with few scattered fine punctures; anterior border of hypomera straight. Mesepimera and mesanepisterna shagreened, unpunctured. Metaventrite smooth on disc, very finely shagreened at sides, with sparse punctation and short appressed, whitish setae. Scutellum arched, longer than wide at base, finely microreticulate, unpunctured. Elytra long, as wide basally as base of pronotum, with humeral angles and sides feebly curved, widest around middle, and regularly curved toward sutural angle at apical third; humeri strongly marked, unpunctured; convexity of elytra feebly depressed at basal 1 / 3; elytra with eight regular striae of fine slit-like punctures on disc and apical half and strong round punctures basally, larger and stronger in subhumeral area; short scutellar stria of 9 – 10 small round punctures reaching prebasal depression and short subhumeral line of strong punctures merged with marginal line of punctures posteriorly; intervals convex, unpunctured. Femora dilated medially; tibiae shorter than corresponding femur, gradually expanded toward apex; tarsi shorter than tibiae, with first pro- and mesotarsomeres enlarged, longer than wide, slightly longer than next two tarsomeres together; claws appendiculate. Penis (Fig. 2 c) short, dorso-ventrally flattened and strongly curved ventrally, slightly compressed laterally at middle; operculum elongate, slightly widened, with curved sides gradually converging toward apex, prolonged as blunt median, elongate lobe. Females (Fig. 4 c): The females of K. costata (Jolivet, Verma et Mille) are less slender than the males, slightly more convex, with prebasal depression on elytra less pronounced, and more or less uniformly darker, particularly on head, pronotum and scutellum. Spermatheca (Fig. 5 a) with long bulbous cornu connected at angle with elongate, curved nodulus bent toward cornu, with spermathecal gland inserted medially and narrowing basally at connection with spermathecal duct.	en	Gómez-Zurita, Jesús (2018): Description of Kumatoeides gen. nov. (Coleoptera: Chrysomelidae, Eumolpinae) from New Caledonia. Zootaxa 4521 (1): 89-115, DOI: 10.11646/zootaxa.4521.1.4
03C587B7FF8F6973C5B9C230FE51FB81.taxon	diagnosis	Diagnosis: This species is very similar to other species belonging to the K. aulacia group, particularly to K. millei sp. nov., collected in close geographic proximity. The scutellar stria of K. costata (Jolivet, Verma et Mille) has individual, recognizable punctures, while the other species in the group have them seamed, almost like a straight line (other striae on elytra show a similar pattern). Moreover, K. megale sp. nov. and K. millei are larger, have a strongly marked supraocular furrows, missing in K. costata, and their punctation of pronotum can be vermiculate, while it is merely strong in K. costata. The males of these species and any other species recognized so far can be easily distinguished by their distinctive penises. The females of K. costata and K. millei are very similar, also in coloration, with the former being smaller and with isolated punctures in scutellar striae; their spermathecae are clearly different too.	en	Gómez-Zurita, Jesús (2018): Description of Kumatoeides gen. nov. (Coleoptera: Chrysomelidae, Eumolpinae) from New Caledonia. Zootaxa 4521 (1): 89-115, DOI: 10.11646/zootaxa.4521.1.4
03C587B7FF8F6973C5B9C230FE51FB81.taxon	distribution	Distribution: The species is distributed in low elevations of the Fonwhary and Moindou river valleys in central Grande Terre (Fig. 3 b).	en	Gómez-Zurita, Jesús (2018): Description of Kumatoeides gen. nov. (Coleoptera: Chrysomelidae, Eumolpinae) from New Caledonia. Zootaxa 4521 (1): 89-115, DOI: 10.11646/zootaxa.4521.1.4
03C587B7FF8D6972C5B9C143FC50FA0D.taxon	description	(Figs 2 d, 3 a, 4 e, 5 h, 6 a)	en	Gómez-Zurita, Jesús (2018): Description of Kumatoeides gen. nov. (Coleoptera: Chrysomelidae, Eumolpinae) from New Caledonia. Zootaxa 4521 (1): 89-115, DOI: 10.11646/zootaxa.4521.1.4
03C587B7FF8D6972C5B9C143FC50FA0D.taxon	materials_examined	Holotype: Male, New Caledonia (N), Tiendanite [Hienghène County], 20 ° 44.8 ' S 164 ° 52.8 ' E, 50 m, 12. i. 2007, at light, leg. M. Wanat & R. Dobosz, HOLOTYPE Kumatoeides leptalei sp. nov. J. Gómez-Zurita det. [red label] (MNHN). Paratype: Male, New Caledonia (N), Tiendanite [Hienghène County], 20 ° 44.8 ' S 164 ° 52.8 ' E, 50 m, 12. i. 2007, at light, leg. M. Wanat & R. Dobosz, PARATYPE Kumatoeides leptalei sp. nov. J. Gómez-Zurita det. [red label] (MNHW). Other material examined: (1) one female, New Caledonia (N), Tiendanite [Hienghène County], 20 ° 44.8 ' S 164 ° 52.8 ' E, 50 m, 12. i. 2007, at light, leg. M. Wanat & R. Dobosz, Kumatoeides leptalei m. J. Gómez-Zurita det. 2018. Body rather uniformly orangish brown, brighter on pronotum; antennae palpi and legs yellowish orange (Fig. 6 a). Length: 2.23 mm (range: 2.23 – 2.36 mm); width: 1.21 mm (range: 1.21 – 1.26 mm). Head hypognathous at rest; frons and vertex regularly convex, very finely microsculptured, glossy, sparsely, but neatly punctured; impressed above eyes but supra-antennal swellings almost undifferentiated; glabrous except for long seta at middle of supraocular furrow. Clypeus as long as wide at apex, smooth, glossy, with few small punctures at base. Labrum small, smooth, about 1.3 x as wide as long, very feebly emarginate at apex. Last maxillary palpomere spindle-shaped, acute at apex. Eyes large, bulging, weakly emarginate at inner border. Antennae slender, reaching beyond middle of elytra; scape nearly cylindrical, slightly bent posteriorly, twice as long as thick, strongly narrowed at base; pedicel cup-shaped, half as long as scape; antennomeres 3 – 10 slender, club-shaped, 3 – 6 glossy with sparse long setae, 7 – 11 feebly incrassate, finely granulose, clothed with dense, short, fine hairs; antennomeres 3 and 4 subequal, slightly longer than pedicel, slightly shorter than sixth; fifth antennomere longer than sixth, nearly as long as seventh; antennomere seventh longest; antennomeres 8 – 10 subequal, about as long as fifth; eleventh antennomere pointed and obliquely cut at apex internally. Pronotum transverse (1.47 x as wide at base than long at middle), convex, with stronger declivity toward anterior angles; anterior border produced forward, straight, unmargined, narrower than posterior border, weakly convex, with very short median lobe before scutellum, and finely margined, more so on median lobe; anterior angles obtuse, with feebly developed calli, not produced forward; posterior angles obtuse, slightly thickened; sides regularly curved, widest at middle, with fine explanate margin; surface glabrous, shiny, very finely microsculptured, more or less uniformly but shallowly punctured with punctures larger than punctures on frons and generally smaller than distance separating them. Hypomera smooth, finely microreticulate, unpunctured; straight at anterior border. Anterior border of prosternum concave, finely margined, narrow before procoxae; prosternal process 0.7 x as wide as transverse diameter of procoxae and expanded apically to enclose procoxae posteriorly, smooth with sparse disheveled fine yellowish hairs. Mesoventrite as wide as prosternal process, spatulate at apex, smooth, unpunctured. Metaventrite relatively short, as long as mesoventrite, smooth, glabrous and finely impressed longitudinally on disc, with deep median apical notch, finely microreticulate with posteriorly recumbent fine yellowish hairs at sides. Scutellum arched, as long as wide at base, round at apex, smooth. Elytra long, basally as wide as base of pronotum, gently curved at sides, widest before middle and progressively curved at apex; elytra regularly convex, without transverse impression at basal 1 / 3; humeri marked, unpunctured; eight complete regular striae on each elytron with longitudinal series of weakly impressed dot-like punctures at basal half and slit-like punctures at apical half of elytra, punctures larger and deeper in humeral area; additional short scutellar stria of seven individual punctures reaching basal 1 / 3 of elytra, and short subhumeral curved row of 5 – 6 punctures joining marginal stria before middle of elytron; intervals nearly flat basally and on disc, and feebly convex laterally and apically on elytra; intervals unpunctured, smooth. Abdominal ventrites finely shagreened, unpunctured, with sparse posteriorly recumbent fine hairs; apical visible ventrite emarginate at apex and slightly impressed medially. Penis (Fig. 2 d) stocky, curved at base and apically in lateral view, sinuous at apex; sides widened and curved at level with operculum, with small projecting blunt tooth medially at apex. (Both type specimens had their penises poorly sclerotized, with only their tips darkened.) Females (Fig. 4 e): The only female available for study is very similar to the two males, except for the unmodified tarsi and the slightly darker color of head and pronotum. Spermatheca (Fig. 5 h) very small and poorly sclerotized, with cornu bent, slightly dilated and round at apical half, nodulus small, nearly spherical with lateral, slightly bent tubular connection to spermathecal duct.	en	Gómez-Zurita, Jesús (2018): Description of Kumatoeides gen. nov. (Coleoptera: Chrysomelidae, Eumolpinae) from New Caledonia. Zootaxa 4521 (1): 89-115, DOI: 10.11646/zootaxa.4521.1.4
03C587B7FF8D6972C5B9C143FC50FA0D.taxon	diagnosis	Diagnosis: This is by far the smallest species of Kumatoeides gen. nov., but can be distinguished from all other species in the genus by the lack of transverse impressions on elytra, which are more or less regularly curved, and by the spaced punctation on pronotum, only found in K. tarsalis sp. nov., but this species is larger, darker and has the typical prebasal impression on elytra of this genus.	en	Gómez-Zurita, Jesús (2018): Description of Kumatoeides gen. nov. (Coleoptera: Chrysomelidae, Eumolpinae) from New Caledonia. Zootaxa 4521 (1): 89-115, DOI: 10.11646/zootaxa.4521.1.4
03C587B7FF8D6972C5B9C143FC50FA0D.taxon	etymology	Derivatio nominis: The name derives from the Latin transliteration of the Greek adjective (fem.) λεπταλέῃ, meaning fine and delicate, referring both to the small size of the species and the relative subtleness of its punctation.	en	Gómez-Zurita, Jesús (2018): Description of Kumatoeides gen. nov. (Coleoptera: Chrysomelidae, Eumolpinae) from New Caledonia. Zootaxa 4521 (1): 89-115, DOI: 10.11646/zootaxa.4521.1.4
03C587B7FF8D6972C5B9C143FC50FA0D.taxon	distribution	Distribution: Only three specimens of K. leptalei sp. nov. were available and both from the same low elevation locality, Tiendanite (Hienghène County), near Poindimié (Fig. 3 a).	en	Gómez-Zurita, Jesús (2018): Description of Kumatoeides gen. nov. (Coleoptera: Chrysomelidae, Eumolpinae) from New Caledonia. Zootaxa 4521 (1): 89-115, DOI: 10.11646/zootaxa.4521.1.4
03C587B7FF8C6975C5B9C0C5FADAF8DC.taxon	description	(Figs 1 b, 2 e, 3 b)	en	Gómez-Zurita, Jesús (2018): Description of Kumatoeides gen. nov. (Coleoptera: Chrysomelidae, Eumolpinae) from New Caledonia. Zootaxa 4521 (1): 89-115, DOI: 10.11646/zootaxa.4521.1.4
03C587B7FF8C6975C5B9C0C5FADAF8DC.taxon	materials_examined	Holotype: Male, New Caledonia (N), Col des Roussettes, nr. refuge, - 21.4074 165.5250, 530 m, rainforest, 2. xii. 2010, leg. M. Wanat & R. Ruta, HOLOTYPE Kumatoeides megale sp. nov. J. Gómez-Zurita det. [red label] (MNHN). Paratype: Male, New Caledonia (N), Col des Roussettes (refuge), - 21.4070 165.5265, 520 m, 2. xii. 2010, at light, leg. M. Wanat & R. Ruta, PARATYPE Kumatoeides megale sp. nov. J. Gómez-zurita det. [red label] (MNHW). Other material examined: (1) two males, New Caledonia (N), - 21.4079 165.5249, 520 m, Col des Roussettes (refuge), 1. xii. 2010, at light, leg. M. Wanat & R. Ruta, Kumatoeides megale m. J. Gómez-Zurita det. 2018 (MNHW); (2) one male, IBE-JGZ- 3974, New Caledonia (N), - 21.4079 165.5249, 520 m, Col des Roussettes (refuge), 1. xii. 2010, at light, leg. M. Wanat & R. Ruta, Kumatoeides megale m. J. Gómez-Zurita det. 2018 (IBE-JGZ). Labrum, head, elytra, hypomera and abdominal ventrites hazelnut brown; pronotum, meso- and metathoracic ventrites and epimera with darker brownish areas and faint metallic reflection on pronotum and scutellum; antennae, palpi and legs yellowish orange (Fig. 1 b). Length: 3.20 mm (range: 3.07 – 3.20 mm); width: 1.54 mm. Head hypognathous at rest; frons and vertex weakly convex, microsculptured, finely sulcate medially with scattered strong punctures; deeply furrowed above eyes with supraantennal swellings only indicated by deep punctures posteriorly; glabrous except for long seta at middle of supraocular furrow and sparse, very short setae near and on short genae. Clypeus as long as wide at apex, microsculptured, with punctures medially. Labrum about 1.5 x as wide as long, microreticulate and very feebly emarginate at apex. Last maxillary palpomere elongate, conical at apical half. Eyes large, bulging, weakly emarginate at inner border. Antennae slender, reaching beyond middle of elytra; scape elongate, nearly cylindrical, more than twice longer than thick, bent posteriorly near base, rather gradually narrowing at base; pedicel cup-shaped, half as long as scape; antennomeres 3 – 10 slender, clubshaped, 3 – 6 glossy with sparse long setae, 7 – 11 slightly incrassate, finely granulose, clothed with dense, short hairs; antennomeres 3, 4 and 6 subequal, slightly longer than pedicel; fifth antennomere as long as antennomeres 8 – 10, slightly shorter than seventh; eleventh antennomere longest, pointed, obliquely cut at apex internally. Pronotum transverse (1.47 x as wide at base than long at middle), convex but with slight transverse impressions at sides of apical half; anterior border convex, slightly produced forward medially, with very fine margin at sides, narrower than posterior border, weakly bisinuate, with very short median lobe before scutellum, and margined; anterior angles subround, largely explanate, not produced forward, with setigerous pore anteriorly at middle; posterior angles obtuse, with trichobothria at angle and long obliquely and dorsally bent golden setae; sides slightly sinuate at basal half, widest after middle, with rather wide explanate margin; surface glabrous, shiny, very finely microsculptured, slightly irregular, with dense deep elongate punctures as big as and generally larger than distance separating them. Hypomera smooth, unpunctured; anterior border very feebly convex. Prosternal process relatively narrow, about 0.7 x as wide as transverse diameter of procoxae, transversally expanded at apex to enclose procoxae posteriorly, smooth with scattered disheveled fine hairs. Mesosternal process as wide as prosternal process, spatulate at apex, smooth, unpunctured. Metaventrite shorter than mesoventrite at middle, smooth on disc, with fine longitudinally impressed suture and short apical median notch; sides finely microreticulate with scattered short fine appressed hairs. Scutellum arched, as long as wide at base, round at apex, very finely microsculptured. Elytra long, basally as wide as base of pronotum, gently curved at sides, widest at middle and regularly curved at apex; elytra with transverse impression at basal 1 / 3; humeri strongly marked, unpunctured; eight complete regular striae on each elytron with longitudinally conjoined slit-like small punctation or dot-like individual, large punctures in striae 5 – 9 basally; additional short scutellar stria reaching transversely depressed area and short subhumeral curved stria of individual punctures joining marginal bead of punctures before middle of elytron; intervals convex, more convex at apical and lateral declivities of elytra; intervals unpunctured, very finely microreticulate. First pro- and mesotarsomeres elongate, as long as next two tarsomeres combined, and widened; claws appendiculate. Abdominal ventrites finely shagreened, with scattered posteriorly recumbent fine translucent hairs; last ventrite weakly emarginate at apex, with short median longitudinal impression. Penis (Fig. 2 e) long, stout, narrower close to base in dorso-ventral view and gradually widening, widest at level with operculum, nearly straight in lateral view, widest near base and progressively narrowing toward apex; apex sinuous in lateral view, with tip bent dorsally; dorsal flap of operculum subrectangular, transverse; sides of operculum feebly curved, converging apically to short, broad axehead apical expansion. Females: Unknown.	en	Gómez-Zurita, Jesús (2018): Description of Kumatoeides gen. nov. (Coleoptera: Chrysomelidae, Eumolpinae) from New Caledonia. Zootaxa 4521 (1): 89-115, DOI: 10.11646/zootaxa.4521.1.4
03C587B7FF8C6975C5B9C0C5FADAF8DC.taxon	diagnosis	Diagnosis: Kumatoeides megale sp. nov. is externally nearly identical to K. millei sp. nov., but they can be easily recognized based on conspicuous differences in their male genitalia (see Figs 2 e and 2 g). This species is also like a larger version of K. aulacia sp. nov. Apart from the larger size, both species can be distinguished by the relative proportions of first protarsomeres in males, longer in K. megale. From other species, it can be distinguished by the lack of metallic shine, by the close, strong punctation of pronotum, and by the completely ordered punctation of elytra.	en	Gómez-Zurita, Jesús (2018): Description of Kumatoeides gen. nov. (Coleoptera: Chrysomelidae, Eumolpinae) from New Caledonia. Zootaxa 4521 (1): 89-115, DOI: 10.11646/zootaxa.4521.1.4
03C587B7FF8C6975C5B9C0C5FADAF8DC.taxon	etymology	Derivatio nominis: The name is the Latin transliteration of the Greek adjective (fem.) µεγάλη, meaning large, to indicate that this is the largest species of the genus known so far.	en	Gómez-Zurita, Jesús (2018): Description of Kumatoeides gen. nov. (Coleoptera: Chrysomelidae, Eumolpinae) from New Caledonia. Zootaxa 4521 (1): 89-115, DOI: 10.11646/zootaxa.4521.1.4
03C587B7FF8C6975C5B9C0C5FADAF8DC.taxon	distribution	Distribution: Only five male specimens of the new species are currently known, and all from nearby localities in Col de Roussettes, in the heart of the Massif de la Boghen in the Central Range (Chaîne centrale) (Fig. 3 b).	en	Gómez-Zurita, Jesús (2018): Description of Kumatoeides gen. nov. (Coleoptera: Chrysomelidae, Eumolpinae) from New Caledonia. Zootaxa 4521 (1): 89-115, DOI: 10.11646/zootaxa.4521.1.4
03C587B7FF886969C5B9C5B0FD7CFBED.taxon	description	(Figs 1 e, 2 f, 3 a, 4 d, 5 d)	en	Gómez-Zurita, Jesús (2018): Description of Kumatoeides gen. nov. (Coleoptera: Chrysomelidae, Eumolpinae) from New Caledonia. Zootaxa 4521 (1): 89-115, DOI: 10.11646/zootaxa.4521.1.4
03C587B7FF886969C5B9C5B0FD7CFBED.taxon	materials_examined	Holotype: Male, New Caledonia (N), Mandjélia, summit, 20 ° 23.9 ' S 164 ° 31.9 ' E, 750 – 780 m, montane rainforest, 12. i. 2007, beating, leg. M. Wanat & R. Dobosz, HOLOTYPE Kumatoeides metallica sp. nov. J. Gómez-Zurita det. [red label] (MNHN). Paratypes: Two males, one female, New Caledonia (N), 20 ° 23.9 ' S 164 ° 32.0 ' E, Mandjélia (subsummit), 700 – 750 m, night beating, 11. i. 2007, leg. M. Wanat, Muzeum Przyrodnicze Uniwersytetu Wroclawskiego, PARATYPE Kumatoeides metallica sp. nov. J. Gómez-Zurita det. [red label] (MNHW). Other material examined: (1) two females, New Caledonia (N), - 20.40058 164.52678, Mt Mandjélia, 700 – 780 m, montane rainforest, 19. xi. 2008, leg. M. Wanat, Kumatoeides metallica m. J. Gómez-Zurita det. 2018 (MNHW); (2) three males, New Caledonia (N), - 20.40058 164.52678, Mt Mandjélia, 700 m, at light, 19 – 20. xi. 2008, leg. M. Wanat, Kumatoeides metallica m. J. Gómez-Zurita det. 2018 (MNHW); (3) one male, New Caledonia (N), - 20.40058 164.52678, Mt Mandjélia, 700 m, at light, 19 – 20. xi. 2008, leg. M. Wanat, Kumatoeides metallica m. J. Gómez-Zurita det. 2018 (IBE-JGZ); (4) one male, three females, New Caledonia (N), - 20.40007 164.52788, Mt Mandjélia, 700 – 780 m, montane rainforest, 20. xi. 2008, leg. M Wanat, Kumatoeides metallica m. J. Gómez-Zurita det. 2018 (MNHW); (5) one male, three females, New Caledonia (N), - 20.40007 164.52788, Mt Mandjélia, 700 – 780 m, montane rainforest, 21. xi. 2008, leg. M Wanat, Kumatoeides metallica m. J. Gómez-Zurita det. 2018 (MNHW). Head, pronotum, scutellum and elytra dark reddish brown with strong dark greenish metallic reflections, less intense on head; ventral surfaces dark reddish brown, darker and with hint of metallic luster on meso- and metathoracic ventrites and episterna; labrum, palpi, antennae and legs yellowish orange (Fig. 1 e). Length: 2.67 mm (range: 2.48 – 3.10 mm); width: 1.38 mm (range: 1.20 – 1.61 mm). Head hypognathous at rest; frons and vertex regularly convex, finely microsculptured, densely, deeply punctured, except medially on frons, sulcate above eyes with sulcus impressed, also punctured above transverse supra-antennal swellings, nearly touching at middle; glabrous except for long seta at middle of supraocular furrow and sparse short, fine appressed whitish setae on short genae. Clypeus nearly as long as wide at apex, punctured at base with smaller punctation than frons. Labrum small, smooth, about 1.5 x as wide as long, weakly emarginate at apex. Last maxillary palpomere elongate, gradually narrowing at apical half toward blunt apex. Eyes large, bulging, entire. Antennae slender, reaching beyond middle of elytra; scape nearly cylindrical, slightly bent posteriorly, twice as long as thick, strongly narrowed at base; pedicel short, nearly spherical, less than half as long as scape; antennomeres 3 – 10 elongate, club-shaped, 3 – 6 glossy with sparse golden setae, 7 – 11 slightly incrassate, finely granulose, clothed with dense, short, fine hairs; antennomeres 3 – 4 subequal, longer than pedicel; fifth antennomere longer than fourth and sixth, and slightly shorter than seventh; antennomeres 8 – 10 subequal, as long as sixth; eleventh antennomere longest, pointed, obliquely cut at apex internally. Pronotum transverse (1.45 x as wide at base than long at middle), convex with stronger declivity toward anterior angles; anterior border straight, produced forward medially, with very fine margin near anterior angles, narrower than posterior border, weakly bisinuate, with very short median lobe before scutellum, finely margined; anterior angles obtuse, slightly produced forward, with trichobothria displaced towards anterior border bearing long anterior setae; posterior angles obtuse, with trichobothria at angle and long obliquely, posteriorly and dorsally bent golden setae; sides more or less regularly curved, widest at middle, with narrow explanate margin; surface glabrous, shiny, very finely microsculptured, slightly irregular, with dense, deep, slightly elongate punctures, larger than distance separating them and sometimes confluent, forming irregular strigose areas on disc. Hypomera smooth, very finely shagreened, with few scattered punctures near basal border; anterior border straight. Anterior border of prosternum concave, very narrow before procoxae; prosternal process about half as wide as transverse diameter of procoxae, transversely expanded at apex to enclose procoxae posteriorly, finely rugose, with relatively long, erect fine whitish hairs. Mesoventrite as wide as prosternal process basally, shortly spatulate at apex, glossy with fine punctures and fine short whitish hairs. Metaventrite slightly longer than mesoventral process, glossy and glabrous on disc, with sparse punctures basally and deep median longitudinal notch at posterior border; sides moderately rugose-punctate, with relatively dense short appressed fine whitish hairs. Scutellum arched, slightly longer than wide at base, round at apex, glossy with few scattered punctures on disc. Elytra long, slightly wider at base than base of pronotum, gently curved at sides, broadest at middle and regularly curved at apex; elytra with very weak transverse impression at basal 1 / 3; humeri strongly marked, concealing elytral margin from above, unpunctured; eight regular striae on each elytron, with only striae 3 – 5 slightly disarranged at basal third, impressed with longitudinally conjoined slit-like small punctation mostly on disc and inner striae or dot-like deep punctures elsewhere, larger in posthumeral area; additional short scutellar stria of small punctures, double basally, reaching basal 1 / 3 of elytra and short subhumeral dotted, curved stria joining marginal bead of punctures before middle of elytron; intervals weakly convex basally and on disc, more convex at apical and lateral declivities of elytra; intervals glossy, with very fine, scattered dot-like punctures. Femora dilated at middle; tibiae weakly curved, shorter than corresponding femur; tarsi shorter than corresponding tibia, with first pro- and mesotarsomeres strongly dilated in males; claws appendiculate. Abdominal ventrites finely shagreened; basal process and sides of first ventrite with fine punctures and sparse fine whitish hairs; ventrites 2 – 4 with sparse short whitish hairs at apical half; last visible ventrite emarginate at apical border with sparse erect fine setae close to border. Penis (Fig. 2 f) proportionally small, narrow and slender, strongly curved ventrally at base, nearly straight at middle, tapering and slightly curved ventrally at apex, with tip pointing ventrally; sides subparallel in dorso-ventral view, widened preapically at level with operculum, smoothly curved and slightly elongate and round at apex. Females (Fig. 4 d): The metallic shine of females is bronzy instead of green, and besides the lack of tarsal modifications, they also have a slightly heavier built, the last visible ventrite of their abdomen is entire and, as most females of Kumatoeides gen. nov., they have the intervals in lateral declivities of elytra more strongly raised. Spermatheca (Fig. 5 d) weakly sigmoidal; cornu bulbous, longer than wide, slightly strangled at base, connected to nodulus, twice as long as cornu, strongly narrowed at middle, at level with insertion of spermathecal gland, bulbous at basal half, with spermathecal duct inserted prebasally, opposite to cornu orientation.	en	Gómez-Zurita, Jesús (2018): Description of Kumatoeides gen. nov. (Coleoptera: Chrysomelidae, Eumolpinae) from New Caledonia. Zootaxa 4521 (1): 89-115, DOI: 10.11646/zootaxa.4521.1.4
03C587B7FF886969C5B9C5B0FD7CFBED.taxon	diagnosis	Diagnosis: This is the only species of Kumatoeides gen. nov. with dorsal metallic surfaces. A similar species, and possibly the closest known relative, K. anomala sp. nov., which is also found in the Mt. Mandjélia, has a hint of metallic shine on head and pronotum, but otherwise it is perfectly recognizable by the scrambled punctation basally on elytra. The two species have very similar spermathecae, more stylized in the case of K. metallica sp. nov.	en	Gómez-Zurita, Jesús (2018): Description of Kumatoeides gen. nov. (Coleoptera: Chrysomelidae, Eumolpinae) from New Caledonia. Zootaxa 4521 (1): 89-115, DOI: 10.11646/zootaxa.4521.1.4
03C587B7FF886969C5B9C5B0FD7CFBED.taxon	etymology	Derivatio nominis: The name is a feminine form for the adjectives metallicum (Latin) or µεταλλικóς (Greek), referred to the metallic appearance of dorsal integuments of the insect.	en	Gómez-Zurita, Jesús (2018): Description of Kumatoeides gen. nov. (Coleoptera: Chrysomelidae, Eumolpinae) from New Caledonia. Zootaxa 4521 (1): 89-115, DOI: 10.11646/zootaxa.4521.1.4
03C587B7FF886969C5B9C5B0FD7CFBED.taxon	distribution	Distribution: Kumatoeides metallica sp. nov. is only known from mid to high elevations in Mt. Mandjélia, on the northern part of the Massif du Panié (Fig. 3 a).	en	Gómez-Zurita, Jesús (2018): Description of Kumatoeides gen. nov. (Coleoptera: Chrysomelidae, Eumolpinae) from New Caledonia. Zootaxa 4521 (1): 89-115, DOI: 10.11646/zootaxa.4521.1.4
03C587B7FF97696BC5B9C1E8FD35FC59.taxon	description	(Figs 1 c, 2 g, 3 b, 4 f, 5 c)	en	Gómez-Zurita, Jesús (2018): Description of Kumatoeides gen. nov. (Coleoptera: Chrysomelidae, Eumolpinae) from New Caledonia. Zootaxa 4521 (1): 89-115, DOI: 10.11646/zootaxa.4521.1.4
03C587B7FF97696BC5B9C1E8FD35FC59.taxon	materials_examined	Holotype: Male, New Caledonia (S), - 21.5962 165.7765, Parc des Grandes Fougères, N of Pic Vincent, 650 m, night, 5. xi. 2010, leg. M. Wanat & R. Ruta, HOLOTYPE Kumatoeides millei sp. nov. J. Gómez-Zurita det. [red label] (MNHN). Paratype: Female, New Caledonia (S), - 21.5962 165.7765, Parc des Grandes Fougères, N of Pic Vincent, 650 m, night, 5. xi. 2010, leg. M. Wanat & R. Ruta, PARATYPE Kumatoeides millei sp. nov. J. Gómez- Zurita det. [red label] (MNHW). Other material examined: (1) one male, New Caledonia (S), 21 ° 37 ' S 165 ° 49 ' E, Col d'Amieu (top of hill), 450 m, ad lucem, 9. ii. 2004, leg. M. Wanat, Kumatoeides millei m. J. Gómez-Zurita det. 2018 (MNHW); (2) one female, IBE-JGZ- 3976, New Caledonia (S), - 21.58383 165.79667, Col d'Amieu, 450 m (2.5 km to gate), 16. xi. 2008, leg. M. Wanat, Kumatoeides millei m. J. Gómez-Zurita det. 2018 (IBE-JGZ); (3) three males, five females, New Caledonia (S), - 21.61176 165.75406, Farino, Parc des Grandes Fougères, 400 m, Camp de la Houe, 13. xi. 2008, leg. M. Wanat, Kumatoeides millei m. J. Gómez-Zurita det. 2018 (MNHW); (4) one female, New Caledonia (S), - 21.58536 165.79319, Col d'Amieu (3.0 km to gate), 500 m, at light, 14. xi. 2008, leg. M. Wanat, Kumatoeides millei m. J. Gómez-Zurita det. 2018 (MNHW); (5) one male, three females, New Caledonia (S), - 21.61859 165.75570, Farino, Parc des Grandes Fougères, 400 m, Aire des Araucarias, 14. xi. 2008, leg. M. Wanat, Kumatoeides millei m. J. Gómez-Zurita det. 2018 (MNHW); (6) one male, three females, New Caledonia (S), - 21.60948 165.77459, Farino, Parc des Grandes Fougères, 600 – 670 m, Pic Vincent track, 17. xi. 2008, leg. M. Wanat, Kumatoeides millei m. J. Gómez-Zurita det. 2018 (MNHW); (7) one male, three females, New Caledonia (S), - 21.59929 165.77519, Farino, Parc des Grandes Fougères, 670 m, track & forest N of Pic Vincent, 17. xi. 2008, leg. M. Wanat, Kumatoeides millei m. J. Gómez-Zurita det. 2018 (MNHW); (8) one female, New Caledonia (S), - 21.59929 165.77519, Farino, Parc des Grandes Fougères, 670 m, track & forest N of Pic Vincent, 17. xi. 2008, leg. M. Wanat, Kumatoeides millei m. J. Gómez-Zurita det. 2018 (IBE-JGZ); (9) one male, one female, New Caledonia (S), - 21.6006 165.7749, Parc des Grandes Fougères, N of Pic Vincent, 680 m, at light, 5. xi. 2010, leg. M. Wanat & R. Ruta, Kumatoeides millei m. J. Gómez-Zurita det. 2018 (MNHW); (10) one female, New Caledonia (S), - 21.6152 165.7736, Parc des Grandes Fougères, Aire des Carpolepis, 550 m, 5. xi. 2010, leg. M. Wanat & R. Ruta, Kumatoeides millei m. J. Gómez-Zurita det. 2018 (MNHW); (11) one male, New Caledonia (S), - 21.6152 165.7736, Parc des Grandes Fougères, Aire d Carpolepis, 550 m, at light, 6. xi. 2010, leg. M. Wanat & R. Ruta, Kumatoeides millei m. J. Gómez-Zurita det. 2018 (MNHW); (12) one male, New Caledonia (S), - 21.5918 165.7745, Parc des Grandes Fougères, N of Pic Vincent, 600 – 680 m, 6. xi. 2010, leg. M. Wanat & R. Ruta, Kumatoeides millei m. J. Gómez-Zurita det. 2018 (IBE-JGZ); (13) one female, New Caledonia (S), - 21.5918 165.7745, Parc des Grandes Fougères, N of Pic Vincent, 600 – 680 m, 6. xi. 2010, leg. M. Wanat & R. Ruta, Kumatoeides millei m. J. Gómez-Zurita det. 2018 (MNHW); (14) five females, New Caledonia (S), - 21.6095 165.7747, Parc des Grandes Fougères, N of Aire des Carpolepis, 550 – 600 m, 7. xi. 2010, leg. M. Wanat & R. Ruta, Kumatoeides millei m. J. Gómez-Zurita det. 2018 (MNHW). Head, including labrum and most of mandibles, pronotum, scutellum, elytra and ventral surfaces chestnut brown, with dim darker areas at sides of pronotum, broad margin of scutellum, basal third of elytra medially, apical half of elytra on disc, coxae and thoracic ventrites. Mouth appendages, antennae and legs brownish orange, with apical end of last antennomeres darkened (Fig. 1 c). Length: 3.10 mm (range: 2.48 – 3.10 mm); width: 1.58 mm (range: 1.29 – 1.58 mm). Head hypognathous at rest; frons microreticulate, with sparse punctures concentrated behind antennal calli and near eyes; supraocular furrows accentuated, marking external border of antennal calli posteriorly. Clypeus as long as wide at anterior border, microreticulate, with sparse dot-like punctures concentrated in posterior half; anterior border entire. Labrum transverse, feebly emarginate anteriorly, microreticulate, unpunctured. Eyes large, bulging, almost entire, feebly emarginate at inner border. Last maxillary palpomere spindle-shaped. Antennae long, slender, feebly clavate beyond seventh antennomere, reaching behind middle of elytra; scape subcylindrical, weakly bent posteriorly; pedicel 0.7 x as long as scape, widest at middle; third antennomere slender, weakly clavate, nearly glabrous, slightly longer than pedicel; fourth antennomere 1.2 x longer than third, with few sparse whitish semierect hairs; fifth and sixth antennomeres subequal, 1.5 x longer than third, sparsely but more abundantly setose than fourth; antennomeres 7 – 11 slightly more dilated than basal antennomeres, granulose and hairier; seventh antennomere longest of interstitial antennomeres, 1.8 x longer than third; antennomeres 8 – 10 subequal, slightly longer than fifth; last antennomere long, conical at apical third. Pronotum transverse (1.44 x as wide as long along middle), markedly slopping toward anterior angles and weakly cordiform, widest in front of middle; anterior border nearly straight, very finely margined, with margin incrassate near and at anterior angles; anterior angles obtuse, with large trichobothria on anterior border; sides and convex posterior border finely margined; posterior angles obtuse, with large trichobothria at angle; surface nearly smooth, densely covered by slightly elongate punctures, finer on disc. Hypomera finely shagreened, with few scattered punctures; anterior border of hypomera straight. Anterior border of prosternum concave, finely margined, narrow in front of procoxae; prosternal process large, nearly as wide as transverse diameter of procoxae, expanded apically to enclose procoxae behind, glossy, with sparse disheveled long, fine translucent hairs. Metaventral process as wide as prosternal process, enlarged apically with lateral blunt lobes, convex at apex, smooth, unpunctured. Metaventrite shorter than mesoventrite, smooth, unpunctured on disc, with fine median longitudinal impression and apical median notch; sides convex, finely microreticulate, with sparse short appressed translucent hairs. Scutellum arched, as long as wide at base, with microreticulate surface, unpunctured. Elytra long, about as wide at base as base of pronotum, with humeral angles and sides weakly curved, widest at middle and regularly curved toward sutural angle at apical third; humeri marked, unpunctured, and elytral convexity weakly depressed transversally at basal third; elytra with eight regular striae of slit-like, seamed-like small punctures, larger, round and free on striae 3 – 8 basally; short subhumeral row of eight strong round punctures posteriorly joining marginal bead of punctures; short and nearly continuously impressed scutellar row, with punctures individualized only basally; elytral intervals convex, finely microreticulate, unpunctured. Femora smooth, strongly dilated, widest at middle; tibiae feebly curved, slightly shorter than corresponding femur, expanded toward apex; tarsi shorter than corresponding tibia, with first pro- and mesotarsomeres longer than wide and as long as next two segments together; claws appendiculate. Abdominal ventrites finely shagreened, with sparse fine translucent hairs in apical half; last visible ventrite emarginate at apex with small preapical impression at middle. Penis (Fig. 2 g) stout, gradually and rather regularly curved ventrally from base to apex, with tip recurved dorsally in lateral view; sides gradually narrowing at middle in dorso-ventral view, expanding toward operculum and more strongly curved to apex, with projecting blunt, as long as wide at base tooth medially; dorsal flap of operculum subquadrate, with round angles. Females (Fig. 4 f): The females of K. millei sp. nov. show some distinctive traits, such as the sides of elytra slightly rounder, more slender, if shorter antennae and tarsi, without modified first pro- and mesotarsomeres; darker in coloration, particularly on pronotum, head, scutellum and ventral surfaces. Spermatheca (Fig. 5 c) with cornu bulbous, round at apex, and nodulus slightly longer than cornu, at angle with axis of cornu and gradually narrowing basally; spermathecal duct apparently connected to base of cornu through widened sclerotized round, short funnel expansion.	en	Gómez-Zurita, Jesús (2018): Description of Kumatoeides gen. nov. (Coleoptera: Chrysomelidae, Eumolpinae) from New Caledonia. Zootaxa 4521 (1): 89-115, DOI: 10.11646/zootaxa.4521.1.4
03C587B7FF97696BC5B9C1E8FD35FC59.taxon	diagnosis	Diagnosis: Kumatoeides millei sp. nov. is most similar to K. megale sp. nov. and the males of both species can be distinguished easily based on their genitalia, the tip of which in K. millei is narrower than its base and in K. megale it is broader. K. millei is larger but similar in appearance to K. aulacia sp. nov. and K. costata (Jolivet, Verma et Mille). From the former, they can be also distinguished by their characteristic penises in the case of males and in part by coloration and the marked posthumeral costa in K. aulacia in the case of females. From the latter, there are several diagnostic differences, including the separate versus seamed punctation of the scutellar stria or the absence versus presence of supraocular furrow, apart from marked differences in both male genitalia and spermathecae.	en	Gómez-Zurita, Jesús (2018): Description of Kumatoeides gen. nov. (Coleoptera: Chrysomelidae, Eumolpinae) from New Caledonia. Zootaxa 4521 (1): 89-115, DOI: 10.11646/zootaxa.4521.1.4
03C587B7FF97696BC5B9C1E8FD35FC59.taxon	etymology	Derivatio nominis: This species is dedicated with affection to Dr. Christian Mille, researcher at the Institut Agronomique néo-Calédonien (La Foa, New Caledonia), and greatly committed to promoting and increasing the knowledge of the entomofauna of New Caledonia.	en	Gómez-Zurita, Jesús (2018): Description of Kumatoeides gen. nov. (Coleoptera: Chrysomelidae, Eumolpinae) from New Caledonia. Zootaxa 4521 (1): 89-115, DOI: 10.11646/zootaxa.4521.1.4
03C587B7FF97696BC5B9C1E8FD35FC59.taxon	distribution	Distribution: The species is present at mid elevations of the southern edges of the Massif de la Boghen in the Central Range of the Grande Terre (Fig. 3 b).	en	Gómez-Zurita, Jesús (2018): Description of Kumatoeides gen. nov. (Coleoptera: Chrysomelidae, Eumolpinae) from New Caledonia. Zootaxa 4521 (1): 89-115, DOI: 10.11646/zootaxa.4521.1.4
03C587B7FF95696DC5B9C13BFBC9FDF1.taxon	description	(Figs 2 h, 3 c, 4 g, 5 f, 6 b)	en	Gómez-Zurita, Jesús (2018): Description of Kumatoeides gen. nov. (Coleoptera: Chrysomelidae, Eumolpinae) from New Caledonia. Zootaxa 4521 (1): 89-115, DOI: 10.11646/zootaxa.4521.1.4
03C587B7FF95696DC5B9C13BFBC9FDF1.taxon	materials_examined	Holotype: Male, New Caledonia (S), Mt. Koghi 22 ° 10.7 ' S 166 ° 30.4 ' E, 450 – 500 m, rainforest, 16. xii. 2006, leg. M. Wanat & R. Dobosz, HOLOTYPE Kumatoeides tarsalis sp. nov. J. Gómez-Zurita det. [red label] (MNHN). Paratypes: Ten males and three females, New Caledonia (S), Mt. Koghi 22 ° 10.7 ' S 166 ° 30.4 ' E, 450 – 500 m, rainforest, 16. xii. 2006, leg. M. Wanat & R. Dobosz, PARATYPE Kumatoeides tarsalis sp. nov. J. Gómez-Zurita det. [red label] (nine males and two females at MNHW, one male and one female at IBE-JGZ). Other material examined: (1) four males and three females, New Caledonia (S), 22 ° 11 ' S 166 ° 30 ' E, Mts. Koghi, 450 – 600 m, track to Cascade, forest, 21. i. 2004, leg. M. Wanat, Kumatoeides tarsalis m. J. Gómez-Zurita det. 2018 (MNHW); (2) four males and seven females, New Caledonia (S), 22 ° 11 ' S 166 ° 30 ' E, Mts. Koghi, 500 – 550 m, humid forest, 21. i. 2004, leg. M. Wanat, Kumatoeides tarsalis m. J. Gómez-Zurita det. 2018 (MNHW); (3) one female, New Caledonia (S), 22 ° 11 ' S 166 ° 30 ' E, Mts. Koghi, 450 – 600 m, humid forest, 11. ii. 2004, leg. M. Wanat, Kumatoeides tarsalis m. J. Gómez-Zurita det. 2018 (MNHW); (4) one male, New Caledonia (S), 22 ° 11 ' S 166 ° 30 ' E, Mts. Koghi, 450 – 600 m, humid forest, 12. ii. 2004, leg. M. Wanat, Kumatoeides tarsalis m. J. Gómez-Zurita det. 2018 (MNHW); (5) four males, one female, New Caledonia (S), - 22.17809 166.50569, Mts. Koghi, 470 – 500 m, Auberge & forest, 24. x. 2008, leg. M. Wanat, Kumatoeides tarsalis m. J. Gómez-Zurita det. 2018 (MNHW); (6) one male, New Caledonia (S), - 22.17809 166.50569, Koghi Mts., 500 – 550 m, rainforest, 27. x. 2008, leg. M. Wanat, Kumatoeides tarsalis m. J. Gómez-Zurita det. 2018 (MNHW); (7) four males and four females, New Caledonia (S), - 22.17809 166.50569, Koghi Mts., 500 – 550 m, Auberge to Vallée des Houps, 3. xii. 2008, leg. M. Wanat, Kumatoeides tarsalis m. J. Gómez-Zurita det. 2018 (MNHW); (8) one male, IBE-JGZ- 3977, New Caledonia (S), - 22.17809 166.50569, Mts. Koghi, 500 – 550 m, Auberge to Vallée des Houps, 3. xii. 2008, leg. M. Wanat, Kumatoeides tarsalis m. J. Gómez-Zurita det. 2018 (IBE-JGZ); (9) seven males and six females, New Caledonia (S), - 22.18177 166.50936, Upper Mts. Koghi, Vallée des Houps, 520 – 550 m, 3. xii. 2008, leg. M. Wanat, Kumatoeides tarsalis m. J. Gómez-Zurita det. 2018 (MNHW); (10) one male and one female, New Caledonia (S), - 22.18177 166.50936, Upper Mts. Koghi, Vallée des Houps, 520 – 550 m, 3. xii. 2008, leg. M. Wanat, Kumatoeides tarsalis m. J. Gómez-Zurita det. 2018 (IBE-JGZ); (11) two males, New Caledonia (S), - 22.18038 166.51313, Upper Mts. Koghi, 600 – 700 m, vicinity of Pic Malaoui track junction, 3. xii. 2008, leg. M. Wanat, Kumatoeides tarsalis m. J. Gómez-Zurita det. 2018 (MNHW); (12) six males and two females, New Caledonia (S), - 22.17809 166.50569, Mts. Koghi, 500 – 550 m., track to Cascade, forest, 4. xii. 2008, leg. M. Wanat, Kumatoeides tarsalis m. J. Gómez-Zurita det. 2018 (MNHW). Pronotum, scutellum, ventral thoracic parts, trochanters and elytral punctures chestnut brown; head and elytra slightly paler; palpi, antennae and legs yellowish orange, with five apical antennomeres, base of tibiae and onychia slightly darkened (Fig. 6 b). Length: 2.74 mm (range: 2.38 – 3.10 mm); width: 1.38 mm (range: 1.18 – 1.59 mm). Head hypognathous at rest; frons and vertex regularly convex, very finely microsculptured, glossy, with obsolete median longitudinal furrow and sparsely but neatly punctured, except at middle, sulcate above eyes with sulcus gently curved and weakly impressed above supra-antennal swellings; glabrous except for long seta at middle of supraocular furrow and sparse short, very fine appressed yellowish setae near and on short genae. Clypeus as long as wide at apex, smooth, glossy, with small punctures at base and along median line. Labrum small, smooth, about 1.5 x as wide as long, emarginate at apex. Apical maxillary palpomere spindle-shaped, obliquely cut shortly at apex. Eyes large, bulging, weakly emarginate at inner border. Antennae slender, reaching beyond middle of elytra; scape nearly cylindrical, slightly bent posteriorly, twice as long as thick, strongly narrowed at base; pedicel cup-shaped, half as long as scape; antennomeres 3 – 10 club-shaped, 3 – 6 glossy with sparse long setae, 7 – 11 feebly incrassate, finely granulose, clothed with dense, short hairs; antennomeres 3 – 4 subequal, slightly longer than pedicel; fifth antennomere longer than third and sixth; seventh anntenomere longer than fifth; antennomeres 8 – 10 subequal, as long as sixth; eleventh antennomere as long as seventh, pointed, obliquely cut at apex internally. Pronotum transverse (1.46 x as wide at base than long at middle), feebly convex with stronger slopes toward anterior angles; anterior border produced forward, straight, finely margined near anterior angles, narrower than posterior border, feebly bisinuate, with very short median lobe before scutellum, finely margined; anterior angles obtuse, blunt, explanate, with trichobothria on anterior border and setae directed forward; posterior angles obtuse, with trichobothria at angle and setae oriented laterally; sides very feebly sinuate at basal half, widest after middle, with fine explanate margin; surface glabrous, shiny, very finely microsculptured, with dense, small elongate punctures smaller than distance separating them. Hypomera finely microreticulate, smooth, unpunctured; very feebly convex at anterior border. Prosternal process broad, about 0.8 x as wide as transverse diameter of procoxae, expanded laterally at apex to enclose procoxae posteriorly, smooth with scattered disheveled long fine translucent hairs. Mesoventral process as broad as prosternal process, short, transversally expanded at apex as two short round lobes, convex at apical border. Metaventrite as long as mesoventrite, smooth on disc, with very fine longitudinally impressed suture and deep longitudinal notch at middle of posterior border; sides finely microreticulate, with scattered short appressed translucent setae. Scutellum ogival, slightly longer than wide at base, very finely microreticulate. Elytra long, basally as wide as base of pronotum, gently curved at sides, widest at middle and regularly curved at apex; elytra with very weak transverse impression at basal 1 / 3; humeri strongly marked, unpunctured; eight complete regular striae on each elytron with longitudinal series of dot- or slit-like small punctation or dot-like individual, large punctures in five outer striae basally; additional short scutellar stria reaching transversely depressed area and short subhumeral dotted stria joining premarginal bead of punctures before middle of elytron; intervals nearly flat basally and on disc, outer two intervals slightly more convex; intervals unpunctured, very finely microreticulate. Femora strongly inflated medially; tibiae slightly shorter than corresponding femur; tarsi shorter than corresponding tibia, with first pro- and mesotarsomeres strongly widened; claws appendiculate. Abdominal ventrites finely shagreened with scattered fine translucent hairs; last visible abdominal ventrite shortly emarginate at apex and weakly impressed longitudinally at middle. Penis (Fig. 2 h) slender, regularly curved ventrally from base to apex in lateral view, tapering dorsally at apex from base of operculum, with tip slightly curved dorsally; sides weakly narrowing at middle in dorso-ventral view and widened at level with operculum, following its oval contour and expanding again at apex as relatively narrow and long axehead. Females (Fig. 4 g): The females of Kumatoeides tarsalis sp. nov. are very similar to males in shape and size, but they can be distinguished by the lack of tarsal modification in fore and mid legs, by thinner antennae, more accentuated elytral intervals in lateral declivities of elytra, and also by the darker color of their bodies, particularly on head, pronotum, along elytral suture and sides of elytra (not so much the appendages). Spermatheca (Fig. 5 f) very small, shaped like question mark, with cornu elongate, curved, narrowed at base toward narrower nodulus; nodulus curved at middle and twisted at base, with spermathecal gland insertion after median curvature; spermathecal duct appears as large, tubular, slightly sclerotized section before membranous part.	en	Gómez-Zurita, Jesús (2018): Description of Kumatoeides gen. nov. (Coleoptera: Chrysomelidae, Eumolpinae) from New Caledonia. Zootaxa 4521 (1): 89-115, DOI: 10.11646/zootaxa.4521.1.4
03C587B7FF95696DC5B9C13BFBC9FDF1.taxon	diagnosis	Diagnosis: From most other species of Kumatoeides gen. nov., K. tarsalis sp. nov. can be distinguished by the more spaced punctation on pronotum, which is dense, sometimes confluent, locally vermiculate in most other species. At present, only two other species in the genus have similar pronotal punctation: K. leptalei sp. nov. and K. wanati sp. nov. However, the former is much smaller (ca. 2.3 mm), much paler, and lacks any sign of transverse impression on the elytra, and the latter is also paler and males have proportionally smaller first pro- and mesotarsomeres. The males of the three species have very distinctive penises, and the spermathecae of these species are very different too.	en	Gómez-Zurita, Jesús (2018): Description of Kumatoeides gen. nov. (Coleoptera: Chrysomelidae, Eumolpinae) from New Caledonia. Zootaxa 4521 (1): 89-115, DOI: 10.11646/zootaxa.4521.1.4
03C587B7FF95696DC5B9C13BFBC9FDF1.taxon	etymology	Derivatio nominis: The males of all known species of the genus have strongly dilated first fore and mid tarsomeres. This trait is nonetheless giving name to this species where this feature is highly prominent, and it corresponds to the Latin adjective (fem., sing.) of the term for these foot articulations, tarsus (Latin) or ταρσóς (Greek).	en	Gómez-Zurita, Jesús (2018): Description of Kumatoeides gen. nov. (Coleoptera: Chrysomelidae, Eumolpinae) from New Caledonia. Zootaxa 4521 (1): 89-115, DOI: 10.11646/zootaxa.4521.1.4
03C587B7FF95696DC5B9C13BFBC9FDF1.taxon	distribution	Distribution: Kumatoeides tarsalis sp. nov. has been collected in several localities in the Massif des Koghi, in the southern edge of the large Massif du Sud, not far from the capital, Nouméa (Fig. 3 c).	en	Gómez-Zurita, Jesús (2018): Description of Kumatoeides gen. nov. (Coleoptera: Chrysomelidae, Eumolpinae) from New Caledonia. Zootaxa 4521 (1): 89-115, DOI: 10.11646/zootaxa.4521.1.4
03C587B7FF926961C5B9C5B0FCA5FDF1.taxon	description	(Figs 2 i, 3 b, 4 h, 5 g, 6 c)	en	Gómez-Zurita, Jesús (2018): Description of Kumatoeides gen. nov. (Coleoptera: Chrysomelidae, Eumolpinae) from New Caledonia. Zootaxa 4521 (1): 89-115, DOI: 10.11646/zootaxa.4521.1.4
03C587B7FF926961C5B9C5B0FCA5FDF1.taxon	materials_examined	Other material examined: (1) two males and one female, New Caledonia (N), - 20.9510 165.2912, Pic d'Amoa (Povila [Poindinié County]), 400 m, lower rainforest, 18. xi. 2010, leg. M. Wanat & R. Ruta, Kumatoeides wanati m. J. Gómez-Zurita det. 2018 (MNHW); (2) one female, New Caledonia (N), - 21.1489 165.3235, Aoupinié (refuge), 400 m, at light, 19. xi. 2010, leg. M. Wanat & R. Ruta, Kumatoeides wanati m. J. Gómez-Zurita det. 2018 (MNHW); (3) one male and one female, New Caledonia (N), - 21.1489 165.3235, Aoupinié (refuge), 400 m, at light, 22. xi. 2010, leg. M. Wanat & R. Ruta, Kumatoeides wanati m. J. Gómez-Zurita det. 2018 (MNHW); (4) seven males and six females, New Caledonia (N), - 21.1489 165.3235, Aoupinié (refuge) [Ponérihouen County], 400 m, at light, 25. xi. 2010, leg. M. Wanat, Kumatoeides wanati m. J. Gómez-Zurita det. 2018 (MNHW); (5) two females, New Caledonia (N), - 20.95280 165.29135, Pic d'Amoa (Povila), 400 m, rainforest, 22. xi. 2008, leg. M. Wanat, Kumatoeides wanati m. J. Gómez-Zurita det. 2018 (MNHW); (6) one female, New Caledonia (N), - 21.14890 165.32348, Aoupinié (refuge), 400 m, at light, 25. xi. 2008, leg. M. Wanat, Kumatoeides wanati m. J. Gómez-Zurita det. 2018 (MNHW); (7) one male and one female, New Caledonia (N), - 21.14890 165.32348, Aoupinié (refuge), 400 m, at light, 26. xi. 2008, leg. M. Wanat, Kumatoeides wanati m. J. Gómez-Zurita det. 2018 (MNHW); (8) two females, IBE-JGZ- 3973, New Caledonia (N), - 21.14890 165.32348, Aoupinié (refuge), 400 m, at light, 26. xi. 2008, leg. M. Wanat, Kumatoeides wanati m. J. Gómez-Zurita det. 2018 (IBE-JGZ). Head, including labrum, pronotum, scutellum and ventral surfaces brownish orange; antennae, mouth appendages, elytra and legs pale orange; apical half of last antennomere dark brown (Fig. 6 c). Length: 2.54 mm (range: 2.41 – 3.10 mm); width: 1.36 mm (range: 1.28 – 1.59 mm). Head hypognathous at rest; vertex and frons regularly convex, finely microreticulate, relatively densely covered by fine punctation; supraocular sulci extended above supraantennal calli. Clypeus short, wider at emarginate anterior border than long at middle, finely microreticulate with sparse, fine punctures. Labrum subquadrate, weakly emarginate at apex, microreticulate, unpunctured. Eyes large, bulging, nearly entire, feebly emarginate at inner border. Last maxillary palpomere spindle-shaped, obliquely cut at apex. Antennae slender, reaching middle of elytra; scape subcylindrical, slightly bent backward; pedicel short, subglobose; third antennomere club-shaped, slightly longer than pedicel; fourth antennomere shortly clavate, as long as pedicel; fifth antennomere clavate, 1.4 x longer than third, with sparse, semierect whitish setae; sixth antennomere 1.2 x longer than third, hairier than fifth; antennomeres 7 – 11 weakly enlarged, granulose, pubescent; seventh antennomere clavate, nearly 1.8 x longer than third; antennomeres 8 – 10 subequal, about 1.6 x longer than third; last antennomere acuminate at apical third. Pronotum transverse (1.52 x as wide at base as long along middle); sides round, widest at middle, with slightly explanate margin, larger at anterior, widely obtuse angles, and feebly notched before obtuse posterior angles; anterior border straight, unmargined, with large trichobothria at ends, nearly on angles; posterior border regularly convex, finely margined, with trichobothria at both ends; surface finely microreticulate, covered by fine punctures, denser on disc than at sides, with interspaces larger than diameter of punctures. Hypomera finely microreticulate, unpunctured, with anterior border straight. Prosternum narrow anteriorly; prosternal process about 0.75 x as wide as transverse diameter of procoxae, smooth, glossy, with sparse, relatively long, disheveled fine yellowish hairs, transversely expanded at apex to enclose procoxae posteriorly. Process of mesoventrite as wide as prosternal process, glossy, unpunctured, shortly expanded at apex, posterior border convex. Metaventrite slightly shorter than mesoventrite, flat and shiny on disc, with finely impressed median suture and sharp apical median notch; sides finely shagreened with sparse, short, posteriorly recumbent fine yellowish setae. Scutellum as long as wide at base, regularly round at apex, finely microreticulate, unpunctured. Elytra long, regularly convex, as wide at base as pronotum basally; humeral angles round and sides feebly curved, widest slightly before middle; humeri marked, unpunctured; surface microreticulation subtler than on pronotum, with eight regular striae of fine punctures, slit-like at apical half of elytra and slightly elongate on striae 1 – 5 basally, larger and round on striae 6 – 8 basally and on short subhumeral striae, connected to elytral margin before middle of elytron; short scutellar striae of 11 – 12 small and occasionally geminate punctures present; interstrices nearly flat on disc basally, weakly convex at sides and apex of elytra. Femora enlarged at middle; tibiae shorter than corresponding femur and tarsi shorter than corresponding tibia; first pro- and mesotarsomeres short but conspicuously enlarged in males. Abdominal ventrites finely shagreened, unpunctured, with sparse yellowish hairs, finer than on metaventrite; last visible ventrite emarginate at apex, with short median longitudinal impression. Penis (Fig. 2 i) slender, in lateral view more strongly curved at both ends than at middle, imperceptibly narrowing from base to apex, tapering at distal end beyond operculum; sides slightly concave in dorso-ventral view, widening and following ovoid contour of operculum preapically; apical border with long and narrow median projection, feebly dilated at apex. Females (Fig. 4 h): The females of this species are very similar to males in shape and coloration, although often with scutellum, elytral suture basally, and sometimes disc of pronotum darkened, and can be recognized externally by lack of modifications in first pro- and mesotarsomeres, as well as the entire posterior border of the last visible abdominal ventrite and raised elytral intervals at sides of elytra behind humeri. Spermatheca (Fig. 5 g) with kidney shaped cornu and long, narrow and cylindrical nodulus with pre-median insertion of spermathecal gland.	en	Gómez-Zurita, Jesús (2018): Description of Kumatoeides gen. nov. (Coleoptera: Chrysomelidae, Eumolpinae) from New Caledonia. Zootaxa 4521 (1): 89-115, DOI: 10.11646/zootaxa.4521.1.4
03C587B7FF926961C5B9C5B0FCA5FDF1.taxon	diagnosis	Diagnosis: Kumatoeides wanati sp. nov. is similar to K. tarsalis sp. nov., but it is paler, except for the apical half of the last antennomere, which is conspicuously darkened. Both the female and the male genitalia of the two species are completely different too.	en	Gómez-Zurita, Jesús (2018): Description of Kumatoeides gen. nov. (Coleoptera: Chrysomelidae, Eumolpinae) from New Caledonia. Zootaxa 4521 (1): 89-115, DOI: 10.11646/zootaxa.4521.1.4
03C587B7FF926961C5B9C5B0FCA5FDF1.taxon	etymology	Derivatio nominis: This species is dedicated with gratitude to Dr. Marek Wanat (Muzeum Przyrodnicze, Uniwersytet Wrocławski, Wroclaw, Poland) for generously confiding me the study of the New Caledonian Eumolpinae collected in his several campaigns in the archipelago.	en	Gómez-Zurita, Jesús (2018): Description of Kumatoeides gen. nov. (Coleoptera: Chrysomelidae, Eumolpinae) from New Caledonia. Zootaxa 4521 (1): 89-115, DOI: 10.11646/zootaxa.4521.1.4
03C587B7FF926961C5B9C5B0FCA5FDF1.taxon	distribution	Distribution: The species appears to have a relatively ample distribution in the eastern slopes of the Central Range, from near Poindimié to Canala counties (Fig. 3 b).	en	Gómez-Zurita, Jesús (2018): Description of Kumatoeides gen. nov. (Coleoptera: Chrysomelidae, Eumolpinae) from New Caledonia. Zootaxa 4521 (1): 89-115, DOI: 10.11646/zootaxa.4521.1.4
