taxonID	type	description	language	source
03C2878BFFD0FFD4F3D4D2AC371FFA73.taxon	description	brevicornis Emery 1894 a. Canindeyú (ALWC, INBP, LACM, MHNG). Literature records: Canindeyú (MacKay 2004). kirbyi Emery 1895 b. “ Paraguay ” (s. loc.) (MCSN). Literature records: “ Paraguay ” (s. loc.) (Emery 1895 b). laticornis Forel 1908 b. Cordillera (MCSN, MHNG, NHMB). Literature records: Cordillera, “ Paraguay ” (s. loc.) (Forel 1908 b, Forel 1911 a, MacKay 1996, Santschi 1916). longinodis MacKay 2004. Alto Paraguay (INBP, MCZC, WPMC). Literature records: Alto Paraguay (MacKay 2004). serratulus (F. Smith 1858). Literature records: Cordillera, “ Paraguay ” (s. loc.) (Emery 1894 d, Forel 1895, Forel 1906).	en	Wild, Alexander L. (2007): A catalogue of the ants of Paraguay (Hymenoptera: Formicidae). Zootaxa 1622: 1-55, DOI: 10.5281/zenodo.179222
03C2878BFFD0FFD4F3D4D11C35BAF8C3.taxon	description	bruchi Santschi 1936. Literature records: Pte. Hayes (Fowler 1981).	en	Wild, Alexander L. (2007): A catalogue of the ants of Paraguay (Hymenoptera: Formicidae). Zootaxa 1622: 1-55, DOI: 10.5281/zenodo.179222
03C2878BFFD0FFD4F3D4D3D43455F99B.taxon	description	sp. alw- 01. Canindeyú (ALWC, INBP).	en	Wild, Alexander L. (2007): A catalogue of the ants of Paraguay (Hymenoptera: Formicidae). Zootaxa 1622: 1-55, DOI: 10.5281/zenodo.179222
03C2878BFFD0FFD4F3D4D076375CF915.taxon	description	brasiliensis Emery 1901. Paraguarí, San Pedro (ALWC, PSWC). Literature records: Cordillera (de Andrade 1998, Forel 1906).	en	Wild, Alexander L. (2007): A catalogue of the ants of Paraguay (Hymenoptera: Formicidae). Zootaxa 1622: 1-55, DOI: 10.5281/zenodo.179222
03C2878BFFD0FFD5F3D4D1BC3780FE7D.taxon	description	adrepens Forel 1911 b. Canindeyú, Cordillera, San Pedro (ALWC, IFML, INBP, LACM, MHNG, NHMB). Literature records: Cordillera (Forel 1911 b). alfari Emery 1893 a. Alto Paraná, Caaguazú, Canindeyú, Central, Cordillera, Itapúa, Ñeembucú, “ Paraguay ” (s. loc.) (ALWC, IFML, INBP, LACM, MCSN, MCZC, MHNG). Literature records: Alto Paraná, Cordillera, “ Paraguay ” (s. loc.) (Emery 1893 l, Forel 1908 b, Longino 1989). leuderwaldti Forel 1909. Central, Cordillera, “ Paraná R. ” (Dept. unknown) (ALWC, MCZC, MHNG, NHMB). Literature records: Cordillera (Forel 1909). schimperi Emery 1893 b. Cordillera, “ Paraná R. ” (Dept. unknown) (LACM, MCZC, MHNG, MZSP, NHMB). Literature records: Central, Cordillera, “ Paraguay ” (s. loc.) (Forel 1909 [as “ fiebrigi ”], Fowler 1981 [as “ clariceps ”], Longino 2007, Santschi 1916 [as “ fiebrigi ”]). sp. alw- 01. Alto Paraná, Amambay, Canindeyú, Central, Ñeembucú, Pte. Hayes (ALWC, IFML, INBP, LACM, MZSP, NHMB).	en	Wild, Alexander L. (2007): A catalogue of the ants of Paraguay (Hymenoptera: Formicidae). Zootaxa 1622: 1-55, DOI: 10.5281/zenodo.179222
03C2878BFFD1FFD5F3D4D7DE37D3FC9D.taxon	description	bispinosus (Olivier 1792). Alto Paraná, Canindeyú, Central, Concepción, Cordillera, Guairá, Ñeembucú, San Pedro, “ Paraná R. ” (Dept. unkown), “ Paraguay ” (s. loc.) (ALWC, IFML, INBP, LACM, MCZC, MZSP, NHMB, NHMW). Literature records: Central, Cordillera (Emery 1906, Forel 1909). germaini Emery 1894 a. Alto Paraguay, Alto Paraná, Amambay, Caaguazú, Canindeyú, Central, Concepción, Cordillera, “ Paraguay ” (s. loc.) (ALWC, BMNH, IFML, INBP, LACM, MCSN, MCZC, MZSP). Literature records: “ Paraguay ” (s. loc.) (Emery 1896 b). lamellosus (Mayr 1870 a). Alto Paraná, Caaguazú, Canindeyú, Central (ALWC, INBP, LACM). lutosus (F. Smith 1858). Canindeyú, Cordillera, San Pedro (ALWC, INBP, LACM). Literature records: “ Paraguay ” (s. loc.) (Emery 1894 b).	en	Wild, Alexander L. (2007): A catalogue of the ants of Paraguay (Hymenoptera: Formicidae). Zootaxa 1622: 1-55, DOI: 10.5281/zenodo.179222
03C2878BFFD1FFD5F3D4D5713439FAFD.taxon	description	bituber Santschi 1916. Boquerón (ALWC, IFML). brunneus Forel 1908 b. Caagauzú, Canindeyú, Central, Cordillera, Guairá, Ñeembucú, Paraguarí, San Pedro, “ Chaco ” (Dept. unknown) (ALWC, IFML, INBP, LACM, MZSP, NHMB, NHMW). Literature records: Cordillera, “ Paraguay ” (s. loc.) (Emery 1913 a, Forel 1911 a). exsanguis Forel 1912 d. Boquerón (ALWC, IFML). jheringi Forel 1912 d. Canindeyú (ALWC, IFML). paranensis Santschi 1922 a. Amambay, Caaguazú, Central, Concepción, Cordillera, Paraguarí (ALWC, IFML, INBP, LACM, MHNG, MZSP, NHMB). Literature records: Central, Cordillera, “ Paraguay ” (s. loc.) (Emery 1913 a, Forel 1911 b, Santschi 1922 a). NEW STATUS. spurius Santschi 1929 b. Alto Paraná, Amambay, Boquerón, Caaguazú, Canindeyú, Central, Concepción, Cordillera, Guairá, Paraguarí, Pte. Hayes, San Pedro (ALWC, IFML, INBP, LACM, MHNG). Literature records: Cordillera, “ Paraguay ” (s. loc.) (Emery 1913 a, Forel 1911 b, Kempf 1975 c). thoracicus Gallardo 1916. Boquerón, Canindeyú, Central, Misiones, Ñeembucú, Paraguarí, Pte. Hayes, San Pedro (ALWC, IFML, INBP, LACM). sp. alw- 03. Boquerón (ALWC).	en	Wild, Alexander L. (2007): A catalogue of the ants of Paraguay (Hymenoptera: Formicidae). Zootaxa 1622: 1-55, DOI: 10.5281/zenodo.179222
03C2878BFFD1FFD5F3D4D093322DF8BE.taxon	description	pombero Wild and Cuezzo 2006. Pte. Hayes (ALWC, BMNH, CASC, IFML, LACM, MCZC, MHNG, MZSP, UCDC, USNM). Literature records: Pte. Hayes (Wild and Cuezzo 2006).	en	Wild, Alexander L. (2007): A catalogue of the ants of Paraguay (Hymenoptera: Formicidae). Zootaxa 1622: 1-55, DOI: 10.5281/zenodo.179222
03C2878BFFD1FFDAF3D4D1133502FDDD.taxon	description	aztecoides Wild 2007. Canindeyú, Cordillera (ALWC, BMNH, CASC, IFML, INBP JTLC, LACM, MCZC, MHNG, MZSP, NHMB, NHMW, QCAZ, UCDC, USNM). Literature records: Canindeyú, Cordillera (Wild 2007). cerradense Wild 2007. Canindeyú, Central, Cordillera (ALWC, BMNH, CASC, IFML, INBP, LACM, MCZC, MHNG, MZSP, NHMB, UCDC, USNM). Literature records: Canindeyú, Central, Cordillera (Wild 2007). cryptobioticum Wild 2007. Boquerón (ALWC, BMNH, INBP, LACM, MCZC, MHNG, MZSP, USNM). Literature records: Boquerón (Wild 2007). gallardoi (Brèthes 1914). Canindeyú, Central, Guairá, Itapúa, Pte. Hayes (ALWC, BMNH, CASC, INBP, IFML, LACM, MCZC, MHNG, MZSP, UCDC, USNM). Literature records: Canindeyú, Central, Guairá, Itapúa, Pte. Hayes (Wild 2007). humile (Mayr 1868). Alto Paraguay, Boquerón, Central, Ñeembucú, Pte. Hayes, San Pedro, “ Paraná R. ” (Dept. unknown) (ALWC, BMNH, IFML, INBP, MACN, MCSN, MCZC, MHNG, MZSP, NHMB). Literature records: Alto Paraguay, Boquerón, Central, Ñeembucú, Pte. Hayes, San Pedro (Wild 2004, Wild 2007). iniquum (Mayr 1870 a). Canindeyú, Cordillera, Itapúa (ALWC, BMNH, CASC, INBP, LACM, MCZC, MZSP, UCDC). Literature records: Canindeyú, Cordillera, Itapúa (Wild 2007). micans (Forel 1908 b). Central (ALWC). Literature records: Central (Wild 2007). neotropicum Wild 2007. Alto Paraná, Canindeyú (ALWC, BMNH, CASC, IFML, INBP, JTLC, MCZC, MHNG, MZSP, NHMB, PSWC, QCAZ, UCDC, USNM, WPMC). Literature records: Alto Paraná, Canindeyú (Wild 2007). pulex Wild 2007. Canindeyú (ALWC, BMNH, CASC, IFML, LACM, MCZC, MHNG, MZSP, NHMW, UCDC, USNM). Literature records: Canindeyú (Wild 2007).	en	Wild, Alexander L. (2007): A catalogue of the ants of Paraguay (Hymenoptera: Formicidae). Zootaxa 1622: 1-55, DOI: 10.5281/zenodo.179222
03C2878BFFDEFFDAF3D4D59E3464FADD.taxon	description	burchellii (Westwood 1842). Amambay, Canindeyú, Guairá, “ Paraguay ” (s. loc.) (ALWC, BMNH, INBP, IFML, LACM, MCSN, MZSP, NHMB, NHMW). Literature records: “ Paraguay ” (s. loc.) (Borgmeier 1955, Santschi 1920 a). dulcium Forel 1912 a. Literature records: Canindeyú (Fowler 1981). mexicanum Roger 1863 a. Central, “ Paraguay ” (s. loc.) (MCSN, MHNG, MZSP, NHMB, NHMW). Literature records: Cordillera, “ Paraguay ” (s. loc.) (Borgmeier 1939, Emery 1894 a, Forel 1911 a). mexicanum argentinum Borgmeier 1955. “ San Alfredo, Paraguay ” (Dept. unknown). (MZSP). quadriglume (Haliday 1836). Literature records: Cordillera (Forel 1909). vagans (Olivier 1792). Central, Paraguarí, “ Paraná R. ” (Dept. unknown) (ALWC, INBP, LACM, MCZC). Literature records: “ Paraguay ” (s. loc.) (Borgmeier 1955, Emery 1896 b, Santschi 1920 a). vagans dubitatum Emery 1896 a. Caaguazú, Cordillera, “ Paraguay ” (s. loc) (MCSN, MHNG, MZSP, NHMB, NHMW). Literature records: “ Paraguay ” (s. loc.), “ San Alfredo, Paraguay ” (Dept. unknown). (Borgmeier 1936, Borgmeier 1955, Emery 1910).	en	Wild, Alexander L. (2007): A catalogue of the ants of Paraguay (Hymenoptera: Formicidae). Zootaxa 1622: 1-55, DOI: 10.5281/zenodo.179222
03C2878BFFDEFFDAF3D4D3BE3498F97D.taxon	description	coecus (Latreille 1802). Amambay, Boquerón, Caaguazú, Canindeyú, Central, Concepción, Cordillera, Guairá, Itapúa, Ñeembucú, San Pedro, “ Paraguay ” (s. loc.) (ALWC, IFML, INBP, LACM, MCSN, MZSP, NHMW). Literature records: “ Paraguay ” (s. loc.) (Borgmeier 1955, Emery 1894 a, Emery 1896 b, Santschi 1916, Wheeler 1921). mars (Forel 1912 a). Canindeyú (ALWC, INBP, LACM). praedator (F. Smith 1858). Alto Paraná, Canindeyú, Central, Cordillera, Pte. Hayes, San Pedro, “ Paraná R. ” (Dept. unknown), “ Paraguay ” (s. loc.) (ALWC, IFML, INBP, LACM, MCZC, NHMB, NHMW). Literature records: Central, Concepción, Cordillera, Ñeembucú, Pte. Hayes, San Pedro (Borgmeier 1955, Emery 1906, Forel 1906, Forel 1908 b, Fowler 1979, Santschi 1916).	en	Wild, Alexander L. (2007): A catalogue of the ants of Paraguay (Hymenoptera: Formicidae). Zootaxa 1622: 1-55, DOI: 10.5281/zenodo.179222
03C2878BFFDEFFDAF3D4D4BE3498FCBD.taxon	description	atriceps Emery 1888 a. Canindeyú (ALWC, IFML, INBP). * melanocephalum (Fabricius 1793). Central, Concepción (ALWC, INBP). sp. alw- 01. Canindeyú (ALWC, IFML, INBP).	en	Wild, Alexander L. (2007): A catalogue of the ants of Paraguay (Hymenoptera: Formicidae). Zootaxa 1622: 1-55, DOI: 10.5281/zenodo.179222
03C2878BFFDEFFDBF3D4D0D13295FBDD.taxon	description	angustinodis (Emery 1888 c). Canindeyú, “ Paraguay ” (s. loc.) (ALWC, INBP, LACM, MZSP). Literature records: “ Paraguay Centrale ”, “ Paraguay ” (s. loc.) (Borgmeier 1955, Emery 1894 a, Emery 1894 b). bohlsi (Emery 1896 b). Concepción (?) (MCSN, MHNG, MZSP). Literature records: Concepción (?) (Borgmeier 1955, Emery 1896 b). carettei (Forel 1913). Central (ALWC, INBP, LACM). clavifemur Borgmeier 1953. Cordillera (LACM). diversinodis (Borgmeier 1933). Central, Paraguarí (?), “ Paraguay ” (s. loc.) (ALWC, IFML, INBP, LACM, MZSP, NHMW). Literature records: Central (?) (Borgmeier 1933, Borgmeier 1955). goeldii (Forel 1901 d). Canindeyú (ALWC, INBP, LACM). halidaii (Schuckard 1840 a). Literature records: Central (Santschi 1916). hetschkoi (Mayr 1886 a). Caaguazú, Paraguarí, “ Paraguay ” (s. loc.) (BMNH, MZSP, NHMB). Literature records: “ Paraguay ” (s. loc.) (Watkins 1976) [some records may refer to diversinodis]. hopei (Schuckard 1840 b). Canindeyú (ALWC). Literature records: “ Paraguay ” (s. loc.) (Watkins 1976). jerrmanni (Forel 1901 e). Central, “ Paraguay ” (s. loc.) (MHNG, MZSP). Literature records: “ Paraguay ” (s. loc.) (Borgmeier 1955, Forel 1901 e, Forel 1907 d). laevigatus (Borgmeier 1948). Literature records: Pte. Hayes (Fowler 1981). legionis (F. Smith 1855). Literature records: Alto Paraná, “ Paraguay ” (s. loc.) (Borgmeier 1930, Fowler 1981). orthonotus (Borgmeier 1933). Canindeyú (ALWC, INBP, LACM). pertii (Shuckard 1840 b). Central, “ Paraguay ” (s. loc.) (MCZC, MZSP, NHMB). Literature records: Guairá, “ Paraguay ” (s. loc.) (Borgmeier 1955, Forel 1895). pilosus (F. Smith 1858). Canindeyú, Cordillera, Paraguarí, “ Paraná R. ” (Dept. unkown), “ Paraguay ” (s. loc.) (ALWC, LACM, MCZC, MHNG, MZSP, NHMB). Literature records: Cordillera, “ Paraguay ” (s. loc.) (Borgmeier 1955, Emery 1894 a, Forel 1895, Forel 1909). planidorsus (Emery 1906). Literature records: Guairá (Borgmeier 1955, Emery 1906 c). pseudops (Forel 1909). Cordillera, “ Paraná R. ” (Dept. Unknown) (LACM, MHNG, MZSP, NHMB). Literature records: Cordillera (Borgmeier 1955, Forel 1909). punctaticeps (Emery 1894 a). Canindeyú, Central, Cordillera, “ Paraná R. ” (Dept. unknown) “ Paraguay ” (s. loc.) (ALWC, INBP, LACM, MCSN, MCZC, MHNG, MZSP, NHMB). Literature records: Cordillera, “ Paraguay ” (s. loc.) (Borgmeier 1955, Santschi 1916). romandii (Shuckard 1840 b). Concepción (MZSP). Literature records: Boquerón, Concepción, “ Paraguay ” (s. loc.) (Borgmeier 1955, Emery 1900, Forel 1895, Forel 1907 d). shuckardi (Emery 1900). Itapúa, “ Paraguay ” (s. loc.) (MCSN, MZSP). Literature records: Itapúa, “ Paraguay ” (s. loc.) (Borgmeier 1955, Emery 1900, Emery 1910 b) sulcatus Mayr 1868. Boquerón (MZSP). swainsonii (Shuckard 1840 a). Itapúa, “ Chaco Paraguay ” (Dept. unknown), “ Paraguay ” (s. loc.) (MCSN, MHNG, MZSP). Literature records: Guairá, Itapúa, “ Paraguay ” (s. loc.) (Borgmeier 1955, Emery 1900, Emery 1910, Forel 1895). spp. (indet males) Alto Paraguay, Amambay, Boquerón, Concepción (ALWC, INBP, MCSN).	en	Wild, Alexander L. (2007): A catalogue of the ants of Paraguay (Hymenoptera: Formicidae). Zootaxa 1622: 1-55, DOI: 10.5281/zenodo.179222
03C2878BFFDFFFDBF3D4D3DB378FF846.taxon	description	brunneum F. Smith 1858. Alto Paraguay, Alto Paraná, Amambay, Boquerón, Caaguazú, Canindeyú, Central, Cordillera, Guairá, Itapúa, Misiones, Paraguarí, Pte. Hayes, San Pedro (ALWC, IFML, INBP, LACM, NHMB, NHMW). Literature records: Central, Guairá, “ Paraguay ” (s. loc.) (Emery 1906, Forel 1895 b, Forel 1907 b). edentatum Roger 1863 a. Alto Paraguay, Caaguazú, Canindeyú, Central, Cordillera, Itapúa, Misiones, Ñeembucú, Paraguarí, Pte. Hayes, San Pedro, “ Paraguay ” (s. loc.) (ALWC, BMNH, IFML, INBP, LACM, MCSN, MHNG, MZSP). Literature records: “ Paraguay ” (s. loc.) (Emery 1896 b, Forel 1912 a). opaciventre Roger 1861 b. Amambay (ALWC, INBP, LACM). Literature records: “ Paraguay ” (s. loc.) (Emery 1894 a, Forel 1895, Forel 1907 d, Santschi 1916). permagnum Forel 1908 b. Alto Paraguay, Amambay, Caaguazú, Canindeyú, Central, “ Paraguay ” (s. loc.) (ALWC, INBP, LACM, MCSN, MZSP). Literature records: “ Paraguay ” (s. loc.) (Kempf 1972). planidens Borgmeier 1939. Amambay (INBP). tuberculatum (Olivier 1792). Alto Paraguay, Amambay, Boquerón, Caaguazú, Canindeyú, Central, Concepción, Cordillera, Paraguarí, “ Chaco ” (Dept. unknown) (ALWC, BMNH, IFML, INBP, LACM, MHNG, MZSP, NHMB, NHMW). Literature records: Central, “ Paraná R. ” (Dept. unknown) (Brown 1958, Santschi 1916, Santschi 1921 b).	en	Wild, Alexander L. (2007): A catalogue of the ants of Paraguay (Hymenoptera: Formicidae). Zootaxa 1622: 1-55, DOI: 10.5281/zenodo.179222
03C2878BFFDFFFDBF3D4D2B33489FADE.taxon	description	esenbeckii (Westwood 1842). Caaguazú, Canindeyú, Central, Concepción, Cordillera, “ Paraguay ” (s. loc.) (ALWC, INBP, LACM, MCSN, MZSP, NHMB, NHMW). Literature records: Cordillera, “ Paraguay ” (s. loc.) (Borgmeier 1955, Forel 1906 [as “ crassicornis ”], Forel 1908 b [as “ crassicornis ”], Forel 1909 [as “ crassicornis ”], Santschi 1929 a). hartigii (Westwood 1842). Canindeyú, Central, Concepción (INBP, LACM, NHMB). Literature records: Central, Guairá (Borgmeier 1955, Santschi 1916).	en	Wild, Alexander L. (2007): A catalogue of the ants of Paraguay (Hymenoptera: Formicidae). Zootaxa 1622: 1-55, DOI: 10.5281/zenodo.179222
03C2878BFFDCFFD8F3D4D57633B6FC35.taxon	description	goeldii Forel 1893 b. Canindeyú (ALWC, MHNG). Literature records: Alto Paraná (Emery 1906 [as “ pachycera ”]).	en	Wild, Alexander L. (2007): A catalogue of the ants of Paraguay (Hymenoptera: Formicidae). Zootaxa 1622: 1-55, DOI: 10.5281/zenodo.179222
03C2878BFFDCFFD8F3D4D5963439FA55.taxon	description	aphidicola Forel 1909. Amambay, Boquerón, Canindeyú, Central, Concepción, Cordillera, Itapúa, Pte. Hayes, “ Paraguay ” (s. loc.) (ALWC, BMNH, INBP, LACM, MHNG, NHMB). Literature records: Cordillera, “ Paraguay ” (s. loc.) (Forel 1909, Santschi 1923 a [as “ fallax ”]). NEW STATUS. cordemoyi Forel 1895 a. Caaguazú, Canindeyú, Central, Concepción, Cordillera, Itapúa, Misiones, Ñeembucú, Paraguarí, San Pedro (ALWC, BMNH, IFML, INBP, LACM, MHNG). Literature records: Cordillera (Forel 1908 b). fiebrigi Forel 1908 b. Cordillera, Itapúa (ALWC, MHNG). Literature records: Cordillera, “ Paraguay ” (s. loc.) (Forel 1908 b, Santschi 1923 a). heeri Forel 1874. Canindeyú, Central (ALWC, BMNH, LACM, MHNG). leuderwaldti Santschi 1923 a. Amambay, Boquerón, Canindeyú, Cordillera, Itapúa, Misiones (ALWC, BMNH, INBP, LACM, MHNG, MZSP). patagonicus Mayr 1868. Amambay, Boquerón, Canindeyú, Central, Concepción, Cordillera, Itapúa, Misiones, Ñeembucú, Paraguarí, Pte. Hayes (ALWC, IFML, INBP, LACM, MHNG, NHMW). Literature records: Cordillera (Forel 1906, Forel 1909). termitophilus Forel 1896 e. Boquerón, Central (ALWC). Literature records: “ Paraguay ” (s. loc.) (Kempf 1972). NEW STATUS. sp. alw- 01. Canindeyú (ALWC, INBP, LACM). sp. alw- 02. Boquerón (ALWC).	en	Wild, Alexander L. (2007): A catalogue of the ants of Paraguay (Hymenoptera: Formicidae). Zootaxa 1622: 1-55, DOI: 10.5281/zenodo.179222
03C2878BFFDCFFD8F3D4D6BE3242FE5D.taxon	description	bruchi (Santschi 1922 c). Canindeyú, Paraguarí (ALWC, BMNH, INBP). caelata Kempf 1967 a. Literature records: Alto Paraná (Fowler 1981). continua (Mayr 1887). Alto Paraná (MHNG). moelleri (Forel 1912 a). Boquerón, Caaguazú, Central, Itapúa, Ñeembucú, Paraguarí (ALWC, BMNH, MHNG). Literature records: Ñeembucú (Lattke 1995). rastrata (Mayr 1866 b). Canindeyú, Cordillera, Misiones (ALWC, MHNG, NHMW). regularis Mayr 1870 b. Central (INBP). Literature records: Cordillera (Forel 1909). striatula Mayr 1884. Amambay, Cordillera, “ Paraguay ” (s. loc) (ALWC, INBP, MHNG, NHMB, NHMW). Literature records: Cordillera, “ Paraguay ” (s. loc.) (Forel 1911 g, Santschi 1929 c). sulcata (F. Smith 1858). Central, Guairá, Misiones, Pte. Hayes (ALWC, IFML, INBP).	en	Wild, Alexander L. (2007): A catalogue of the ants of Paraguay (Hymenoptera: Formicidae). Zootaxa 1622: 1-55, DOI: 10.5281/zenodo.179222
03C2878BFFDCFFD8F3D4D43E3571FD1D.taxon	description	clavicornis Emery 1906. Central, Cordillera, Misiones (ALWC, INBP, MHNG). Literature records: Cordillera (Forel 1906 [as “ divergens ”]). pusillus Emery 1894 a. Concepción (MHNG). Literature records: Pte. Hayes (Fowler 1981). rogenhoferi Mayr 1862. Canindeyú (ALWC, INBP, LACM).	en	Wild, Alexander L. (2007): A catalogue of the ants of Paraguay (Hymenoptera: Formicidae). Zootaxa 1622: 1-55, DOI: 10.5281/zenodo.179222
03C2878BFFDCFFDEF3D4D3D63439FC9D.taxon	description	arboreus (F. Smith 1858). Central (ALWC, INBP). atriceps (F. Smith 1858). Caaguazú, Central (ALWC, IFML, INBP, LACM, MZSP). Literature records: Cordillera (Forel 1911 b, Hashmi 1973). balzani Emery 1894 a. Canindeyú (ALWC). bonariensis Mayr 1868. Caaguazú, Canindeyú, Cordillera (ALWC, INBP, LACM, MHNG). borellii Emery 1894 b. Alto Paraguay (ALWC). brasiliensis Mayr 1862. Alto Paraná, Caaguazú, Canindeyú, Central (ALWC, INBP, MZSP). Literature records: “ Paraguay ” (s. loc.) (Kempf 1972). cameranoi Emery 1894 b. Amambay, Canindeyú, Central, Misiones, Paraguarí, “ Paraná R. ” (Dept. uknown), “ Paraguay ” (s. loc.) (ALWC, BMNH, IFML, INBP, LACM, MCSN, MCZC, NHMW). Literature records: Cordillera, “ Paraguay ” (s. loc.) (Emery 1894 b, Emery 1896 c, Forel 1909, Santschi 1922 b). cingulatus Mayr 1862. Canindeyú, Central, Cordillera, Itapúa, Paraguarí (?), San Pedro (ALWC, MCSN, MHNG). Literature records: Central, Cordillera, San Pedro, “ Paraguay ” (s. loc.) (Emery 1894 b, Emery 1896 c [as “ lessonai ”], Forel 1909). coloratus Forel 1904 a. Boquerón (ALWC). Literature records: “ Paraguay ” (s. loc.) (Kempf 1972). NEW STATUS. crassus Mayr 1862. Alto Paraguay, Alto Paraná, Amambay, Boquerón, Caaguazú, Canindeyú, Central, Concepción, Cordillera, Guairá, Itapúa, Misiones, Ñeembucú, Paraguarí, Pte. Hayes, San Pedro, “ Paraguay ” (s. loc.) (ALWC, IFML, INBP, LACM, MCSN, MHNG, MZSP, NHMB). Literature records: Central, Cordillera, Misiones (Forel 1907 b [as “ vezenyii ”], Forel 1911 b, Fowler 1981). crispulus Santschi 1922 b. Boquerón, Central, Concepción, Pte. Hayes (ALWC, MZSP). NEW STATUS. depressus Mayr 1866 a. Alto Paraná, Canindeyú (ALWC, INBP, MCZC). dimorphus Emery 1894 a. Canindeyú (ALWC, INBP, LACM). emeryodicatus Forel 1901 b. Literature records: “ Paraguay ” (s. loc.) (Forel 1901 e). excisus Mayr 1870 a. Alto Paraná, Canindeyú (ALWC, IFML, INBP, LACM). Literature records: Cordillera (Forel 1911 b). fastigatus Roger 1863 b. Canindeyú, Ñeembucú (ALWC, IFML, INBP, LACM). fiebrigi Forel 1906. Canindeyú, Cordillera, Ñeembucú (ALWC, MHNG). Literature records: Cordillera (Forel 1906). germaini Emery 1903. Alto Paraná, Canindeyú, Concepción (ALWC, MCSN). Literature records: Alto Paraná (Emery 1911 [as “ tacuruensis ”]). helleri Emery 1903. Canindeyú. (ALWC, INBP). hispidus Emery 1906. Caaguazú, Central, Ñeembucú (ALWC, IFML, MCSN). Literature records: Ñeembucú (Emery 1906). NEW STATUS. iheringi Forel 1908 b. Canindeyú, Cordillera (ALWC, INBP). Literature records: Cordillera (Emery 1925 [as “ bajulus ”]). innocens Forel 1909. Amambay, Cordillera (ALWC, MHNG). Literature records: Cordillera (Forel 1909). NEW STATUS. lespesii Forel 1886 c. Central, Concepción, Paraguarí, (ALWC, MCSN, MHNG). Literature records: Concepción, Cordillera, “ Paraguay ” (s. loc.) (Emery 1894 b, Emery 1896 c [as “ melancholicus ”], Forel 1906, Forel 1908 b, Forel 1911 b). leydigi Forel 1886 c. Alto Paraguay, Alto Paraná, Amambay, Boquerón, Caaguazú, Canindeyú, Guairá, Pte. Hayes, San Pedro, “ Paraguay ” (s. loc.) (ALWC, IFML, INBP, LACM, MCSN, NHMB). Literature records: “ Paraguay ” (s. loc.) (Emery 1896 j, Forel 1895). macrocephalus Emery 1894 a. Literature records: “ Paraguay ” (s. loc.) (Kempf 1972 [as “ geralensis ”]). melanoticus Emery 1894 a. Alto Paraná, Caaguazú, Canindeyú, Central, Concepción, Cordillera, Guairá, Misiones, “ Paraguay ” (s. loc.) (ALWC, INBP, LACM, MCSN, MZSP, NHMW). Literature records: “ Paraguay ” (s. loc.) (Emery 1894 b, Emery 1896 b). mus Roger 1863 a. Alto Paraguay, Canindeyú, Central, Guairá, Pte. Hayes, San Pedro (ALWC, IFML, INBP, LACM, MCSN). Literature records: Cordillera, “ Paraguay ” (s. loc.) (Emery 1906, Forel 1911 b). novogranadensis Mayr 1870 a. Canindeyú, San Pedro (ALWC). Literature records: Cordillera (Forel 1906). pellitus Mayr 1862. Amambay, Caaguazú, Canindeyú, Central, Concepción, Cordillera, Itapúa (?), Misiones, Paraguarí, San Pedro, “ Paraguay ” (s. loc.) (ALWC, BMNH, IFML, INBP, LACM, MCSN, MHNG, MZSP, NHMB, NHMW). Literature records: “ Paraguay ” (s. loc.) (Emery 1896 b). REVISED STATUS. personatus Emery 1894 d. Amambay, Central, Cordillera, Itapúa, San Pedro (ALWC, IFML, INBP, MCSN, MHNG, NHMB). Literature records: Central, Cordillera (Emery 1925 [as “ alloysii ”], Forel 1906, Forel 1908 b). punctulatus Mayr 1868. Caaguazú, Central, Concepción, Cordillera, Pte. Hayes, Paraguarí (ALWC, INBP, IFML, MZSP). renggeri Emery 1894 b. Alto Paraná, Amambay, Caaguazú, Canindeyú, Central, Concepción, Cordillera, Guairá, Itapúa, Pte. Hayes, “ Paraná R ” (Dept. unknown), “ Paraguay ” (s. loc.) (ALWC, IFML, INBP, LACM, MCSN, MCZC, MHNG, MZSP, NHMB, NHMW). Literature records: Alto Paraná / Caaguazú, Caaguazú, Central, Cordillera, “ Bonpland ” (Dept. unknown), “ Paraná R. ” (Dept. unkown), “ Paraguay ” (s. loc.) (Emery 1894 b, Emery 1906, Forel 1907 b, Forel 1908 b, Forel 1909, Hashmi 1973, Santschi 1916). rosariensis Forel 1912 e. Alto Paraguay, Boquerón, Central, Ñeembucú, Pte. Hayes (ALWC, IFML, INBP, NHMB). Literature records: Misiones (Fowler 1981). NEW STATUS. rufipes (Fabricius 1775). Amambay, Caaguazú, Central, Concepción, Itapúa, Misiones, Ñeembucú, Paraguarí, Pte. Hayes, “ Paraná R ” (Dept. unknown), “ Paraguay ” (s. loc.) (ALWC, IFML, INBP, MCSN, MCZC, MZSP, NHMB). Literature records: Concepción, Cordillera, “ Paraguay ” (s. loc.) (Emery 1894 b, Emery 1896 h, Forel 1906, Forel 1907 b, Hashmi 1973). sanctaefidei Dalla Torre 1892. Caaguazú, Canindeyú, Ñeembucú, San Pedro (ALWC, IFML, INBP, LACM). scipio Forel 1908 b. Amambay, Central, Cordillera (ALWC, MCSN, MHNG, NHMB). Literature records: Central, Cordillera (Forel 1908 b, Santschi 1922 b). sericeiventris (Guérin-Méneville 1838). Alto Paraná, Amambay, Caaguazú, Canindeyú, Central, Cordillera, Guairá, Itapúa, Paraguarí, “ Paraguay ” (s. loc.) (ALWC, IFML, INBP, LACM, MZSP, NHMB, NHMW). Literature records: Central, Cordillera, Itapúa, “ Paraguay ” (s. loc.) (Forel 1906, Forel 1907 d, Forel 1911 a, Santschi 1916, Wheeler 1931). sexguttatus (Fabricius 1793). Alto Paraguay, Alto Paraná, Caaguazú, Canindeyú, Central, Concepción, Cordillera, Paraguarí, Pte. Hayes, “ Paraná R. ” (Dept. unknown), “ Chaco ” (Dept. unknown), “ Paraguay ” (s. loc.) (ALWC, IFML, INBP, LACM, MCSN, MCZC, MHNG, MZSP, NHMB, NHMW). Literature records: Central, Concepción, Cordillera (Emery 1906, Forel 1906, Forel 1907 d, Forel 1911 a). silvicola Forel 1902. Central (ALWC, INBP). Literature records: Central, Cordillera (Emery 1906, Forel 1911 b). substitutus Emery 1894 c. Alto Paraguay, Boquerón, Central, Concepción, Ñeembucú, Paraguarí (?), Pte. Hayes, “ Paraguay ” (s. loc.) (ALWC, IFML, INBP, MCSN, MHNG, NHMW). Literature records: Concepción, “ Paraguay ” (s. loc.) (Emery 1894 f, Emery 1896 b, Forel 1907 d [as “ multipilis ”]). termitarius Emery 1902. Misiones, Ñeembucú (ALWC). NEW STATUS. trapeziceps Forel 1908 b. Boquerón (ALWC). vagulus Forel 1908 b. Canindeyú, Central (ALWC). NEW STATUS. westermanni Mayr 1862. Amambay, Canindeyú, Misiones, San Pedro (ALWC). sp. alw- 01. Boquerón, Pte. Hayes (ALWC). sp. alw- 02. Caaguazú, Canindeyú, Central, Ñeembucú (ALWC, IFML, INBP). sp. alw- 03. Canindeyú (ALWC). sp. alw- 04. Pte. Hayes (ALWC). sp. alw- 05. Canindeyú (ALWC). sp. alw- 06. Boquerón (ALWC). sp. alw- 07. Canindeyú (ALWC). sp. alw- 08. Boquerón, Pte. Hayes (ALWC). sp. alw- 09. Boquerón (ALWC).	en	Wild, Alexander L. (2007): A catalogue of the ants of Paraguay (Hymenoptera: Formicidae). Zootaxa 1622: 1-55, DOI: 10.5281/zenodo.179222
03C2878BFFDAFFDEF3D4D06E3495F9CD.taxon	description	mucronata (Roger 1860). Caaguazú (MZSP).	en	Wild, Alexander L. (2007): A catalogue of the ants of Paraguay (Hymenoptera: Formicidae). Zootaxa 1622: 1-55, DOI: 10.5281/zenodo.179222
03C2878BFFDAFFDEF3D4D08E3274F88D.taxon	description	dolo (Roger 1860). Literature records: Alto Paraná, Caaguazú (Fowler 1981). flava Kempf 1962. Alto Paraná (MHNG). mayri Kempf 1962. Alto Paraná, Central, Itapúa (ALWC, BMNH, MHNG). microps Borgmeier 1957. Alto Paraná, Amambay, Itapúa (ALWC, BMNH, MHNG).	en	Wild, Alexander L. (2007): A catalogue of the ants of Paraguay (Hymenoptera: Formicidae). Zootaxa 1622: 1-55, DOI: 10.5281/zenodo.179222
03C2878BFFDAFFDEF3D4D5713403FB3D.taxon	description	arborea Forel 1909. Cordillera (MHNG). Literature records: Cordillera (Forel 1909). arthuri Forel 1903. Canindeyú (ALWC, IFML, LACM). bambusarum Forel 1903. Canindeyú (ALWC). catharinae Mayr 1887. Canindeyú (ALWC, IFML, INBP, LACM). gallicola Mayr 1887. Misiones (ALWC). nodigera Mayr 1887. Canindeyú, Concepción, Cordillera, San Pedro, “ Paraná R. ” (Dept. unknown), “ Paraguay ” (s. loc.) (ALWC, IFML, INBP, LACM, MCSN, MCZC, MHNG, NHMB). Literature records: Concepción (Emery 1896 b [as “ flavicornis ”], Forel 1907 d, Forel 1909 [as “ pallida ”]). sp. alw- 01. Canindeyú (ALWC).	en	Wild, Alexander L. (2007): A catalogue of the ants of Paraguay (Hymenoptera: Formicidae). Zootaxa 1622: 1-55, DOI: 10.5281/zenodo.179222
03C2878BFFDAFFDEF3D4D291343CFA5D.taxon	description	docilis Forel 1908 b. Canindeyú (ALWC, LACM). NEW STATUS. fulva (Mayr 1862). Alto Paraná, Amambay, Caaguazú, Canindeyú, Central, Concepción, Cordillera, Misiones, Ñeembucú, Paraguarí, Pte. Hayes, San Pedro (ALWC, BMNH, IFML, INBP, LACM, MHNG, MZSP, NHMB). Literature records: Concepción, “ Paraguay ” (s. loc.) (Emery 1906, Forel 1909 [as “ fumata ”]). * longicornis (Latreille 1802). Central, Concepción (ALWC, INBP). sp. alw- 01. Canindeyú (ALWC, BMNH, IFML, INBP, LACM, MHNG). sp. alw- 02. Canindeyú, Central, Pte. Hayes (ALWC, INBP). sp. alw- 03. Canindeyú, Concepcón (ALWC, MHNG). sp. alw- 04. Paraguarí (ALWC).	en	Wild, Alexander L. (2007): A catalogue of the ants of Paraguay (Hymenoptera: Formicidae). Zootaxa 1622: 1-55, DOI: 10.5281/zenodo.179222
03C2878BFFDAFFDFF3D4D1B13259FF7D.taxon	description	boltoni Fernández 2003 a. Itapúa (BMNH). Literature records: Itapúa (Fernández 2003 a).	en	Wild, Alexander L. (2007): A catalogue of the ants of Paraguay (Hymenoptera: Formicidae). Zootaxa 1622: 1-55, DOI: 10.5281/zenodo.179222
03C2878BFFDBFFDFF3D4D6DE3309FE9D.taxon	description	rudis Brown & Kempf 1969. Caaguazú (MZSP). Literature records: “ Paraguay ” (s. loc.) (Brandão 1991).	en	Wild, Alexander L. (2007): A catalogue of the ants of Paraguay (Hymenoptera: Formicidae). Zootaxa 1622: 1-55, DOI: 10.5281/zenodo.179222
03C2878BFFDBFFDFF3D4D77E3403F91D.taxon	description	ambiguus (Emery 1888 a). Literature records: Alto Paraná, Misiones, Ñeembucú (Fowler 1985). balzani (Emery 1890 a). Caaguazú, Central, Concepción (?), Cordillera, Itapúa, Paraguarí, “ Paraguay ” (s. loc.) (ALWC, INBP, LACM, MCSN, MHNG, MZSP, NHMB, NHMW, USNM). Literature records: Amambay, Caaguazú, Caazapá, Central, Concepción (?), Cordillera, Itapúa, “ Paraguay ” (s. loc.) (Emery 1890 a, Emery 1896 b, Forel 1895, Forel 1911 a, Fowler 1985, Santschi 1920 a, Santschi 1925 a). coronatus (Fabricius 1804). Alto Paraná, Canindeyú, Itapúa (ALWC, INBP, LACM, MZSP, USNM). Literature records: Itapúa (Fowler 1985). crassispinus (Forel 1909). Amambay, Canindeyú, Central, Concepción, Cordillera, Guairá, Ñeembucú, Paraguarí, Pte. Hayes, San Pedro, “ Paraguay ” (s. loc.) (ALWC, IFML, INBP, LACM, MHNG, MZSP, NHMB, USNM). Literature records: Canindeyú, Cordillera, Misiones, Ñeembucú, “ Paraguay ” (s. loc.) (Forel 1909 a, Fowler 1985, Santschi 1916 [as “ parallelus ”], Santschi 1925 a). disciger (Mayr 1887). Literature records: Alto Paraná, Misiones (Fowler 1985). fracticornis (Forel 1909). Boquerón, Caaguazú, Central, Concepción, Cordillera, Guairá, Ñeembucú, Pte. Hayes, San Pedro, “ Paraná R. ” (Dept. unknown) (ALWC, IFML, INBP, LACM, MCZC, MHNG, MZSP, USNM). Literature records: Central, Cordillera, Concepción, Guairá, Misiones, Paraguarí, Pte. Hayes, San Pedro, “ Paraguay ” (s. loc.) (Forel 1909, Fowler 1985, Santschi 1925 a). heyeri (Forel 1899 a). Misiones, Ñeembucú (ALWC, IFML). Literature records: Misiones, Pte. Hayes (Fowler 1985). hispidus fallax Santschi 1925 a. Caaguazú (MZSP). Literature records: Alto Paraná, Caaguazú, Central, Guairá, Ñeembucú (Fowler 1985). lobicornis (Emery 1888 a). Literature records: Itapúa, Misiones, Ñeembucú, Paraguarí, Pte. Hayes (Emery 1906, Fowler 1985). lobicornis ferrugineus (Emery 1905). Literature records: “ Paraguay ” (s. loc.) (Kempf 1972). lundii (Guérin-Méneville 1838). Alto Paraguay, Ñeembucú, Pte. Hayes, “ Paraguay ” (s. loc.) (ALWC, INBP, IFML, MCSN, USNM). Literature records: Central, Misiones, Ñeembucú (Fowler 1985). lundii decolor (Emery 1905). Literature records: “ Paraguay ” (s. loc.) (Emery 1905). niger (F. Smith 1858). Canindeyú, San Pedro (ALWC, INBP, USNM). Literature records: Cordillera (Forel 1912 b). nigrosetosus Forel 1908 b. Alto Paraná, Canindeyú (ALWC, MZSP). NEW STATUS. pubescens (Emery 1905). “ Paraguay ” (s. loc.) (MCSN, NHMB). Literature records: Pte. Hayes, “ Paraguay ” (s. loc.) (Emery 1905, Fowler 1985). NEW STATUS. rugosus (F. Smith 1858). Boquerón, Caaguazú, Canindeyú, Central, “ Paraguay ” (s. loc.) (ALWC, INBP, IFML, LACM, MCSN, MHNG, MZSP, NHMB, USNM). Literature records: Alto Paraná, Caaguazú, Canindeyú, Central, Cordillera, Guairá, Misiones, Paraguarí, “ Paraguay ” (s. loc.) (Emery 1905, Fowler 1985, Santschi 1925 a). striatus (Roger 1863 a). Pte. Hayes (ALWC, INBP, LACM, USNM). Literature records: Boquerón, Pte. Hayes (Fowler 1985). subterraneus (Forel 1893 a). Canindeyú, Central, Cordillera (ALWC, INBP, LACM, MHNG, MZSP, USNM). Literature records: Alto Paraná, Caaguazú, Canindeyú, Concepción, Cordillera, Guairá, Paraguarí, “ Paraguay ” (s. loc.) (Emery 1906, Forel 1906 h, Forel 1908 b, Forel 1909, Fowler 1985, Santschi 1925 a). sp. alw- 01. Ñeembucú, Pte. Hayes (ALWC). sp. alw- 02. Pte. Hayes (ALWC). sp. alw- 03. Ñeembucú, Pte. Hayes (ALWC). sp. alw- 04. Canindeyú (ALWC).	en	Wild, Alexander L. (2007): A catalogue of the ants of Paraguay (Hymenoptera: Formicidae). Zootaxa 1622: 1-55, DOI: 10.5281/zenodo.179222
03C2878BFFDBFFDFF3D4D0F132CAF8FD.taxon	description	pilosum Mayr 1865. Alto Paraguay, Alto Paraná, Amambay, Canindeyú, Concepción, Itapúa, Misiones (ALWC, INBP, LACM, MHNG, USNM). Literature records: Alto Paraná (Fowler 1981). steigeri Santschi 1911. Central (NHMB). Literature records: Central (Santschi 1922 a [as “ affinis ”]).	en	Wild, Alexander L. (2007): A catalogue of the ants of Paraguay (Hymenoptera: Formicidae). Zootaxa 1622: 1-55, DOI: 10.5281/zenodo.179222
03C2878BFFDBFFDCF3D4D1513581FF7D.taxon	description	capiguara Gonçalves 1944. Amambay (INBP). Literature records: Alto Paraná, Amambay, Caaguazú (Fowler 1985). laevigata (F. Smith 1858). Amambay, Canindeyú, San Pedro, “ Paraguay ” (s. loc.) (ALWC, IFML, INBP, LACM, MZSP, NHMW, USNM). Literature records: Alto Paraná, Amambay, Caaguazú, Canindeyú, Concepción, Cordillera, San Pedro (Borgmeier 1959 b, Forel 1921, Fowler 1985).	en	Wild, Alexander L. (2007): A catalogue of the ants of Paraguay (Hymenoptera: Formicidae). Zootaxa 1622: 1-55, DOI: 10.5281/zenodo.179222
03C2878BFFD8FFDCF3D4D41E345AFDDD.taxon	description	hispidula Lattke 1990. Central (BMNH).	en	Wild, Alexander L. (2007): A catalogue of the ants of Paraguay (Hymenoptera: Formicidae). Zootaxa 1622: 1-55, DOI: 10.5281/zenodo.179222
03C2878BFFD8FFDCF3D4D4BE358AFD7D.taxon	description	disciger (Mayr 1887). Alto Paraná, Itapúa (ALWC, BMNH, MHNG).	en	Wild, Alexander L. (2007): A catalogue of the ants of Paraguay (Hymenoptera: Formicidae). Zootaxa 1622: 1-55, DOI: 10.5281/zenodo.179222
03C2878BFFD8FFDCF3D4D4DE34B4FCFD.taxon	description	mayri (Forel 1901 e). Literature records: “ Paraguay ” (s. loc.) (Forel 1901 e). sp. alw- 01. Concepción (ALWC, MHNG).	en	Wild, Alexander L. (2007): A catalogue of the ants of Paraguay (Hymenoptera: Formicidae). Zootaxa 1622: 1-55, DOI: 10.5281/zenodo.179222
03C2878BFFD8FFDCF3D4D55135A2FC1D.taxon	description	bicolor Emery 1906. Literature records: Caaguazú (Fowler 1981).	en	Wild, Alexander L. (2007): A catalogue of the ants of Paraguay (Hymenoptera: Formicidae). Zootaxa 1622: 1-55, DOI: 10.5281/zenodo.179222
03C2878BFFD8FFDDF3D4D5F13424FBBD.taxon	description	angustus (Mayr 1862). Central (ALWC). atratus (Linneaus 1758). Alto Paraguay, Alto Paraná, Boquerón, Caaguazú, Canindeyú, Central, Concepción, Cordillera, Itapúa, Ñeembucú, Paraguarí, Pte. Hayes, San Pedro, “ Paraguay ” (s. loc.) (ALWC, LACM, MCSN, MHNG, MZSP, NHMB, NHMW). Literature records: Caaguazú, Central, Cordillera, “ Paraguay ” (s. loc.) (de Andrade & Baroni-Urbani 1999, Emery 1896 b, Forel 1906, Forel 1907 d, Santschi 1920 b). bohlsi (Emery 1896 b). Concepción (?) (MCSN, MHNG, NHMW). Literature records: Concepción (?) (de Andrade & Baroni-Urbani 1999, Emery 1896 b). borgmeieri (Kempf 1951). Alto Paraguay, Alto Paraná, Caaguazú, Canindeyú, Central, Paraguarí, Pte. Hayes, “ Paraguay ” (s. loc.) (ALWC, IFML, INBP, LACM, MCSN, MCZC, MHNG, MZSP). Literature records: Alto Paraná, Caaguazú, “ Paraguay ” (s. loc.) (de Andrade & Baroni-Urbani 1999). bruchi (Forel 1912 b). Central, Pte. Hayes (ALWC, INBP). clypeatus (Fabricius 1804). Alto Paraguay, Amambay, Caaguazú, Canindeyú, Concepción, Cordillera, Guairá, Itapúa, Misiones, Ñeembucú, Paraguarí, Pte. Hayes, “ Paraguay ” (s. loc.) (ALWC, IFML, INBP, LACM, MCSN, MHNG, MZSP, NHMB, NHMW). Literature records: Alto Paraná, Amambay, Caaguazú, Central, Cordillera, Guairá, Misiones, “ Paraguay ” (s. loc.) (de Andrade & Baroni-Urbani 1999, Emery 1896 b, Emery 1906, Forel 1906, Santschi 1916). depressus (Klug 1824). Alto Paraguay, Alto Paraná, Boquerón, Caaguazú, Canindeyú, Central, Cordillera, Itapúa, Ñeembucú, “ Paraguay ” (s. loc.) (ALWC, BMNH, IFML, INBP, LACM, MCSN, MCZC, MZSP, NHMB). Literature records: Caaguazú, Central, Cordillera, Guairá, San Pedro, “ Paraguay ” (s. loc.) (de Andrade & Baroni- Urbani 1999, Emery 1894 b, Emery 1906, Forel 1906, Santschi 1916). eduarduli (Forel 1921). Amambay, Caaguazú, Canindeyú, Central, Concepción, Itapúa, Ñeembucú, Paraguarí (ALWC, INBP, LACM, MHNG, MZSP). Literature records: Caaguazú, Concepción, Cordillera (de Andrade & Baroni-Urbani 1999, Forel 1921). fiebrigi (Forel 1906). Boquerón, Central, Cordillera, Ñeembucú, Pte. Hayes, San Pedro (ALWC, INBP, IFML, MHNG). Literature records: Caaguazú, Cordillera (de Andrade & Baroni-Urbani 1999, Forel 1906). guayaki De Andrade & Baroni-Urbani 1999. Concepción, Ñeembucú (ALWC, BMNH, MHNG). Literature records: Concepción (?) (de Andrade & Baroni-Urbani 1999, Emery 1896 b). incertus (Emery 1906). Central, Ñeembucú (ALWC, INBP, MHNG). Literature records: Ñeembucú (de Andrade & Baroni-Urbani 1999, Emery 1906). jheringi (Emery 1894 a). Central, Ñeembucú (ALWC). Literature records: Central, Concepción (?), “ Paraguay ” (s. loc.) (de Andrade & Baroni-Urbani 1999, Emery 1894 d, Emery 1896 b). lanuginosus (Santschi 1919). Canindeyú (ALWC, INBP, LACM). maculatus (F. Smith 1876). Canindeyú, Central, Ñeembucú, Paraguarí, Pte. Hayes (ALWC, IFML, INBP, LACM). Literature records: Central, Cordillera, “ Paraguay ” (s. loc.) (de Andrade & Baroni-Urbani 1999, Emery 1894 a, Forel 1911 g). minutus (Fabricius 1804). Alto Paraguay, Amambay, Canindeyú, Central, Concepción, Cordillera, Ñeembucú, San Pedro (ALWC, IFML, INBP, LACM, MHNG). Literature records: Caaguazú, Central, Cordillera, “ Paraguay ” (s. loc.) (Forel, 1909 a, de Andrade & Baroni-Urbani 1999). pallidoides de Andrade, in de Andrade & Baroni-Urbani 1999. Canindeyú, Central (ALWC, INBP). pellans de Andrade, in de Andrade & Baroni-Urbani 1999. Canindeyú (?), Cordillera (ALWC, IFML, INBP, LACM, MHNG). Literature records: Boquerón / Alto Paraguay, Cordillera, “ Paraguay ” (s. loc.) (de Andrade & Baroni- Urbani 1999, Emery 1896 b, Forel 1895). persimilis de Andrade, in de Andrade & Baroni-Urbani 1999. Canindeyú, Central, Cordillera, Ñeembucú, Paraguarí, Pte. Hayes (ALWC, IFML, INBP, MHNG, NHMB). Literature records: Central, Concepción, “ Paraguay ” (s. loc.) (de Andrade & Baroni-Urbani 1999, Emery 1896 b, Forel 1907 b, Forel 1907 d [as “ grandinosus ”]). pilosus (Emery 1896 b). Concepción (?) (MCSN, MHNG). Literature records: Concepción (?) (de Andrade & Baroni- Urbani 1999, Emery 1896 b). pinelii (Guérin-Méneville 1844). Literature records: Ñeembucú (de Andrade & Baroni-Urbani 1999). pusillus (Klug 1824). Alto Paraguay, Alto Paraná, Amambay, Boquerón, Caaguazú, Canindeyú, Central, Concepción, Cordillera, Guairá, Itapúa, Misiones, Paraguarí, Pte. Hayes, San Pedro, “ Chaco ” (Dept. unknown), “ Paraguay ” (s. loc.) (ALWC, IFML, INBP, LACM, MCSN, MCZC, MHNG, MZSP, NHMB, NHMW). Literature records: Alto Paraguay / Boquerón, Alto Paraná, Caaguazú, Central, Concepción, Cordillera, “ Chaco ” (Dept. unknown) (de Andrade & Baroni-Urbani 1999, Emery 1906, Forel 1906, Forel 1909, Santschi 1916, Santschi 1921 c). quadratus (Mayr 1868). Canindeyú, Central, Pte. Hayes, “ Paraguay ” (s. loc.) (ALWC, INBP, LACM, MCSN, NHMW). targionii (Emery 1894 a). Canindeyú (ALWC, LACM). Literature records: Cordillera (de Andrade & Baroni-Urbani 1999, Forel 1911 b). sp. alw- 01. Central (ALWC).	en	Wild, Alexander L. (2007): A catalogue of the ants of Paraguay (Hymenoptera: Formicidae). Zootaxa 1622: 1-55, DOI: 10.5281/zenodo.179222
03C2878BFFD9FFE2F3D4D21137D0FE3D.taxon	description	acuta (Fabricius 1804). Canindeyú (ALWC, IFML, INBP, JTLC, LACM). ampla Forel 1912 c. Canindeyú (ALWC). arata Emery 1906. Canindeyú, Guairá, Misiones (ALWC, BMNH, IFML, INBP, JTLC, LACM, MHNG). bruchi Forel 1912 c. Boquerón, Pte. Hayes (ALWC, JTLC). chodati Forel 1921. Alto Paraná, Amambay, Caaguazú, Canindeyú, Central, Concepción, Cordillera, Itapúa, Misiones, Ñeembucú, Paraguarí, Pte. Hayes, San Pedro, “ Paraguay ” (s. loc.) (ALWC, BMNH, INBP, IFML, JTLC, LACM, MHNG). Literature records: Concepción (Forel 1921). cisplatinalis Mayr 1887. Caaguazú, Canindeyú, Central, Misiones (ALWC, BMNH, IFML, JTLC, MHNG). REVISED STATUS. corticicola Mayr 1887. Caaguazú, Canindeyú (ALWC, INBP, JTLC, LACM, MZSP). crinosa Mayr 1862. Caaguazú, Canindeyú, Central, Concepción, Ñeembucú, Pte. Hayes, San Pedro (ALWC, BMNH, IFML, INBP, JTLC, LACM, NHMB). Literature records: Cordillera, Itapúa, “ Paraguay ” (s. loc.) (Emery 1896 b, Emery 1906 c, Forel 1901 a, Forel 1909, Forel 1911 b, Forel 1912 c). curvispinosa Mayr 1862. Amambay, Caaguazú, Canindeyú, Central, Concepción, Cordillera, Ñeembucú, Pte. Hayes, San Pedro (ALWC, BMNH, IFML, INBP, JTLC, LACM, MHNG, MZSP, NHMB). Literature records: Central, Concepción, Cordillera (Forel 1907 b, Forel 1907 d, Forel 1909). montezumia F. Smith 1858. Canindeyú (ALWC, JTLC). nigropilosa Mayr 1870 a. Alto Paraná, Canindeyú, San Pedro (ALWC, IFML, INBP, JTLC, LACM). nitidiceps Emery 1895 a. Caaguazú, Canindeyú, Guairá, San Pedro (ALWC, BMNH, IFML, JTLC, LACM). Literature records: Cordillera (Forel 1911 b [as “ pergens ” (unavailable name, examined)]). NEW STATUS. quadriformis Roger 1863 a. Boquerón, Caaguazú, Canindeyú, Central, Concepción, Guairá, Itapúa, Misiones, Ñeembucú, Pte. Hayes, San Pedro (ALWC, INBP, JTLC, LACM, NHMB). Literature records: Central, Cordillera, “ Paraguay ” (s. loc.) (Forel 1911 b [as “ gracilior ”], Kempf 1972). rochai Forel 1903. Alto Paraguay, Canindeyú, Central, Concepción, Pte. Hayes (ALWC, IFML, INBP, JTLC, LACM, NHMB). Literature records: “ Paraguay ” (s. loc.) (Kempf 1972 [as “ malevolens ”]). scelerata Santschi 1917. Cordillera (ALWC, JTLC, LACM). sericea Forel 1912 c. Canindeyú (ALWC, JTLC). Literature records: “ Paraguay ” (s. loc.) (Kempf 1972). thalia Forel 1911 b. Canindeyú, Cordillera (ALWC, MHNG). Literature records: Cordillera (Forel 1911 b). torosa Mayr 1870 a. Literature records: Central (Forel 1907 b). victima F. Smith 1858. Amambay, Caaguazú, Canindeyú, Central, Paraguarí (ALWC, BMNH, INBP, JTLC, LACM, MHNG). Literature records: “ Paraguay ” (s. loc.) (Emery 1896 b). sp. alw- 01. Canindeyú (ALWC). sp. alw- 02. Boquerón, Paraguarí (ALWC). [brevispinosa sericea var. semisericea Santschi 1923 b, unavailable name]. Literature records: Central (Santschi 1923 b). [brevispinosa tumulifera var. tumulicula Forel 1909, unavailable name]. Cordillera (MHNG). Literature records: Cordillera (Forel 1909).	en	Wild, Alexander L. (2007): A catalogue of the ants of Paraguay (Hymenoptera: Formicidae). Zootaxa 1622: 1-55, DOI: 10.5281/zenodo.179222
03C2878BFFE6FFE2F3D4D79E3570FC1D.taxon	description	laevigatus Weber 1938. Canindeyú (ALWC). lectus Forel 1911 a. Concepción (LACM). minutus Mayr 1862. Canindeyú, Misiones (ALWC, INBP, LACM, USNM). olitor Forel 1893 a. Canindeyú (USNM). rimosus (Spinola 1851). Alto Paraná, Caaguazú (MZSP). Literature records: Canindeyú, “ Paraguay ” (s. loc.) (Emery 1894 a, Fowler 1981). transversus Emery 1894 a. Boquerón, Canindeyú, Central, Concepción, Pte. Hayes (ALWC, INBP, LACM, MHNG, USNM). Literature records: Ñeembucú (Fowler 1981). sp. alw- 01. Canindeyú, Central (ALWC, MHNG). sp. alw- 02. Canindeyú (ALWC). sp. alw- 03. Alto Paraná (ALWC). sp. alw- 04. Amambay (ALWC, USNM). sp. alw- 05. Canindeyú, San Pedro (ALWC, USNM). sp. alw- 06. Boquerón (ALWC, USNM). sp. alw- 07. Amambay, Canindeyú (ALWC, LACM, USNM).	en	Wild, Alexander L. (2007): A catalogue of the ants of Paraguay (Hymenoptera: Formicidae). Zootaxa 1622: 1-55, DOI: 10.5281/zenodo.179222
03C2878BFFE6FFE2F3D4D5F13424FB3D.taxon	description	bruchi (Santschi 1922 c). Itapúa (ALWC). sp. A. Canindeyú (MZSP). [new sp. - Dietz, pers com.] sp. alw- 01. Alto Paraná (ALWC, BMNH, MHNG). sp. alw- 02. Misiones (ALWC, BMNH, MHNG). sp. alw- 03. Central (ALWC).	en	Wild, Alexander L. (2007): A catalogue of the ants of Paraguay (Hymenoptera: Formicidae). Zootaxa 1622: 1-55, DOI: 10.5281/zenodo.179222
03C2878BFFE6FFE2F3D4D2913586FA9D.taxon	description	balzani (Emery 1894 a). Alto Paraná, Canindeyú, Central, Concepción, Itapúa (ALWC, BMNH, LACM, MCSN, MHNG). Literature records: Central (Emery 1894 a). reitteri (Mayr 1887). Alto Paraná, Itapúa (ALWC, BMNH, MHNG).	en	Wild, Alexander L. (2007): A catalogue of the ants of Paraguay (Hymenoptera: Formicidae). Zootaxa 1622: 1-55, DOI: 10.5281/zenodo.179222
03C2878BFFE6FFE2F3D4D37137D0FA5D.taxon	description	drifti Kempf 1961. Alto Paraná, Amambay, Central, Concepción (ALWC, BMNH, MHNG). incisus M. R. Smith 1947. Amambay (ALWC, INBP). silvestrii Wheeler 1909. Amambay (ALWC). Literature records: Pte. Hayes (Fowler 1981). sp. alw- 01. Itapúa (ALWC).	en	Wild, Alexander L. (2007): A catalogue of the ants of Paraguay (Hymenoptera: Formicidae). Zootaxa 1622: 1-55, DOI: 10.5281/zenodo.179222
03C2878BFFE6FFE2F3D4D3D1343EF9DD.taxon	description	* floricola (Jerdon 1851). Canindeyú, Central (ALWC, INBP, LACM). * pharaonis (Linnaeus 1758). Canindeyú, Central, Concepción (ALWC, IFML, INBP). Literature records: Central, Concepción (Fowler 1981).	en	Wild, Alexander L. (2007): A catalogue of the ants of Paraguay (Hymenoptera: Formicidae). Zootaxa 1622: 1-55, DOI: 10.5281/zenodo.179222
03C2878BFFE6FFE2F3D4D0B135C0F95D.taxon	description	parallelus (Emery 1906). Central, Misiones, Ñeembucú (ALWC, USNM). sp. alw- 01. Caaguazú, Canindeyú (ALWC, IFML, INBP, LACM, USNM).	en	Wild, Alexander L. (2007): A catalogue of the ants of Paraguay (Hymenoptera: Formicidae). Zootaxa 1622: 1-55, DOI: 10.5281/zenodo.179222
03C2878BFFE6FFE2F3D4D13133B7F8FD.taxon	description	emeryi (Forel 1907 a). Boquerón (ALWC, IFML, INBP, MZSP, USNM). Literature records: Central (Fowler 1981).	en	Wild, Alexander L. (2007): A catalogue of the ants of Paraguay (Hymenoptera: Formicidae). Zootaxa 1622: 1-55, DOI: 10.5281/zenodo.179222
03C2878BFFE6FFE2F3D4D15E3403F85D.taxon	description	clorindae Kusnezov 1949. Literature records: Misiones (Fowler 1981). explicata Kempf 1968. Boquerón (ALWC). sp. alw- 01. Canindeyú (USNM).	en	Wild, Alexander L. (2007): A catalogue of the ants of Paraguay (Hymenoptera: Formicidae). Zootaxa 1622: 1-55, DOI: 10.5281/zenodo.179222
03C2878BFFE7FFE3F3D4D6BE32B5FE9D.taxon	description	goeldii (Forel 1893 b). Amambay, Caaguazú, Canindeyú, Central, Concepción, Paraguarí, San Pedro (ALWC, LACM, USNM). Literature records: Caaguazú (Fowler 1981). smithii (Forel 1893 b). Alto Paraná, Amambay, Canindeyú, Central, Concepción, Misiones, Ñeembucú (ALWC, BMNH, LACM, MHNG, MZSP, USNM). Literature records: Misiones (Fowler 1981).	en	Wild, Alexander L. (2007): A catalogue of the ants of Paraguay (Hymenoptera: Formicidae). Zootaxa 1622: 1-55, DOI: 10.5281/zenodo.179222
03C2878BFFE7FFE3F3D4D77E3407FE5D.taxon	description	squamosa F. Smith 1860. Canindeyú, Central, Concepción, Cordillera, Itapúa, Ñeembucú, Pte. Hayes, “ Paraguay ” (s. loc.) (ALWC, BMNH, INBP, LACM, MHNG, NHMB, NHMW, USNM). Literature records: Cordillera (Forel 1911 a [as “ uncinata ”], Forel 1911 b). sp. alw- 01. Amambay (USNM).	en	Wild, Alexander L. (2007): A catalogue of the ants of Paraguay (Hymenoptera: Formicidae). Zootaxa 1622: 1-55, DOI: 10.5281/zenodo.179222
03C2878BFFE7FFE3F3D4D43E3400FCBD.taxon	description	asper (Mayr 1887). Alto Paraná, Canindeyú, Paraguarí (ALWC, IFML, INBP, MZSP). echinatinodis (Forel 1886 b). Canindeyú, Ñeembucú (ALWC, IFML, INBP, LACM). spininodis (Mayr 1887). Canindeyú, Central, Cordillera, Misiones, Ñeembucú, Paraguarí, Pte. Hayes (ALWC, IFML, INBP, LACM, MZSP). Literature records: “ Paraguay ” (s. loc.) (Emery 1896 b). vicinus (Mayr 1887). Caaguazú, Canindeyú (ALWC, MZSP). sp. alw- 01. Central, Pte. Hayes (ALWC). sp. alw- 02. Pte. Hayes (ALWC).	en	Wild, Alexander L. (2007): A catalogue of the ants of Paraguay (Hymenoptera: Formicidae). Zootaxa 1622: 1-55, DOI: 10.5281/zenodo.179222
03C2878BFFE7FFE3F3D4D51E327AFCDD.taxon	description	semipolitus Mayr 1878. Literature records: Misiones (Fowler 1981 [as “ argentinus ”]).	en	Wild, Alexander L. (2007): A catalogue of the ants of Paraguay (Hymenoptera: Formicidae). Zootaxa 1622: 1-55, DOI: 10.5281/zenodo.179222
03C2878BFFE7FFE3F3D4D5B13364FBDD.taxon	description	balzani (Emery 1894 a). Alto Paraná, Caaguazú, Canindeyú, Central, Concepción, Guairá, Itapúa (ALWC, BMNH, INBP, LACM, MHNG, MZSP). iheringi (Emery 1888 a). Canindeyú, Concepción (ALWC, BMNH, INBP, LACM, MZSP). rugifera (Mayr 1887). Alto Paraná, Canindeyú, Itapúa (ALWC, BMNH, MHNG). sp. M. Paraguarí (MZSP). [undescribed species, B. Dietz pers com.] sp. S. Canindeyú, Concepción (ALWC, BMNH, MHNG, MZSP). [undescribed species, B. Dietz pers com.]	en	Wild, Alexander L. (2007): A catalogue of the ants of Paraguay (Hymenoptera: Formicidae). Zootaxa 1622: 1-55, DOI: 10.5281/zenodo.179222
03C2878BFFE7FFE3F3D4D2B13298FA9D.taxon	description	bruchi Santschi 1926. Central (ALWC, BMNH). rastratus (Mayr 1887). Canindeyú (ALWC, INBP, LACM). reticulatus Kempf 1974. Itapúa (ALWC, MHNG). vezenyii (Forel 1907 b). Central (ALWC, INBP). Literature records: Concepción (Forel 1907 d).	en	Wild, Alexander L. (2007): A catalogue of the ants of Paraguay (Hymenoptera: Formicidae). Zootaxa 1622: 1-55, DOI: 10.5281/zenodo.179222
03C2878BFFE7FFE0F3D4D3713370F83D.taxon	description	aberrans Mayr 1868. Caaguazú, Central, Cordillera (ALWC, IFML, INBP, MZSP). Literature records: Central (Fowler 1981). auropilosa Mayr 1887. Caaguazú (MZSP). Literature records: Alto Paraná (Fowler 1981). bergi Mayr 1887. Literature records: Misiones (Fowler 1981). bruchi Forel 1914 b. Literature records: Central (Fowler 1981). chrysops Wilson 2003. Amambay, Canindeyú, Central, Concepción (ALWC, IFML, INBP, MCZC, NHMB). Literature records: Canindeyú, Central (Santschi 1923 c [as “ deflexa ” {invalid name}], Wilson 2003). cornicula Wilson 2003. Amambay, Canindeyú, Concepción (ALWC). cyrtostela Wilson 2003. Amambay (ALWC, INBP). flavens Roger 1863 a. Boquerón, Canindeyú, Central, Ñeembucú, Paraguarí, Pte. Hayes, San Pedro (ALWC, IFML, INBP, LACM). Literature records: “ Paraguay ” (s. loc.) (Emery 1894 a). fimbriata Roger 1863 a. Alto Paraná, Canindeyú, Central, Cordillera, Guairá, Itapúa, “ Paraguay ” (s. loc.) (ALWC, BMNH, INBP, LACM, MCZC, MHNG, NHMW). Literature records: “ Paraguay ” (s. loc.) (Emery 1896 b, Forel 1895). fracticeps Wilson 2003. Amambay, Canindeyú, Pte. Hayes (ALWC, INBP, LACM, MCZC). Literature records: Canindeyú (Wilson 2003). gertrudae Forel 1886 b. Amambay, Canindeyú, Concepción (ALWC, INBP, LACM, MCZC). Literature records: Canindeyú (Wilson 2003). gigaflavens Wilson 2003. Boquerón, Canindeyú, Central, Concepción (ALWC, IFML, INBP). haywardi Kusnezov 1952 a. Central, Ñeembucú (ALWC). jelskii Mayr 1884. Alto Paraguay, Boquerón, Caaguazú, Central, Concepción, Ñeembucú, Paraguarí, Pte. Hayes (ALWC, IFML, INBP, MCSN). lignicola Mayr 1887. Cordillera, Itapúa (ALWC, MCZC, NHMB). Literature records: Canindeyú, Cordillera, “ Paraguay ” (s. loc.) (Forel 1906, Forel 1908 b, Kempf 1972, Wilson 2003). mosenopsis Wilson 2003. Alto Paraná, Canindeyú (ALWC, IFML, INBP, LACM, MCZC). Literature records: Canindeyú (Wilson 2003). obscurior Forel 1886 b. Pte. Hayes (ALWC). obscurithorax Naves 1985. Caaguazú, Canindeyú, Central, Cordillera, Paraguarí, Pte. Hayes, San Pedro, “ Paraná R. ” (Dept. unknown), “ Paraguay ” (s. loc.) (ALWC, IFML, INBP, LACM, MCZC, MZSP, NHMB, NHMW). Literature records: Canindeyú (Wilson 2003). oxyops Forel 1908 b. Amambay, Caaguazú, Canindeyú, Central, Cordillera, San Pedro, “ Paraná R. ” (Dept. unknown) (ALWC, IFML, LACM, MCZC, MHNG, MZSP, NHMB). Literature records: Cordillera (Forel 1908 b, Wilson 2003). radoszkowskii Mayr 1884. Amambay, Boquerón, Canindeyú, Central, Concepción, Cordillera, Itapúa, Misiones, Pte. Hayes, San Pedro (ALWC, INBP, LACM, MCZC). Literature records: Cordillera, “ Paraguay ” (s. loc.) (Emery 1896 b, Forel 1906 d, Forel 1909). reflexans Santschi 1933. Canindeyú, Central, Itapúa (ALWC, BMNH, IFML, INBP, LACM, MCZC). NEW STA- TUS. rudigenis Emery 1906. Central, Misiones (ALWC). rugatula Santschi 1933. Alto Paraná, Canindeyú, Itapúa (ALWC, BMNH, IFML, INBP, LACM, MCZC, MHNG). scapulata Santschi 1923 c. Amambay, Boquerón (ALWC). sigillata Wilson 2003. Itapúa (ALWC). spininodis Mayr 1887. Amambay, Boquerón, Canindeyú, Central, San Pedro (ALWC, IFML, INBP, MCZC, NHMB). subarmata Mayr 1884. Alto Paraná, Boquerón, Caaguazú, Canindeyú, Central, Cordillera, Itapúa, Misiones, Pte. Hayes (ALWC, BMNH, INBP, MCSN, MHNG, MZSP, NHMB). Literature records: Central, Cordillera (Emery 1890 b [as “ cornutula ”], Forel 1909). triconstricta Forel 1886 b. Literature records: Cordillera, “ Paraguay ” (s. loc.) (Forel 1909 a, Wilson 2003). tristis (F. Smith 1858). Canindeyú (ALWC). vafra Santschi 1923 c. Boquerón, Central, Itapúa, Ñeembucú (ALWC, INBP, LACM, MCZC). Literature records: “ Paraguay ” (s. loc.) (Wilson 2003). vallifica Forel 1901 b. Amambay, Boquerón, Central (ALWC, INBP, MCZC). Literature records: Amambay (Wilson 2003). sp. alw- 02. Central, Cordillera, Itapúa, Pte. Hayes (ALWC, INBP, MCZC). sp. alw- 03. Boquerón, Central, Itapúa (ALWC). sp. alw- 04. Canindeyú, Itapúa (ALWC, BMNH, MHNG). sp. alw- 05. Itapúa, Misiones (ALWC). sp. alw- 06. Central (ALWC). sp. alw- 07. Canindeyú, Central (ALWC). sp. alw- 08. Canindeyú, Central (ALWC). sp. alw- 10. Canindeyú (ALWC). sp. alw- 11. Boquerón, Pte. Hayes (ALWC). sp. alw- 12. Canindeyú (ALWC, INBP). sp. alw- 14. Central, Itapúa (ALWC). sp. alw- 15. Alto Paraná (ALWC, BMNH). sp. alw- 16. Amambay, Canindeyú (ALWC). sp. alw- 17. Canindeyú (ALWC, LACM). sp. alw- 19. Concepción, Central (ALWC). sp. alw- 22. Canindeyú. (ALWC, LACM). sp. alw- 25. Canindeyú, Cordillera, San Pedro (ALWC, INBP, LACM). sp. alw- 26. Boquerón (ALWC). spp. (diligens - group indet). Amambay, Boquerón, Caaguazú, Canindeyú, Central, Cordillera, Itapúa, San Pedro (ALWC, MZSP). spp. (fallax - group indet). Canindeyú, Pte. Hayes (ALWC). spp. (flavens- group indet). Central, Paraguarí (ALWC). [flavens tuberculata var. jheringi Emery 1894 a [invalid name]. Literature records: Ñeembucú (Emery 1906).]	en	Wild, Alexander L. (2007): A catalogue of the ants of Paraguay (Hymenoptera: Formicidae). Zootaxa 1622: 1-55, DOI: 10.5281/zenodo.179222
03C2878BFFE4FFE1F3D4D19E3222FEFD.taxon	description	cunicularius Mayr 1887. Boquerón, Pte. Hayes (ALWC, IFML, INBP). micans Forel 1914 b. Boquerón (ALWC). naegelii Forel, in Emery 1878. Amambay, Boquerón, Caaguazú, Canindeyú, Central, Cordillera, Guairá, Paraguarí, Pte. Hayes, San Pedro, “ Paraguay ” (s. loc.) (ALWC, IFML, INBP, LACM, MSCN, MCZC). Literature records: Paraguarí, “ Paraguay ” (s. loc.) (Emery 1906, Forel 1895, Taber 1998). tenuipubens Santschi 1936. Literature records: Caaguazú (Fowler 1981, Taber 1998). uruguayensis Mayr 1887. Boquerón, Ñeembucú, Pte. Hayes (ALWC, IFML).	en	Wild, Alexander L. (2007): A catalogue of the ants of Paraguay (Hymenoptera: Formicidae). Zootaxa 1622: 1-55, DOI: 10.5281/zenodo.179222
03C2878BFFE5FFE1F3D4D75E348BFE5D.taxon	description	hylaeus Kempf 1951. Caaguazú, Canindeyú, Central, Cordillera, Guairá, San Pedro (ALWC, IFML, INBP, LACM). Literature records: Guairá (Kempf 1964 b). montanus Kempf 1957. Canindeyú (ALWC).	en	Wild, Alexander L. (2007): A catalogue of the ants of Paraguay (Hymenoptera: Formicidae). Zootaxa 1622: 1-55, DOI: 10.5281/zenodo.179222
03C2878BFFE5FFE1F3D4D43E37B6FBDD.taxon	description	appretiata Borgmeier 1954 b. Itapúa (ALWC, MHNG). crassicornis (Mayr 1887). Alto Paraná, Canindeyú, Concepción, Itapúa, Paraguarí (ALWC, MHNG). Literature records: Alto Paraná, Canindeyú, Concepción, Itapúa, Paraguarí (Bolton 2000). denticulata (Mayr 1887). Canindeyú, Concepción (ALWC, BMNH, INBP, LACM, MHNG). Literature records: Alto Paraná, Canindeyú, Concepción, Pte. Hayes (Bolton 2000, Fowler 1981). eggersi (Emery 1890 b). Canindeyú, Central, Ñeembucú, Paraguarí, Pte. Hayes (ALWC, BMNH, MHNG). Literature records: Canindeyú, Central, Ñeembucú, Paraguarí (Bolton 2000). emilae (Forel 1907 b). Literature records: Central (Bolton 2000, Forel 1907 b). epinotalis (Weber 1934). Canindeyú (MHNG). fridericimuelleri (Forel 1886 a). Alto Paraná, Amambay, Canindeyú, Itapúa, Ñeembucú (ALWC, BMNH, INBP, LACM, MHNG). Literature records: Alto Paraná, Amambay, Ñeembucú (Bolton 2000). metrix Bolton 2000. Canindeyú (ALWC, BMNH, INBP). Literature records: Canindeyú (Bolton 2000). siagodens Bolton 2000. Itapúa (MHNG). Literature records: Itapúa (Bolton 2000). subedentata (Mayr 1887). Canindeyú (ALWC, INBP, LACM). tanymastax (Brown 1964). Boquerón, Canindeyú (ALWC, BMNH). Literature records: Alto Paraná, Canindeyú (Bolton 2000). xenochelyna Bolton 2000. Canindeyú, Concepción (BMNH, MHNG). Literature records: Canindeyú, Concepción (Bolton 2000).	en	Wild, Alexander L. (2007): A catalogue of the ants of Paraguay (Hymenoptera: Formicidae). Zootaxa 1622: 1-55, DOI: 10.5281/zenodo.179222
03C2878BFFE5FFE1F3D4D2B1357BFB7D.taxon	description	sp. NHM _ A. [Dietz, ms name]. Central, Itapúa (BMNH).	en	Wild, Alexander L. (2007): A catalogue of the ants of Paraguay (Hymenoptera: Formicidae). Zootaxa 1622: 1-55, DOI: 10.5281/zenodo.179222
03C2878BFFE5FFE1F3D4D2D13439F9FD.taxon	description	alzatei Kugler 1994. Amambay, Central, Paraguarí, Pte. Hayes (ALWC, BMNH, INBP, LACM, MHNG). Literature records: Central, Concepción, Paraguarí (Kugler 1994). besucheti Kugler 1994. Alto Paraná (MHNG). Literature records: Alto Paraná (Kugler 1994). bruchi Santschi 1922 a. Concepción, Misiones (BMNH, MCZC, MHNG). Literature records: Concepción, Misiones (Kugler 1994). germaini Emery 1894 a. Amambay, Canindeyú, Central, Itapúa, Misiones (ALWC, BMNH, INBP, LACM, MCZC, MHNG). Literature records: Alto Paraná, Central, Itapúa, Misiones (Kugler 1994). scobinata Kugler 1994. Alto Paraná, Amambay, Caaguazú, Canindeyú, Concepción (ALWC, BMNH, INBP, LACM, MHNG). Alto Paraná, Amambay, Caaguazú, Canindeyú, Concepción (Kugler 1994). sp. alw- 01. Boquerón (ALWC).	en	Wild, Alexander L. (2007): A catalogue of the ants of Paraguay (Hymenoptera: Formicidae). Zootaxa 1622: 1-55, DOI: 10.5281/zenodo.179222
03C2878BFFE5FFE1F3D4D05134A5F91D.taxon	description	sp. alw- 01. Canindeyú (ALWC, USNM).	en	Wild, Alexander L. (2007): A catalogue of the ants of Paraguay (Hymenoptera: Formicidae). Zootaxa 1622: 1-55, DOI: 10.5281/zenodo.179222
03C2878BFFE5FFE6F3D4D0F13448F99D.taxon	description	albidula Emery 1906. Alto Paraná, Canindeyú, Central, Ñeembucú (ALWC, IFML, INBP, LACM). clytemnestra Emery 1896 a. Caaguazú, Canindeyú, Central, Concepción (ALWC, IFML, INBP, LACM). Literature records: “ Paraguay ” (s. loc.) (Emery 1896 a). daguerrei (Santschi 1930). Canindeyú (ALWC). franki Forel 1908 b. Cordillera (MCZC). Literature records: Cordillera (Forel 1909). iheringi Forel 1908 b. Canindeyú, Concepción, Misiones, Ñeembucú, Paraguarí (ALWC, INBP, MHNG). invicta Buren 1972. Boquerón, Caaguazú, Canindeyú, Central, Guairá, Ñeembucú, Paraguarí, Pte. Hayes (ALWC, LACM). Literature records: Central (Santschi 1923 c), “ all of Paraguay ” (Trager 1991). loretana Santschi 1936. Amambay, Caaguazú, Canindeyú, Central, San Pedro (ALWC, IFML, INBP, LACM, MZSP). macdonaghi Santschi 1916. Alto Paraguay, Canindeyú, Central, Cordillera, Guairá, Paraguarí, San Pedro (ALWC, INBP, LACM, MCSN). Literature records: “ Paraguay ” (s. loc.) (Trager 1991). medioclara Santschi 1923 b. Canindeyú (ALWC, INBP, LACM). megergates Trager 1991. Boquerón (ALWC). richteri Forel 1909. Literature records: Ñeembucú (Fowler 1981). saevissima (F. Smith 1855). Canindeyú, Central, Ñeembucú (ALWC, INBP, LACM). Literature records: Cordillera (Forel 1911 b). scelesta Forel 1908 b. Concepción, Cordillera, Pte. Hayes (ALWC, MHNG). Literature records: Cordillera (Forel 1911 b). schmalzi Forel 1901 a. Canindeyú (ALWC, INBP). steigeri Santschi 1916. Pte. Hayes (ALWC). stricta Emery 1896 a. Canindeyú (ALWC, LACM). substituta Santschi 1925 d. Amambay (ALWC). subtilis Emery 1896 a. Central, “ Paraguay ” (s. loc.) (MCZC, MZSP, NHMW). Literature records: Central, “ Paraguay ” (s. loc.) (Emery 1896 a, Santschi 1923 b). tetracantha Emery 1906. Canindeyú, Itapúa (ALWC). wasmannii Emery 1894 a. Caaguazú, Central, Misiones, Pte. Hayes, “ Paraná R. ” (Dept. unknown), “ Paraguay ” (s. loc.) (ALWC, BMNH, IFML, INBP, MCZC, MHNG, MZSP, NHMW). Literature records: Central, Concepción, Cordillera, Itapúa, Guairá, “ Paraguay ” (s. loc.) (Emery 1894 a, Emery 1896 b, Emery 1906, Forel 1911 e [as “ transformis ”], Fowler 1981). sp. alw- 01. Canindeyú (ALWC, INBP, LACM). sp. alw- 02. Boquerón, Central, Pte. Hayes (ALWC, IFML, INBP). sp. alw- 03. Canindeyú (ALWC, IFML, INBP, LACM). sp. alw- 05. Canindeyú (ALWC). sp. alw- 06. Canindeyú (ALWC, INBP, LACM). sp. alw- 07. Canindeyú (ALWC, IFML, INBP, LACM, MHNG). sp. alw- 08. Pte. Hayes (ALWC, INBP). sp. alw- 09. Boquerón, Caaguazú (ALWC, INBP). sp. alw- 10. Canindeyú, Central (ALWC, INBP). sp. alw- 11. Canindeyú, Central (ALWC, INBP). sp. alw- 12. Canindeyú (ALWC). sp. alw- 13. Caaguazú, Central, Paraguarí, Pte. Hayes (ALWC, INBP). sp. alw- 14. San Pedro (ALWC). sp. alw- 15. Canindeyú (ALWC). sp. alw- 16. Central, Itapúa, Paraguarí (ALWC, MHNG). sp. alw- 18. Ñeembucú (ALWC). sp. alw- 19. Canindeyú (ALWC). sp. alw- 20. Amambay (ALWC). sp. alw- 21. Amambay (ALWC). sp. alw- 22. Itapúa (ALWC). spp. (saevissima - complex, indet.). Alto Paraguay, Alto Paraná, Amambay, Boquerón, Caaguazú, Canindeyú, Central, Concepción, Cordillera, Guairá, Pte. Hayes, San Pedro, “ Rinconada ” (Dept. unknown) (ALWC, INBP, LACM, MCSN, NHMB, NHMW).	en	Wild, Alexander L. (2007): A catalogue of the ants of Paraguay (Hymenoptera: Formicidae). Zootaxa 1622: 1-55, DOI: 10.5281/zenodo.179222
03C2878BFFE2FFE7F3D4D0713403FE9D.taxon	description	carinithorax Borgmeier 1934. Canindeyú, Concepción (ALWC, BMNH, MHNG). Literature records: Canindeyú, Concepción (Bolton 2000). cordovensis Mayr 1887. Concepción (BMNH, MHNG). Literature records: Concepción (Bolton 2000). elongata Roger 1863 a. Canindeyú, Central Concepción (ALWC, BMNH, INBP, LACM, MHNG). Literature records: Canindeyú, Central, Concepción (Bolton 2000, Fowler 1981). epelys Bolton 2000. Paraguarí (MHNG). Literature records: Paraguarí (Bolton 2000). hindenburgi Forel 1915. Literature records: Itapúa (Fowler 1981). insolita Bolton 2000. Central (MHNG). Literature records: Central (Bolton 2000). louisianae Roger 1863 a. Alto Paraná, Canindeyú, Central, Concepción, Itapúa, Paraguarí (ALWC, BMNH, MHNG). Literature records: Alto Paraná, Canindeyú, Central, Concepción, Itapúa, Paraguarí (Bolton 2000, Fowler 1981). ogloblini Santschi 1936. Literature records: Central (Fowler 1981). prospiciens Emery 1906. Literature records: Alto Paraná (Fowler 1981). saliens Mayr 1887. Literature records: Alto Paraná (Fowler 1981). schmalzi Emery 1906. Canindeyú (ALWC, MHNG). silvestrii Emery 1906. Boquerón (ALWC). sp. alw- 01. Canindeyú (ALWC).	en	Wild, Alexander L. (2007): A catalogue of the ants of Paraguay (Hymenoptera: Formicidae). Zootaxa 1622: 1-55, DOI: 10.5281/zenodo.179222
03C2878BFFE3FFE7F3D4D0FC3772F8E3.taxon	description	altisquamis Mayr 1887. Itapúa (ALWC, BMNH, MHNG). neglectus Emery 1894 a. Canindeyú, Central, Concepción, Itapúa, Pte. Hayes (ALWC, BMNH, INBP, LACM, MHNG).	en	Wild, Alexander L. (2007): A catalogue of the ants of Paraguay (Hymenoptera: Formicidae). Zootaxa 1622: 1-55, DOI: 10.5281/zenodo.179222
03C2878BFFE3FFE7F3D4D15C349FF803.taxon	description	brachycola (Roger 1861 a). Amambay (INBP).	en	Wild, Alexander L. (2007): A catalogue of the ants of Paraguay (Hymenoptera: Formicidae). Zootaxa 1622: 1-55, DOI: 10.5281/zenodo.179222
03C2878BFFE3FFE7F3D4D03E32FBF9FD.taxon	description	clavata (Fabricius 1775). Amambay (INBP). Literature records: “ Paraguay ” (s. loc.) (Forel 1895).	en	Wild, Alexander L. (2007): A catalogue of the ants of Paraguay (Hymenoptera: Formicidae). Zootaxa 1622: 1-55, DOI: 10.5281/zenodo.179222
03C2878BFFE3FFE7F3D4D77E35DBFE3D.taxon	description	* simillimum (F. Smith 1851). Literature records: Concepción (Fowler 1981).	en	Wild, Alexander L. (2007): A catalogue of the ants of Paraguay (Hymenoptera: Formicidae). Zootaxa 1622: 1-55, DOI: 10.5281/zenodo.179222
03C2878BFFE3FFE7F3D4D79E3400FCFD.taxon	description	dichrous Kempf 1967 a. Amambay (ALWC, INBP, LACM, USNM). fiebrigi Santschi 1916. “ Paraguay ” (s. loc.) (NHMB). Literature records: “ Paraguay ” (s. loc.) (Santschi 1916). holmgreni Wheeler 1925. Canindeyú (ALWC, USNM). kempfi Fowler 1982. Boquerón, Canindeyú, Paraguarí (ALWC, INBP, LACM, MZSP, USNM). Literature records: Boquerón, Canindeyú (Fowler 1981, Mayhé-Nunes and Brandão 2005). sp. alw- 01. Canindeyú, Central, Concepción, Ñeembucú (ALWC, INBP, LACM, MHNG, USNM). sp. alw- 02. Amambay, Canindeyú, Concepción, Cordillera (ALWC, INBP, LACM, USNM). sp. alw- 03. Amambay (USNM). sp. alw- 04. Canindeyú (ALWC). sp. alw- 05. Pte. Hayes (ALWC). sp. alw- 06. Pte. Hayes (ALWC). sp. alw- 07. Pte. Hayes (ALWC).	en	Wild, Alexander L. (2007): A catalogue of the ants of Paraguay (Hymenoptera: Formicidae). Zootaxa 1622: 1-55, DOI: 10.5281/zenodo.179222
03C2878BFFE3FFE7F3D4D55137AEFC5D.taxon	description	gilva Mayr 1866 a. Alto Paraguay, Cordillera, “ Paraguay ” (s. loc.) (ALWC, INBP, MCZC, MHNG, NHMB, NHMW). Literature records: Cordillera, “ Paraguay ” (s. loc.) (Emery 1919, Fernández 2003 b, Forel 1909 [as “ amblyops ”]).	en	Wild, Alexander L. (2007): A catalogue of the ants of Paraguay (Hymenoptera: Formicidae). Zootaxa 1622: 1-55, DOI: 10.5281/zenodo.179222
03C2878BFFE3FFE7F3D4D2313228FABD.taxon	description	auropunctata (Roger 1863 a). Boquerón, Caaguazú, Canindeyú, Central, Pte. Hayes (ALWC, IFML, INBP, LACM, MHNG, MSCN, MZSP). Literature records: Alto Paraná, Concepción (Emery 1894 a, Emery 1906, Forel 1907 b; [also as “ australis ”, “ nigricans ”, and “ rugosa ”]). lutzi Forel 1908 b. Itapúa (ALWC). sulcaticeps Emery 1894 a. Caaguazú, Central, Itapúa (ALWC, BMNH, MHNG). sp. alw- 01. Alto Paraná, Canindeyú, Concepción (ALWC, MHNG). sp. alw- 02. Amambay, Canindeyú, Concepción, Ñeembucú (ALWC, MHNG).	en	Wild, Alexander L. (2007): A catalogue of the ants of Paraguay (Hymenoptera: Formicidae). Zootaxa 1622: 1-55, DOI: 10.5281/zenodo.179222
03C2878BFFE3FFE7F3D4D3113403FADD.taxon	description	sp. alw- 01. Canindeyú (ALWC).	en	Wild, Alexander L. (2007): A catalogue of the ants of Paraguay (Hymenoptera: Formicidae). Zootaxa 1622: 1-55, DOI: 10.5281/zenodo.179222
03C2878BFFE3FFE4F3D4D1FC3531FEBD.taxon	description	australis Emery 1901. Amambay, Canindeyú, Concepción, Guairá, Itapúa, “ Paraguay ” (s. loc.) (ALWC, INBP, LACM, MZSP, NHMB, NHMW). Literature records: Concepción, Itapúa, “ Paraguay ” (s. loc.) (Borgmeier 1937, Emery 1896 b, Emery 1901 b, Forel 1907 d, Santschi 1929 a).	en	Wild, Alexander L. (2007): A catalogue of the ants of Paraguay (Hymenoptera: Formicidae). Zootaxa 1622: 1-55, DOI: 10.5281/zenodo.179222
03C2878BFFE0FFE4F3D4D71E352EFB7D.taxon	description	argentina (Santschi 1922 c). Canindeyú, Central, Cordillera, San Pedro, “ Paraguay ” (s. loc.) (ALWC, INBP, LACM, MCSN). distinguenda (Emery 1890 a). Alto Paraná, Amambay, Caaguazú, Canindeyú, Concepción, Itapúa (ALWC, INBP, LACM, MHNG, MZSP). Literature records: “ Paraguay ” (s. loc.) (Emery 1890 a, Forel 1895). fiebrigi (Forel 1908 b). Cordillera (MHNG). Literature records: Cordillera, Misiones (Forel 1908 b). foreli (Mayr 1887). Alto Paraná, Amambay, Canindeyú, Itapúa (ALWC, INBP, LACM, MHNG). opaciceps (Mayr 1887). Alto Paraná, Amambay, Caaguazú, Canindeyú, Central, Concepción, Cordillera, Ñeembucú, Paraguarí, Pte. Hayes (ALWC, INBP, MHNG, MZSP). Literature records: Cordillera (Forel 1908 b [as “ postangustata ”]). * punctatissima (Roger 1859). Central, Concepción (ALWC, MHNG). Literature records: Central (Forel 1907 b [as “ ergatandria ”]). reichenspergeri (Santschi 1923 c). Alto Paraná, Canindeyú, Paraguarí (ALWC). schmalzi (Emery 1986 g). Amambay (ALWC). trigona (Mayr 1887). Itapúa (ALWC). Literature records: Ñeembucú, “ Paraguay ” (s. loc.) (Emery 1896 a, Fowler 1981). wilsoni (Santschi 1925 b). Literature records: “ Paraguay ” (s. loc.) Brandão 1991. sp. alw- 01. Canindeyú, Concepción (ALWC, INBP, LACM, MZSP). sp. alw- 02. Caaguazú, Canindeyú, Central, Itapúa, Pte. Hayes (ALWC, INBP, LACM, MZSP). sp. alw- 03. Alto Paraná, Amambay, Canindeyú, Central, Concepción (ALWC, INBP, LACM, MHNG). sp. alw- 04. Amambay, Boquerón, Canindeyú, Central, Concepción, Pte. Hayes (ALWC, NBP, LACM). sp. alw- 05. Amambay, Canindeyú (ALWC, INBP, LACM). sp. alw- 06. Canindeyú, Itapúa (ALWC, MHNG). sp. alw- 08. Canindeyú, Misiones, Ñeembucú (ALWC). sp. alw- 09. Paraguarí (ALWC). sp. alw- 10. Alto Paraná, Canindeyú, Central, Concepción (ALWC). sp. alw- 11. Itapúa (ALWC). sp. alw- 12. Concepción (ALWC). sp. alw- 13. Caaguazú, Misiones, Ñeembucú (ALWC).	en	Wild, Alexander L. (2007): A catalogue of the ants of Paraguay (Hymenoptera: Formicidae). Zootaxa 1622: 1-55, DOI: 10.5281/zenodo.179222
03C2878BFFE0FFE4F3D4D2D134EAFAFD.taxon	description	bohlsi Emery 1896 b. Concepción, Ñeembucú, Pte. Hayes (ALWC, BMNH, INBP MCSN, MHNG, NHMW). Literature records: Concepción (?) (Emery 1896 b).	en	Wild, Alexander L. (2007): A catalogue of the ants of Paraguay (Hymenoptera: Formicidae). Zootaxa 1622: 1-55, DOI: 10.5281/zenodo.179222
03C2878BFFE0FFE4F3D4D35132E8F91D.taxon	description	bauri Emery 1892 f. Boquerón (ALWC, INBP). Literature records: Cordillera (Forel 1909 a). chelifer (Latreille 1802). Amambay, Caaguazú, Canindeyú, Central, Concepción, Cordillera, Itapúa, Misiones, Paraguarí, “ Paraguay ” (s. loc.) (ALWC, INBP, LACM, MCSN, MHNG, MZSP). Literature records: Cordillera (Forel 1909). haematodus (Linneaus 1758). Canindeyú, Central, Guairá, Misiones, Ñeembucú, Pte. Hayes, “ Paraguay ” (s. loc.) (ALWC, BMNH, IFML, INBP, LACM, MCSN, MHNG). Literature records: Cordillera (Forel 1909). meinerti Forel 1905. Alto Paraná, Canindeyú, Central, Cordillera, Itapúa, Paraguarí, San Pedro (ALWC, BMNH, IFML, INBP, LACM, MHNG). Literature records: “ Paraguay ” (s. loc.) (Kempf 1972). sp. alw- 01. Alto Paraguay, Caaguazú, Central, Cordillera, Itapúa (ALWC, IFML, INBP, MZSP).	en	Wild, Alexander L. (2007): A catalogue of the ants of Paraguay (Hymenoptera: Formicidae). Zootaxa 1622: 1-55, DOI: 10.5281/zenodo.179222
03C2878BFFE0FFE5F3D4D0F1371FFD5D.taxon	description	commutata (Roger 1860). Literature records: Central (Fowler 1981, Wild 2003). crassinoda (Latreille 1802). Concepción (INBP). Literature records: Concepción (Wild 2003). crenata (Roger 1861 a). Alto Paraná, Amambay, Caaguazú, Canindeyú, Cordillera, Guairá, Itapúa, Paraguarí, San Pedro (ALWC, INBP, LACM, MHNG, MZSP). Literature records: Alto Paraná, Amambay, Canindeyú, Cordillera, Guairá, Paraguarí, San Pedro (Forel 1906, Forel 1912 a, Wild 2003). harpax (Fabricius 1804). Amambay, Canindeyú, Concepción, Cordillera, Guairá, Itapúa (ALWC, BMNH, IFML, INBP, MHNG). Literature records: Amambay, Canindeyú, Cordillera “ Paraguay ” (s. loc.) (Emery 1890 a, Forel 1895, Wild 2003). lunaris (Emery 1896 a). Canindeyú, Concepción, Itapúa, Paraguarí (ALWC, BMNH, MHNG). Literature records: Canindeyú, “ Paraguay ” (s. loc.) (Emery 1896 a, Wild 2003). marginata (Roger 1861 a). Canindeyú, Central, Cordillera, Misiones, Paraguarí, “ Paraguay ” (s. loc.) (ALWC, INBP, LACM, MCSN, MHNG, NHMW). Literature records: Cordillera, Misiones, Paraguarí “ Paraguay ” (s. loc.) (Borgmeier 1959 a, Emery 1890 a, Forel 1895, Wild 2003). obscuricornis Emery 1890. Canindeyú (ALWC, INBP). Literature records: Canindeyú (Wild 2005). stigma (Fabricius 1804). Boquerón, Caaguazú, Canindeyú, Central, Concepción, San Pedro (ALWC, INBP, LACM, MZSP). Literature records: Boquerón, Canindeyú, Central, Concepción, San Pedro (Wild 2003). striata F. Smith 1858. Amambay, Caaguazú, Canindeyú, Cordillera, Guairá, Itapúa, Misiones, Paraguarí, San Pedro, “ Paraguay ” (s. loc.) (ALWC, BMNH, IFML, INBP, LACM, MCSN, MHNG, MZSP, NHMB, NHMW). Literature records: Amambay, Canindeyú, Cordillera, Guairá, Itapúa, Misiones, Paraguarí, San Pedro, “ Paraguay ” (s. loc.) (Forel 1895, Forel 1906, Wild 2003). verenae (Forel 1922). Amambay, Caaguazú, Canindeyú, Misiones, Paraguarí (ALWC, INBP, LACM, MCZC, MZSP). Literature records: Amambay, Caaguazú, Canindeyú, Misiones, Paraguarí (Wild 2005). villosa (Fabricius 1804). Alto Paraguay, Amambay, Caaguazú, Canindeyú, Central, Concepción, Cordillera, Guairá, Itapúa, Paraguarí, San Pedro, “ Paraguay ” (s. loc.) (ALWC, IFML, INBP, LACM, MCSN, MHNG, NHMB, NHMW). Literature records: Alto Paraguay, Alto Paraná, Amambay, Caaguazú, Canindeyú, Central, Concepción, Cordillera, Paraguarí, San Pedro, “ Paraguay ” (s. loc.) (Emery 1906, Forel 1895, Forel 1907 b, Wild 2003).	en	Wild, Alexander L. (2007): A catalogue of the ants of Paraguay (Hymenoptera: Formicidae). Zootaxa 1622: 1-55, DOI: 10.5281/zenodo.179222
03C2878BFFE1FFE5F3D4D25E3545FB7D.taxon	description	neotropica Bruch 1919. Itapúa (BMNH). sexarticulata Borgmeier 1954 a. Alto Paraná, Itapúa (BMNH).	en	Wild, Alexander L. (2007): A catalogue of the ants of Paraguay (Hymenoptera: Formicidae). Zootaxa 1622: 1-55, DOI: 10.5281/zenodo.179222
03C2878BFFE1FFE5F3D4D53E327DFCDD.taxon	description	exigua Kempf 1964 a. Canindeyú (ALWC). pilosula (F. Smith 1858). Canindeyú, Central, Cordillera, Pte. Hayes (ALWC, INBP).	en	Wild, Alexander L. (2007): A catalogue of the ants of Paraguay (Hymenoptera: Formicidae). Zootaxa 1622: 1-55, DOI: 10.5281/zenodo.179222
03C2878BFFE1FFE5F3D4D5B134B2FC7D.taxon	description	mutilatus Mayr 1887. Amambay (BMNH).	en	Wild, Alexander L. (2007): A catalogue of the ants of Paraguay (Hymenoptera: Formicidae). Zootaxa 1622: 1-55, DOI: 10.5281/zenodo.179222
03C2878BFFE1FFEAF3D4D37C3403FBBD.taxon	description	acanthobius (Emery 1896 b). Alto Paraguay, Alto Paraguay / Boquerón, Concepción, Pte. Hayes, “ Paraguay ” (s. loc.) (ALWC, BMNH, MCSN, MCZC, MHNG, PSWC). Literature records: Central, Concepción, Cordillera (Emery 1896 b, Forel 1907 b, Forel 1909). cubaensis (Forel 1901 b). Canindeyú, Central (ALWC, INBP). denticollis (Emery 1890 b). Alto Paraguay, Boquerón, Central, Ñeembucú, Pte. Hayes, “ Paraguay ” (s. loc.) (ALWC, BMNH, IFML, INBP, MHNG, MZSP). Literature records: Central, “ Paraguay ” (s. loc.) (Emery 1890 b, Emery 1896 b, Kempf 1960). elongatus (Mayr 1870 a). Canindeyú, Central, Ñeembucú, Pte. Hayes, San Pedro (ALWC, IFML, INBP). euryblemma (Forel 1899 b). Canindeyú (ALWC). fiebrigi (Forel 1908 b). Cordillera (MHNG). Literature records: Cordillera (Forel 1908 b, Forel 1909). filiformis (Fabricius 1804). Canindeyú, San Pedro (ALWC). flavidulus (F. Smith 1858). Amambay, Boquerón, Caaguazú, Canindeyú, Central, Cordillera, Misiones, Ñeembucú, Paraguarí, Pte. Hayes, San Pedro (ALWC, IFML, INBP, MZSP, NHMB). gracilis (Fabricius 1804). Alto Paraguay / Boquerón, Alto Paraná, Amambay, Caaguazú, Canindeyú, Central, Concepción, Cordillera, Guairá, Itapúa, Misiones, Ñeembucú, Paraguarí, San Pedro, “ Paraguay ” (s. loc.) (ALWC, BMNH, IFML, INBP, MCSN, MCZC, MZSP, NHMB, PSWC). Literature records: Caaguazú, Central, Cordillera, “ Paraguay ” (s. loc.) (Emery 1890 b, Emery 1896 b, Forel 1907 d, Forel 1908 b, Forel 1909, Fowler 1981, Santschi 1916, Santschi 1934 [as “ atrinodis ”]). holmgreni (Wheeler 1925). Amambay, Caaguazú, Canindeyú, Central, Itapúa, Ñeembucú, Paraguarí (ALWC, INBP). ita (Forel 1906). Literature records: “ Paraguay ” (s. loc.) (Kempf 1972). kuenckeli (Emery 1890 b). Central, Guairá, Misiones (ALWC, IFML, MZSP). Literature records: Guairá (Kempf 1961, Ward 1999). laevivertex (Forel 1906). Paraguarí (ALWC). lisus (Enzmann 1944). Canindeyú (ALWC). lizeri (Santschi 1922 a). Concepción, San Pedro (ALWC, INBP). NEW STATUS. longus Forel 1912 c. Canindeyú (ALWC). phyllophilus (F. Smith 1858). Caaguazú, Canindeyú, Central, Guairá, Itapúa, San Pedro (ALWC, INBP, IFML, MZSP). Literature records: Pte. Hayes (Fowler 1981). rufiventris (Forel 1911 b). Amambay, Caaguazú, Canindeyú, Central, Cordillera, Guairá, Itapúa, Paraguarí, San Pedro (ALWC, IFML, INBP, PSWC). Literature records: Cordillera (Forel 1911 b). schuppi (Forel 1901 a). Caaguazú, Canindeyú, Central, Cordillera, Guairá (ALWC, IFML, INBP, MHNG, MZSP, NHMB, PSWC). Literature records: Caaguazú, Central, Cordillera (Forel 1906, Forel 1908 b, Forel 1911 b, Ward 1989). sericeus (Mayr 1870 a). Canindeyú, Central (ALWC, INBP). simplex (F. Smith 1877). Canindeyú, Central, “ Paraná R. ” (Dept. unknown) (ALWC, IFML, INBP, MCZC). subater (Wheeler & Mann 1914). Canindeyú (ALWC, INBP). tenuis (Fabricius 1804). Pte. Hayes (ALWC). tenuissimus (Emery 1906). Canindeyú, Ñeembucú (ALWC, INBP). termitarius (F. Smith 1855). Alto Paraná, Amambay, Caaguazú, Canindeyú, Central, Guairá, Itapúa, Paraguarí, Pte. Hayes, San Pedro (ALWC, IFML, INBP, MCZC, MZSP, PSWC). Literature records: Alto Paraná, Caaguazú, Concepción, Cordillera, Guairá (Emery 1906, Forel 1907 d, Forel 1911 a, Fowler 1981). unicolor (F. Smith 1855). Guairá (IFML, MZSP). Literature records: Cordillera, Guairá (Forel 1911 b, Kempf 1958). urbanus (F. Smith 1877). Canindeyú, “ Paraguay ” (s. loc.) (ALWC, MHNG). Literature records: Caaguazú, Misiones, “ Paraguay ” (s. loc.) (Forel 1921, Fowler 1981, Ward 1989). sp. PSW- 18. Canindeyú (ALWC). sp. PSW- 57. Canindeyú (ALWC). sp. alw- 01. Canindeyú (ALWC).	en	Wild, Alexander L. (2007): A catalogue of the ants of Paraguay (Hymenoptera: Formicidae). Zootaxa 1622: 1-55, DOI: 10.5281/zenodo.179222
03C2878BFFEEFFEAF3D4D29E336BFB5D.taxon	discussion	Literature records: Central, Cordillera, Misiones, Ñeembucú, “ Paraguay ” (s. loc.) (Emery 1905, Fowler 1985). Identity of these records unclear, could pertain to A. laticeps or A. nigrosetosus.	en	Wild, Alexander L. (2007): A catalogue of the ants of Paraguay (Hymenoptera: Formicidae). Zootaxa 1622: 1-55, DOI: 10.5281/zenodo.179222
03C2878BFFEEFFEAF3D4D2DE37A3FAFD.taxon	discussion	Literature records: Cordillera (Santschi 1916). Misidentification of A. crassispinus; specimens examined (MHNG). Acromyrmex lundii parallelus may be a junior synonym of A. crassipinus (Wild, pers. obs.).	en	Wild, Alexander L. (2007): A catalogue of the ants of Paraguay (Hymenoptera: Formicidae). Zootaxa 1622: 1-55, DOI: 10.5281/zenodo.179222
03C2878BFFEEFFEAF3D4D37E3337FA3D.taxon	discussion	Literature records: “ Paraguay ” (s. loc.) (Emery 1906, Santschi 1923 a). Emery’s Paraguay record pertains to B. cordemoyi; specimen examined (MCSN). Santschi’s record not verified.	en	Wild, Alexander L. (2007): A catalogue of the ants of Paraguay (Hymenoptera: Formicidae). Zootaxa 1622: 1-55, DOI: 10.5281/zenodo.179222
03C2878BFFEEFFEAF3D4D3BE3483FA7D.taxon	discussion	Literature records: Cordillera (Forel 1909). Misidentification of B. patagonicus; specimens examined (MHNG).	en	Wild, Alexander L. (2007): A catalogue of the ants of Paraguay (Hymenoptera: Formicidae). Zootaxa 1622: 1-55, DOI: 10.5281/zenodo.179222
03C2878BFFEEFFEAF3D4D3FE375AF9BD.taxon	discussion	Literature records: Central (Emery 1906). Possible misidentification of C. pellitus or C. rosariensis.	en	Wild, Alexander L. (2007): A catalogue of the ants of Paraguay (Hymenoptera: Formicidae). Zootaxa 1622: 1-55, DOI: 10.5281/zenodo.179222
03C2878BFFEEFFEAF3D4D03E37D6F9FD.taxon	discussion	Literature records: Pte. Hayes (Fowler 1981). Possible misidentification of C. pellitus or C. rosariensis.	en	Wild, Alexander L. (2007): A catalogue of the ants of Paraguay (Hymenoptera: Formicidae). Zootaxa 1622: 1-55, DOI: 10.5281/zenodo.179222
03C2878BFFEEFFEAF3D4D07E3321F93D.taxon	discussion	Literature records: “ Paraguay? ” (Kempf 1972). Camponotus sericeiventris rex has a Central American distribution (Wheeler 1931); record probably pertains to C. sericeiventris.	en	Wild, Alexander L. (2007): A catalogue of the ants of Paraguay (Hymenoptera: Formicidae). Zootaxa 1622: 1-55, DOI: 10.5281/zenodo.179222
03C2878BFFEEFFEAF3D4D0BE3449F95D.taxon	discussion	Literature records: Central (?), Concepción, “ Paraguay ” (s. loc.) (Emery 1896 b, Forel 1907 b, Forel 1907 d). Records probably pertain to persimillis, C. grandinosus has a more northerly distribution (De Andrade and Baroni-Urbani 1999).	en	Wild, Alexander L. (2007): A catalogue of the ants of Paraguay (Hymenoptera: Formicidae). Zootaxa 1622: 1-55, DOI: 10.5281/zenodo.179222
03C2878BFFEEFFEAF3D4D0DE3419F8FD.taxon	discussion	Literature records: Cordillera, “ Paraguay ” (s. loc.) (Forel 1911 b, Forel 1912 b). Records refer to C. borgmeieri; specimens examined (MHNG). True pavonii has a more northerly distribution (De Andrade and Baroni Urbani 1999).	en	Wild, Alexander L. (2007): A catalogue of the ants of Paraguay (Hymenoptera: Formicidae). Zootaxa 1622: 1-55, DOI: 10.5281/zenodo.179222
03C2878BFFEEFFEAF3D4D17E37DCF83D.taxon	discussion	Literature records: Cordillera (Forel 1909, as “ leviusculus Emery ”). Probable misidentification of D. germaini.	en	Wild, Alexander L. (2007): A catalogue of the ants of Paraguay (Hymenoptera: Formicidae). Zootaxa 1622: 1-55, DOI: 10.5281/zenodo.179222
03C2878BFFEEFFEBF3D4D1FE33A3FF1D.taxon	discussion	Literature records: Cordillera (Forel 1909). Dorymyrmex flavus is a North American species unlikely to be conspecific with South American forms, record probably refers to Dorymyrmex paranensis.	en	Wild, Alexander L. (2007): A catalogue of the ants of Paraguay (Hymenoptera: Formicidae). Zootaxa 1622: 1-55, DOI: 10.5281/zenodo.179222
03C2878BFFEEFFEAF3D4D1BE3537F87D.taxon	discussion	Literature records: “ Paraguay ” (s. loc.) (Kempf 1972). Record may refer to D. paranensis. See discussion under D. paranensis below.	en	Wild, Alexander L. (2007): A catalogue of the ants of Paraguay (Hymenoptera: Formicidae). Zootaxa 1622: 1-55, DOI: 10.5281/zenodo.179222
03C2878BFFEFFFEBF3D4D69E33A2FF5D.taxon	discussion	Literature records: “ Paraguay ” (s. loc.) (Emery 1894 d). Record probably refers to E. burchelli sensu strictu, true E. foreli is known from Central America and northern South America (Watkins 1976).	en	Wild, Alexander L. (2007): A catalogue of the ants of Paraguay (Hymenoptera: Formicidae). Zootaxa 1622: 1-55, DOI: 10.5281/zenodo.179222
03C2878BFFEFFFEBF3D4D6DE3488FE9D.taxon	discussion	Literature records: Central (Forel 1907 b). Misidentification of Hypoponera punctatissima; specimen examined (MHNG).	en	Wild, Alexander L. (2007): A catalogue of the ants of Paraguay (Hymenoptera: Formicidae). Zootaxa 1622: 1-55, DOI: 10.5281/zenodo.179222
03C2878BFFEFFFEBF3D4D71E35C2FEDD.taxon	discussion	Literature records: Central (Forel 1907 b). Monomorium minimum is a North American species. Record may refer to the introduced species M. floricola.	en	Wild, Alexander L. (2007): A catalogue of the ants of Paraguay (Hymenoptera: Formicidae). Zootaxa 1622: 1-55, DOI: 10.5281/zenodo.179222
03C2878BFFEFFFEBF3D4D75E34E4FE7D.taxon	discussion	Literature records: Caaguazú (Fowler 1981). Klingenberg (2006) states that M. simplex is known only from coastal dune areas and was unable to verify Fowler’s (1981) record, which probably refers to the common inland species M. emeryi.	en	Wild, Alexander L. (2007): A catalogue of the ants of Paraguay (Hymenoptera: Formicidae). Zootaxa 1622: 1-55, DOI: 10.5281/zenodo.179222
03C2878BFFEFFFEBF3D4D7FE34E6FDFD.taxon	discussion	Literature records: Cordillera (Forel 1911 a). The relationship of M. squamosa uncinata and M. squamosa squamosa is unclear (J. Sosa-Calvo, pers. comm.). I have recorded a single common species in Paraguay related to M. squamosa, and Forel’s record (MHNG, examined) belongs to this species. M. squamosa is the older name, so I use it here.	en	Wild, Alexander L. (2007): A catalogue of the ants of Paraguay (Hymenoptera: Formicidae). Zootaxa 1622: 1-55, DOI: 10.5281/zenodo.179222
03C2878BFFEFFFEBF3D4D47E37C0FD1D.taxon	discussion	Literature records: “ Paraguay ” (s. loc.) (Brown 1976). Odontomachus brunneus is a North American species unlikely to be conspecific with South American forms. Name almost certainly refers to Odontomachus sp. alw- 01.	en	Wild, Alexander L. (2007): A catalogue of the ants of Paraguay (Hymenoptera: Formicidae). Zootaxa 1622: 1-55, DOI: 10.5281/zenodo.179222
03C2878BFFEFFFEBF3D4D49E33ADFD7D.taxon	discussion	Literature records: Cordillera (Forel 1911 b). Misplaced locality (see Bolton 1995).	en	Wild, Alexander L. (2007): A catalogue of the ants of Paraguay (Hymenoptera: Formicidae). Zootaxa 1622: 1-55, DOI: 10.5281/zenodo.179222
03C2878BFFEFFFEBF3D4D4FE3305FCBD.taxon	discussion	Literature records: “ Paraguay ” (s. loc.) (Kempf 1972). Pachycondyla stigma var. attrita is a Caribbean form unlikely to be conspecific with southern South American species.	en	Wild, Alexander L. (2007): A catalogue of the ants of Paraguay (Hymenoptera: Formicidae). Zootaxa 1622: 1-55, DOI: 10.5281/zenodo.179222
03C2878BFFEFFFEBF3D4D53E32E3FCDD.taxon	discussion	Literature records: Pte. Hayes, “ Paraguay ” (s. loc.) (Fowler 1981, Kempf 1972). Pheidole fallax is a Central American / Caribbean species highly unlikely to be conspecific with southern South American forms; Wilson (2003) gives distribution. Records probably refer to P. jelskii or P. obscurithorax.	en	Wild, Alexander L. (2007): A catalogue of the ants of Paraguay (Hymenoptera: Formicidae). Zootaxa 1622: 1-55, DOI: 10.5281/zenodo.179222
03C2878BFFEFFFEBF3D4D55E35FBFC7D.taxon	discussion	Literature record: Central (Forel 1907 b). Forel described Prionopelta mocsaryi from “ Asunción, Paraguay. ” Brown (in Wilson 1958) concluded that this was a misplaced locality and synonymized P. mocsaryi with the asian species P. opaca, with which it is nearly identical.	en	Wild, Alexander L. (2007): A catalogue of the ants of Paraguay (Hymenoptera: Formicidae). Zootaxa 1622: 1-55, DOI: 10.5281/zenodo.179222
03C2878BFFEFFFEBF3D4D5FE3445FB9D.taxon	discussion	Literature records: Alto Paraná (Emery 1906). Pseudomyrmex pallidus is a North and Central American form unlikely to be conspecific with southern South American species (Ward 1985); records probably refer to P. flavidulus.	en	Wild, Alexander L. (2007): A catalogue of the ants of Paraguay (Hymenoptera: Formicidae). Zootaxa 1622: 1-55, DOI: 10.5281/zenodo.179222
03C2878BFFEFFFEBF3D4D21E35EAFB3D.taxon	discussion	Literature records: Cordillera, “ Paraguay ” (s. loc.) (Emery 1906, Forel 1906, Forel 1908 b, Forel 1909). Solenopsis geminata is native from southern North Americs to northern South America (Trager 1991); records probably refer to S. saevissima or S. macdonaghi.	en	Wild, Alexander L. (2007): A catalogue of the ants of Paraguay (Hymenoptera: Formicidae). Zootaxa 1622: 1-55, DOI: 10.5281/zenodo.179222
03C2878BFFEFFFEBF3D4D2BE36DAFB5D.taxon	discussion	Literature records: Cordillera (Forel 1911 a). Solenopsis pylades was synonomized by Trager (1991) with S. xyloni, a North American species. May be a misidentification of S. macdonaghi or S. saevissima.	en	Wild, Alexander L. (2007): A catalogue of the ants of Paraguay (Hymenoptera: Formicidae). Zootaxa 1622: 1-55, DOI: 10.5281/zenodo.179222
03C2878BFFECFFE8F3D4D6C4376FFC5B.taxon	discussion	Conomyrma pyramicus var. paranensis (Santschi). Kempf 1972: 79. Dorymyrmex (Conomyrma) pyramicus paranensis Santschi. Bolton et al. 2006: catalogue. Dorymyrmex paranensis, described by Forel from Fiebrig’s Paraguayan material and known from additional recent collections in eastern Parguay, can be distinguished from all other Dorymyrmex in the region by the following combination of character states: compound eyes relatively small, about the same diameter as the distance between the frontal carinae; head of largest workers relatively broad, about as broad as long; head, mesosoma, and anterior gastric terga clear yellow to yellow-orange in color; and psammophore present but weakly developed. This ant shares the same small eye size as the common D. brunneus, but that species has a more strongly produced propodeal tooth, a more angular mesonotum, and a dark brown body color. The other common light-colored Dorymyrmex in Paraguay, the chacoan species D. exsanguis, is more gracile; the head of the largest workers at least 1.2 x as long as broad and supports a fully-developed psammophore bearing setae roughly equal in length to the head. The identity of Dorymyrmex pyramicus (Roger 1863 a), described from Bahia, Brazil, is unclear. Snelling (1995) discusses the problems with this and other names in this taxonomically problematic genus. Apparently, no subsequent taxonomists have seen Roger’s material and Roger’s description of D. pyramicus does not provide enough detail to discern anything other than a bicolored red / brown Dorymyrmex. Kusnezov (1952 b) and Fabiana Cuezzo (pers comm.) use the name for a robust, bicolored, large-eyed species that I have collected near Buenos Aires, Argentina but that has not yet been recorded from Paraguay. In any case, I find no justification for retaining the current trinomen given the uncertainty of D. pyramicus.	en	Wild, Alexander L. (2007): A catalogue of the ants of Paraguay (Hymenoptera: Formicidae). Zootaxa 1622: 1-55, DOI: 10.5281/zenodo.179222
03C2878BFFECFFE8F3D4D26332F2F88D.taxon	discussion	The collections of Nomamyrmex that I have examined from Paraguay fall neatly into two species corresponding most commonly to N. esenbeckii and more rarely to N. hartigii. Watkins (1976) and Borgmeier (1955) provide characters to separate the species. In contrast, literature records of Nomamyrmex from Paraguay pertain to three forms, N. hartigii (Borgmeier 1955, Santschi 1916), N. esenbeckii (Borgmeier 1955, Santschi 1929 a), and N. esenbeckii crassicornis (Forel 1906, Forel 1908 b, Forel 1909). The literature records of N. esenbeckii crassicornis, a subspecies generally known from the more tropical regions of South America, are based on Forel’s identifications. Thus, the presence of three forms in the literature is likely an artifact of differing conceptions on the part of earlier taxonomists. Regardless of these conceptual differences, the subspecies of N. esenbeckii are largely allopatric across the Neotropics (Watkins 1976). Gordon Snelling, who has examined material from throughout the region, provides a detailed case for the synonymy of all of them at www. armyants. org. I concur with Snelling’s reasoning and synonymize N. crassicornis under N. esenbeckii here.	en	Wild, Alexander L. (2007): A catalogue of the ants of Paraguay (Hymenoptera: Formicidae). Zootaxa 1622: 1-55, DOI: 10.5281/zenodo.179222
03C2878BFFECFFE9F3D4D1BC333EFDFD.taxon	discussion	There is little reason to consider Brachymyrmex heeri and B. aphidicola as conspecific, as both species are locally sympatric and morphologically distinct in eastern Paraguay. Brachymyrmex aphidicola is one of the most abundant ground and litter nesting ant species in Paraguayan forests. Brachymyrmex heeri is more rarely collected, the two nest records I have from Paraguay are both arboreal. The two species are easily separated by the pubescence on gastric tergite 1 (= abd. tergite 3), with the pubescence dense on B. heeri and sparse on B. aphidicola. Santschi’s type of B. heeri fallax (NHMB, examined) falls within the range of variation exhibited by B. aphidicola in Paraguay and is synonymized here. It is worth mentioning here that Paraguayan specimens of B. heeri are a close match, in both the worker and queen castes, to Forel’s types of B. heeri (MHNG, examined) described from an introduced population in Switzerland (Forel 1874). Given the notoriously poor taxonomic state of Brachymyrmex and the ubiquity of B. aphidicola in the region, there may well exist an older name for this species. A thorough taxonomic revision of this genus is sorely needed.	en	Wild, Alexander L. (2007): A catalogue of the ants of Paraguay (Hymenoptera: Formicidae). Zootaxa 1622: 1-55, DOI: 10.5281/zenodo.179222
03C2878BFFEDFFE9F3D4D4833561FBCD.taxon	discussion	Although B. termitophilus shares the dense pubescence and light coloration of B. heeri, this ant is more compact than Forel’s B. heeri types (MHNG, examined). In contrast to the relatively long antennal scapes of B. heeri, the scapes of B. termitophilus are shorter than the head length, in repose only barely meeting the posterior margin of the head in full-face view. Additionally, the propodeum of B. termitophilus in lateral view is only slightly or not at all depressed below the level of the promesonotum, whereas in B. heeri the propodeum is always noticeably lower than the promesonotum. Both species occur in Paraguay with no evidence of intergrades, an observation that supports the elevation of termitophilus to species. Brachymyrmex fiebrigi Forel (MHNG, types examined) has smaller eyes in the worker caste than B. termitophilus, but is otherwise a very similar ant and may merely represent the low end of intraspecific eye size variation in B. termitophilus. As only a few specimens of both the large- and small-eyed variants were available in the present study, I prefer to retain B. fiebrigi as valid until more material becomes available.	en	Wild, Alexander L. (2007): A catalogue of the ants of Paraguay (Hymenoptera: Formicidae). Zootaxa 1622: 1-55, DOI: 10.5281/zenodo.179222
03C2878BFFEDFFE9F3D4D2FC3581F863.taxon	discussion	Worker head shape in Myrmelachista nodigera shows slight allometric variation within nest series, with larger workers having somewhat broader heads, and this variation may have misled Emery and Forel in defining their intraspecific taxa flavicornis and pallida, both from Paraguay. Mayr’s M. nodigera type is a larger and more robust worker than the types of M. flavicornis and M. pallida, yet single nest series I have collected in Canindeyú and San Pedro span the range of variation encompassed by the type specimens for all three taxa. Color is also variable within a nest, presumably related to worker age, so Forel’s distinction of M. pallida on color is baseless. In my estimation the known Paraguayan collections related to M. nodigera comprise a single species. Forel’s type of Myrmelachista arborea (MHNG, examined) from San Bernadino is a single queen unassociated with workers. Judging from the shape of the petiole, M. arborea may be the queen of M. nodigera. Further collections to obtain complete nest series will help establish the identity of this species.	en	Wild, Alexander L. (2007): A catalogue of the ants of Paraguay (Hymenoptera: Formicidae). Zootaxa 1622: 1-55, DOI: 10.5281/zenodo.179222
03C2878BFFEAFFEEF3D4D3AC359BF903.taxon	discussion	Both C. innocens and C. trapeziceps (w syntypes examined, MHNG) occur in Paraguay, but there is little justification for viewing these sympatric forms as conspecific. To my knowledge major workers of C. innocens have never been collected, but the minor of C. innocens has a markedly more slender head in full-face view than C. trapeziceps, a more elongate mesosoma, and thicker, longer pubescence on the head and mesosoma.	en	Wild, Alexander L. (2007): A catalogue of the ants of Paraguay (Hymenoptera: Formicidae). Zootaxa 1622: 1-55, DOI: 10.5281/zenodo.179222
03C2878BFFEAFFEEF3D4D484336BFA9D.taxon	discussion	Paratrechina fulva is among the most abundant ants in Paraguay, particularly in open and riparian habitats. It can be distinguished from all other Paratrechina species in Paraguay by the presence of dense pubescence on the meso- and metapleura. Forel’s P. fumata worker syntypes are also densely pubescent, but they are darker than most collections of P. fulva. However, Forel’s specimens fall well within the variation of forms that I consider to be conspecific with P. fulva, so I synonymize P. fumata here. Fernández (2000), who provides a useful redescription and discussion of P. fulva, reached a similar conclusion about the identity of “ P. fumata ” from Brazil but stopped short of formal synonymy.	en	Wild, Alexander L. (2007): A catalogue of the ants of Paraguay (Hymenoptera: Formicidae). Zootaxa 1622: 1-55, DOI: 10.5281/zenodo.179222
03C2878BFFEAFFEFF3D4D12B3338FB0B.taxon	discussion	The names associated with Camponotus blandus present one of the most difficult taxonomic problems in the Paraguayan fauna, and one that is particularly frustrating given the abundance of these ants in open habitats throughout the region. Multiple closely-related species certainly occur in Paraguay, but aside from Santschi’s C. crispulus (see separate discussion under that species), the taxonomic status of these forms is ambiguous and the arrangement in the present study is tenuous. In eastern Paraguay, most collections are of dark-colored ants closely matching Mayr’s type of C. pellitus from Brazil and Forel’s types of C. scintillans from Paraguay. Workers from western Paraguay are almost universally bicolored with a red head and mesosoma and dark legs and gaster. The major workers of these chaco forms have finer suberect hairs on the sides of the head, spaced more densely, than do workers of C. pellitus, and the postero-lateral corners of the head of the largest workers are often more produced. Minor workers of the bicolored forms have smaller eyes than do minors of C. pellitus, and they have a slightly lower, less convex propodeum. Both forms co-occur in eastern Paraguay near the Paraguay River with little evidence of interbreeding. Forel’s types of C. rosariensis fall within the range of variation of these bicolored chacoan forms, so I adopt here a two-species scheme with C. rosariensis representing the bulk of the chacoan collections and C. pellitus representing most of the eastern collections. The chacoan forms, while difficult to subdivide further on the basis of external worker morphology, show two distinct nest phenotypes in reproductively mature colonies. I have observed these nest forms within a few meters of each other. One form constructs large, conspicuous crater mounds while the other has cryptic nest entrances in the soil consisting of a single small hole with a slight scattering of excavated earth placed up to half a meter away, a pattern similar to the nests of C. pellitus. This sympatric difference in nesting behavior suggests the existence of cryptic species. F. Smith’s holotype of C. blandus, a minor worker from Pará (BMNH, examined), is not a close match to any Paraguayan material. This ant has the smaller eyes and the lower propodeum more similar to C. rosariensis than to C. pellitus, but with denser pubescence on gastric tergites 3 and 4 (= abd. tergites 5 and 6) than in most specimens of C. rosariensis. The C. blandus type is also colored differently, with the posterior portion of the head quite dark, as in C. pellitus, the mesosoma lighter, as in C. rosariensis, and the first gastric tergite concolorous with the mesosoma and infuscated posteriorly. However, I have not examined any material occurring between the type locality in Pará and Paraguay, so I cannot dismiss the eventuality that the name C. blandus may apply one of these subtropical populations.	en	Wild, Alexander L. (2007): A catalogue of the ants of Paraguay (Hymenoptera: Formicidae). Zootaxa 1622: 1-55, DOI: 10.5281/zenodo.179222
03C2878BFFEBFFEFF3D4D3333445FA46.taxon	discussion	See discussion under Camponotus pellitus.	en	Wild, Alexander L. (2007): A catalogue of the ants of Paraguay (Hymenoptera: Formicidae). Zootaxa 1622: 1-55, DOI: 10.5281/zenodo.179222
03C2878BFFEBFFECF3D4D076332EFECD.taxon	discussion	Camponotus crispulus is locally sympatric in Paraguay with at least two other species in the C. blandus complex, most frequently the similarly-colored C. rosariensis. Both species share a similarly dense pubsence over most of the body and a common bicoloration consisting of a red head and mesosoma and a dark gaster. However, C. crispulus is structurally more compact, with shorter appendages and a relatively tall propodeum. In my experience, the easiest character to separate C. crispulus from other bicolored blandus- complex species is the unique pattern of pubescense on the gaster. While all species in the complex show some degree of medial convergence in the appressed setae on the gastric terga, in C. crispulus this convergence is extreme, starting on the first gastric tergite (= abd. tergite 3) well ahead of the posterior margin and continuing to the apex of the gaster almost as a visible line. Because of the distinct morphological separation in sympatry between C. crispulus and other closely-related forms, I find unambiguous the elevation of C. crispulus to species. Material from Paraguay is a close match to Santschi’s types from Córdoba, although Santschi’s material is somewhat more pubescent. F. Smith’s C. blandus holotype worker (BMNH, examined- see discussion under C. pellitus) from Pará is clearly a different species, as that ant is more gracile than C. crispulus and lacks the distinct pubescence.	en	Wild, Alexander L. (2007): A catalogue of the ants of Paraguay (Hymenoptera: Formicidae). Zootaxa 1622: 1-55, DOI: 10.5281/zenodo.179222
03C2878BFFE8FFECF3D4D7FC32A6FB1D.taxon	discussion	Camponotus crassus is the most frequently recorded Camponotus from Paraguay. This ant appears to be found in every terrestrial habitat type with woody vegetation of sufficient diameter to house the arboreal nests in hollow twigs. The coloration of the antennal scapes shows slight variation among collections, from testaceous to dark reddish brown. The amount of pilosity on the mesosomal dorsum also varies somewhat. Forel’s type of C. vezenyii from Asunción is slightly more pilose and has lighter colored scapes than Mayr’s type of C. crassus, but it still falls well within the range of variation observed in Paraguay and is synonymized here. Santschi (1936) distinguished the variety C. picticornis from C. crassus on the basis of antennal scape color. I was unable to view Santschi’s types of C. picticornis from Loreto, Argentina, but several other series from northern Argentina at NHMB identified by Santschi as C. picticornis are within my conception of C. crassus.	en	Wild, Alexander L. (2007): A catalogue of the ants of Paraguay (Hymenoptera: Formicidae). Zootaxa 1622: 1-55, DOI: 10.5281/zenodo.179222
03C2878BFFE8FFECF3D4D32C3584F885.taxon	discussion	Forel’s C. latinota types from San Bernadino (the replacement name is C. bajulus Emery) have slightly longer pilosity on the mesosomal dorsum and gastric terga than Forel’s iheringi type from São Paulo, but the two forms are otherwise similar. Forel (1909 a) separated the two based on surface sculpture and mesosomal structure, but the sculpture differences are slight and full nest series of this species I have collected show enough allometric variation in mesosomal structure to call into question the reliability of Forel’s distinction. Since C. iheringi and C. bajulus are allopatric and the differences are ambiguous at best, I find little reason to treat these forms as separate.	en	Wild, Alexander L. (2007): A catalogue of the ants of Paraguay (Hymenoptera: Formicidae). Zootaxa 1622: 1-55, DOI: 10.5281/zenodo.179222
03C2878BFFE8FFEDF3D4D1B434E3FDDE.taxon	discussion	Camponotus sexguttatus is a common ant throughout the neotropics, particularly in disturbed or edge habitats, that has accumulated an unfortunate number of infraspecific taxa. These names usually pertain to minor color differences. While C. sexguttatus in the broad sense may actually contain a number of cryptic species over the full range of the group, there is little chance that the current haphazard assortment of subspecies and varieties accurately represents them. Literature records from Paraguay include both C. sexguttatus (Forel 1906, Forel 1907 b [as “ bimaculatus ”], Kempf 1972) and C. sexguttatus var. fusciceps (Emery 1906, Kempf 1972). Examination of the specimens at MHNG on which the records are based shows both to belong to the single common species that I have been calling C. sexguttatus. A proper taxonomic study may eventually show that the forms from subtropical South America should have a different name, as the type of C. sexguttatus is Antillean in origin, far to the north.	en	Wild, Alexander L. (2007): A catalogue of the ants of Paraguay (Hymenoptera: Formicidae). Zootaxa 1622: 1-55, DOI: 10.5281/zenodo.179222
03C2878BFFE9FFEDF3D4D4EE3261FB5D.taxon	discussion	Hashmi (1973) never saw the types of C. lessonai and subsequently excluded C. lessonai from Myrmothrix. I located Emery’s lessonai types from San Pedro at MCSN and they are nearly identical to Forel’s type of C. damocles from San Bernadino (MHNG, examined). Workers of these forms are considerably darker than Mayr’s type of C. cingulatus and match several recent collections from eastern Paraguay. I have also encountered light-colored C. cingulatus in San Pedro. Whether the presence of sympatric color morphs indicates mere intraspecific variation or the existence of cryptic species is not clear. However, Hashmi (1973) considered C. damocles a junior synonym of C. cingulatus, and if one adopts Hashmi’s broad concepcion of the species then C. lessonai should also be synonymized.	en	Wild, Alexander L. (2007): A catalogue of the ants of Paraguay (Hymenoptera: Formicidae). Zootaxa 1622: 1-55, DOI: 10.5281/zenodo.179222
03C2878BFFE9FFEDF3D4D36C32CCF88D.taxon	discussion	Several species in the Camponotus punctulatus complex occur in Paraguay and are among the more common groundnesting ants in open habitats. Camponotus hispidus can be distinguished from the locally sympatric C. punctulatus Mayr (syntype w examined, NHMW) by the more extensive pilosity on the head of the major. Camponotus punctulatus majors normally lack standing setae on the sides of the head and between the eye and the mandibular insertion, while C. hispidus majors have numerous subdecumbent to suberect setae on the sides of the head and> 10 between the eye and the mandibular insertion. As the pilosity of C. hispidus is more extensive than that of C. punctulatus, the integument of C. hispidus appears more densely and deeply punctate. Camponotus hispidus is piceous to black in color, while C. punctulatus in Paraguay is ordinarily bicolored with a reddish brown head and mesosoma a dark brown gaster. Since these two species are sympatric and distinct in Paraguay, there is no justification for retaining C. hispidus as a subspecies.	en	Wild, Alexander L. (2007): A catalogue of the ants of Paraguay (Hymenoptera: Formicidae). Zootaxa 1622: 1-55, DOI: 10.5281/zenodo.179222
03C2878BFFE9FFF2F3D4D1BC378AFD23.taxon	discussion	The aptly-named Camponotus termitarius is most commonly collected from old termite mounds in low-lying pastures and pastizales. This ant should be treated as a full species. Camponotus termitarius is locally sympatric with C. punctulatus (syntype w examined, NHMW), shows consistent differences in nesting site (C. punctulatus is a generalist groundnesting ant), and is morphological distinct in a number of characters. First, C. termitarius is more pilose, being intermediate in this regard between the relatively hirsute C. hispidus and the relatively bare C. punctulatus, lacking extensive standing hairs on the side of the head but bearing> 8 between the eye and the mandibular insertions. Second, the propodeum of C. termitarius is lower, more elongate, and with a shorter declivitous face than that of C. punctulatus. Third, the microsculpture of the gastric tergites is more extensive, giving the gaster a duller appearance than in either C. hispidus or C. termitarius. Finally, the pubescence is more extensive in C. termitarius than in either C. hispidus or C. termitarius, with the fine appressed setae on the first gastric tergite separated by distances much shorter than the length of the setae and overlapping in places.	en	Wild, Alexander L. (2007): A catalogue of the ants of Paraguay (Hymenoptera: Formicidae). Zootaxa 1622: 1-55, DOI: 10.5281/zenodo.179222
03C2878BFFF6FFF2F3D4D3DC3774F813.taxon	discussion	Camponotus coloratus cannot be considered conspecific with C. substitutus. Both ants are locally sympatric in the Paraguayan chaco without signs of hybridization. These species share a similar striking bicoloration (see discussion under C. substitutus), but workers of C. coloratus, in addition to being on average larger than workers of C. substitutus, have disproportionately longer legs, a more opaque microsculpture, and considerably reduced pilosity with the sides of the head of the major bearing fewer than 5 standing setae. Paraguayan material is lighter in color and with a broader head in the major caste than Forel’s type from Colombia, but given the allopatric nature of the variation I see no reason to divide C. coloratus further.	en	Wild, Alexander L. (2007): A catalogue of the ants of Paraguay (Hymenoptera: Formicidae). Zootaxa 1622: 1-55, DOI: 10.5281/zenodo.179222
03C2878BFFF6FFF2F3D4D4CB35D7FA2D.taxon	discussion	Camponotus substitutus is among the more commonly encountered Camponotus in open and disturbed habitats, particularly in the chaco and near the Rio Paraguay in eastern Paraguay. This ant can be distinguished from other large Paraguayan Camponotus (Tanaemyrmex) by the following combination of character states: numerous (> 20) suberect to subdecumbent setae on the sides of the head in the major caste; scape of the major simple, not flattened at base; and striking bicoloration in all female castes, with gastric tergites 1 - 3 (= abd. tergites 3 - 6) bearing yellowish-white antero-lateral spots contrasting with a dark brown to black central area. Forel’s types of C. substitutus multipilis are more pilose than most Paraguayan collections of C. substitutus, but still within the range of variation of the species. As I can find no other characters that distinguish C. multipilis from C. substitutus I synonymize it here.	en	Wild, Alexander L. (2007): A catalogue of the ants of Paraguay (Hymenoptera: Formicidae). Zootaxa 1622: 1-55, DOI: 10.5281/zenodo.179222
03C2878BFFF7FFF3F3D4D5AB3309F98D.taxon	discussion	In my opinion the literature records of C. lespesii and C. lespesii melancholicus from Paraguay pertain to a single species, and the differences in identification stem from alternate taxonomic interpretations on the part of Forel and Emery. All of the material I have examined is rather homogenous and closely matches Emery’s melancholicus types. The distinct appearance of this ant renders it unlikely to be confused with any other species in Paraguay. Camponotus lespesii is large (mesosomal length> 3.5 mm in major workers) and more gracile than any other Paraguayan Camponotus of similar size. Additional diagnostic characters include: head of major worker considerably longer than broad; sides of head of major bearing> 15 standing setae; antennal scapes lacking standing setae; antennal scapes longer than head length; mesosoma low and elongate, declivitous propodeal face less than 1 / 3 length of dorsal face; sculpture of head, mesosoma, and gaster opaque. Forel’s lespesii type, a large minor worker from Colombia, is somewhat lighter in color than the Paraguayan specimens but is otherwise similar. For the purpose of eliminating a trinomen I synonymize C. melancholicus here.	en	Wild, Alexander L. (2007): A catalogue of the ants of Paraguay (Hymenoptera: Formicidae). Zootaxa 1622: 1-55, DOI: 10.5281/zenodo.179222
03C2878BFFF7FFF3F3D4D6BE322AFC83.taxon	discussion	Camponotus fastigatus has been interpreted broadly by Forel, Emery, and Santschi, who each named subspecies based on forms that are distinct from C. fastigatus in head shape, mesonotal structure, and pilosity. It is unlikely that any of these subspecies, including Forel’s C. vagulus, are truly conspecific as they maintain their differences in broad sympatry in southern South America. With minor workers approaching only 3 mm in length, Camponotus vagulus is one of the smallest Camponotus in the region. This ant co-occurs with C. fastigatus throughout eastern Paraguay and shows similar associations with open or edge habitats. Both ants share a robust build, a strong medial clypeal carina, and short antennal scapes that in repose fall well short of the posterior cephalic margin. However, differences between the two are numerous. In full face view, the head of the major of C. vagulus is rectangular, over 1.1 x longer than broad, with sides straight and subparallel, while the head of C. fastigatus is rounded, about as long as broad, with the sides clearly convex. The integument of C. vagulus over most of the body has little microsculpture, giving the ant a characteristic shiny appearance, while that of C. fastigatus is opaque. The pubescence of C. vagulus is shorter and more difficult to see than that of C. fastigatus, and the head of both worker castes in C. vagulus is lighter in color near the mandibular insertions. In light of the numerous differences in sympatry between the forms, the case for raising C. vagulus to species is unambiguous.	en	Wild, Alexander L. (2007): A catalogue of the ants of Paraguay (Hymenoptera: Formicidae). Zootaxa 1622: 1-55, DOI: 10.5281/zenodo.179222
03C2878BFFF7FFF0F3D4D0BC3573FEEE.taxon	discussion	Camponotus germaini is a soil-nesting species recorded from low forests, second growth forests, and edge habitats. Emery (1911 d) distinguished C. tacuruensis from C. germaini on the basis of size and head shape of the major worker. I have examined Emery’s types for both taxa. Although it is true that C. germaini tacuruensis is slightly larger than most collections of C. germaini in Paraguay, I can find no other difference to distinguish C. tacuruensis. The head shape of the major worker in spite of Emery’s written description is scarcely different between the two forms and both forms fall within the observed range of variation of this species in Paraguay.	en	Wild, Alexander L. (2007): A catalogue of the ants of Paraguay (Hymenoptera: Formicidae). Zootaxa 1622: 1-55, DOI: 10.5281/zenodo.179222
03C2878BFFF4FFF0F3D4D79E3563FC3B.taxon	discussion	Fowler (1985 a) described the differing habitat associations of Acromyrmex lundii and Acromyrmex pubescens, noting that the former is found in open habitats and the latter in the patchy forest “ islands ” that occur in the chaco savannah. The two forms are sympatric and structurally similar, both bearing elongate lateral pronotal spines that are longer than the mesonotal spines. However, these ants are distinguishable in pubescence. Much of the integument of A. pubescens is covered in a dense decumbent pubescence while the integument of A. lundi is relatively bare. This difference is easiest to diagnose on the mesopleura, as the mesopleural pubescence of A. pubescens comprises overlapping hairs, while that of A. lundii is sparse and the hairs non-overlapping. Given the ecological and morphological differences in sympatry between these two forms, I elevate A. pubescens to species here.	en	Wild, Alexander L. (2007): A catalogue of the ants of Paraguay (Hymenoptera: Formicidae). Zootaxa 1622: 1-55, DOI: 10.5281/zenodo.179222
03C2878BFFF4FFF0F3D4D5C33406F9F5.taxon	discussion	Crematogaster cisplatinalis is a twig-nesting species recorded most frequently from low-lying open habitats in the Paraguay river floodplain. This species is distinct in a number of characters from C. victima, a slightly larger ant that tends to be found in the better-drained habitats of the eastern forests and cerrados. There is little reason to retain the subspecific status of C. cisplatinalis as the two forms overlap substantially in distribution but differ consistently in numerous characters. Among Paraguayan collections these differences include: standing setae present on tibiae and antennal scapes of C. cisplatinalis (absent in C. victima); dorsum of head almost entirely punctate in C. cisplatinalis, or if a smooth medial strip present the strip is narrower than the width of the compound eye (smooth medial strip present, usually wider than width of compound eye, in C. victima); color brown to black in C. cisplatinalis (testaceous to medium brown in C. victima). Jack Longino, who has conducted extensive taxonomic work on Neotropical Crematogaster (Longino 2003), confirmed the identifications for both C. victima and C. cisplatinalis in Paraguay and concurs with a change in status of C. cisplatinalis (Longino, pers. comm.).	en	Wild, Alexander L. (2007): A catalogue of the ants of Paraguay (Hymenoptera: Formicidae). Zootaxa 1622: 1-55, DOI: 10.5281/zenodo.179222
03C2878BFFF4FFF1F3D4D083329DFEEE.taxon	discussion	Crematogaster nitidiceps and Crematogaster victima occur in local sympatry across eastern Paraguay. There is no reason to consider the two forms conspecific, and indeed the two forms are likely not even close relatives. I have not examined type material of either form, but Jack Longino has confirmed the identification of both species in my Paraguayan material and concurs with a change in status for C. nitidiceps (Longino, pers. comm.). In Paraguay, the two species differ in a number of characters, including: antennal scapes in repose fail to meet posterior cephalic margin in C. nitidiceps but surpass posterior margin in C. victima; head in full-face view longer than broad in C. niticeps but broader than long in C. victima; sculpture of lateral face of pronotum smooth and shining in C. nitidiceps but punctate in C. victima; dorsum of head bearing only one or two pairs of standing setae in C. niticeps but bearing> 20 standing clavate setae in C. victima. The two forms show additional differences in eye size, petiolar structure, and mesosomal structure.	en	Wild, Alexander L. (2007): A catalogue of the ants of Paraguay (Hymenoptera: Formicidae). Zootaxa 1622: 1-55, DOI: 10.5281/zenodo.179222
03C2878BFFF5FFF1F3D4D79E370CFA8E.taxon	discussion	Crematogaster quadriformis is the most frequently recorded ground-nesting Crematogaster in Paraguay. This species can be diagnosed by the following combination of character states: propodeum in profile only slightly or not at all depressed below level of mesonotum; propodeal spines elongate but shorter than length of petiole; antennal scapes equal or greater in length than head length, in repose well exceeding the posterior margins of the head in full-face view; head sculpture densely punctate / striate except for a smooth frontal strip;> 20 subdecumbent to suberect setae on the first gastric tergite. Crematogaster quadriformis shows considerable variation even within nest series in overall size, pilosity, eye size, and extent of punctation / striation on the head. Jack Longino examined some recent Paraguayan collections and confirmed the identity of this species as C. quadriformis. I have subsequently viewed Forel’s types of C. quadriformis gracilior from São Paulo and found them to fall well within the range of variation I have observed in Paraguay, so I synonymize C. gracilior here. I was not able to view Forel’s types of C. quadriformis vezenyii, described from material collected in Asunción. Forel’s description notes small size, a more shiny head, and lack of a well-developed tooth in the propodeal suture. These character states are common in the C. quadriformis material I have examined towards the small end of the size distribution of nest series, so I find it likely that C. quadriformis vezenyii is a collection of a young nest or small workers of C. quadriformis.	en	Wild, Alexander L. (2007): A catalogue of the ants of Paraguay (Hymenoptera: Formicidae). Zootaxa 1622: 1-55, DOI: 10.5281/zenodo.179222
03C2878BFFF5FFF6F3D4D3BE35BBFEEE.taxon	discussion	Crematogaster rochai can be diagnosed by the following combination of characters: antennal scapes short, less than. 8 x head length, in repose failing to reach posterior margin of head in full-face view; dorsal surface of head largely devoid of erect setae; promesonotum slightly arched in profile; propodeal suture present but not deep; postero-dorsal face of propodeum forming a single concave slope, not broken into distinct dorsal and posterior faces; propodeal spines short and upturned; first gastric tergite (= abd. tergite 4) with <6 standing setae exclusive of posterior row. Mature colonies in the Crematogaster crinosa complex, including C. rochai, are polymorphic in the worker caste and occasionally produce rather large workers. Santschi’s C. malevolens seems to be little other than a large C. rochai. I can find no consistent characters to separate C. malevolens from the C. rochai type or from Paraguayan material that Jack Longino has identified as C. rochai. Longino (2003) elevated C. malevolens to species with the intent to clear trinomials from the C. crinosa complex but did not provide additional reasoning. If C. malevolans is indeed conspecific with what I call C. rochai in Paraguay, an alternate resolution would be to split these southern populations off from C. rochai under the name C. malevolens. Specimens from southern South America are more pilose and bear a stronger propodeal suture than more northerly material Longino (2003), character states that apply equally to C. malevolens. Considering the allopatric nature of the variation, however, I prefer to retain a single species.	en	Wild, Alexander L. (2007): A catalogue of the ants of Paraguay (Hymenoptera: Formicidae). Zootaxa 1622: 1-55, DOI: 10.5281/zenodo.179222
03C2878BFFF2FFF6F3D4D79E3782FC15.taxon	discussion	Wilson (2003) took a broad view of Pheidole radoszkowskii and synonymized several names, including Santschi’s variety reflexans. The evidence for resurrecting P. reflexans to species is strong, however. Both P. radoszkowskii and P. reflexans are locally sympatric across eastern Paraguay, with the former species commonly found in open habitats such as campo cerrado, roadside, and pasture, while the later is restricted to closed forests. Pheidole reflexans can be one of the most common forest-floor ants and is frequently recovered in sifted litter samples from subtropical tall forests. The morphological differences between the two forms are consistent across eastern Paraguay. Pheidole reflexans is more gracile, with considerably longer legs and antennal scapes, and in Paraguay is also somewhat more pilose in the major caste, supporting> 10 suberect setae on gastric tergite 1 (= abd. tergite 4). This pilosity difference appears to be regional, as Mayr’s P. radoszkowskii types from French Guiana (NHMW, examined) are more pilose than the Paraguayan populations of the same species.	en	Wild, Alexander L. (2007): A catalogue of the ants of Paraguay (Hymenoptera: Formicidae). Zootaxa 1622: 1-55, DOI: 10.5281/zenodo.179222
03C2878BFFF2FFF6F3D4D2243233FAFD.taxon	discussion	Solenopsis scelesta is a relatively gracile species that can be recognized by the following combination of character states: head width> 0.4 mm; compound eyes comprised of about 8 - 10 ommatidia; dorsal propodeal face about twice as long as declivitous face; dorsal face of propodeum smooth and without sculpture. Material from Paraguay is a close match to Forel’s types from São Paulo. I have not examined Emery’s type of S. decipiens, but Forel’s description of S. scelesta based on differences in thoracic structure, antennal scape length, and eye size suggests that it is unlikely that both forms are conspecific. I change the status of Solenopsis scelesta to eliminate the trinomen.	en	Wild, Alexander L. (2007): A catalogue of the ants of Paraguay (Hymenoptera: Formicidae). Zootaxa 1622: 1-55, DOI: 10.5281/zenodo.179222
03C2878BFFF2FFF7F3D4D38C328FFEA5.taxon	discussion	Solenopsis wasmannii shows a more pronounced worker polymorphism than any other Solenopsis in Paraguay. Major workers of this species can be diagnosed as follows: head width> 0.8 mm; head in full-face view chordate, wider posterior than anterior to the compound eyes, posterior margin concave; integument between compound eye and antennal insertions bearing deep piligerous punctures; compound eye small, with approximately 20 ommatidia; mesosomal dorsum in profile forming a single convexity, propodeum not depressed below level of mesonotum; propodeal suture distinct; petiole and post-petiole subequal in width; body covered in erect setae of varying length; head and body concolorous light brown to dark reddish brown in color. Minor workers are considerably smaller and lighter in color the the major but share the similarly diagnostic continuous mesosomal profile. Forel (1911 e) distinguished his subspecies S. transformis from S. wasmannii on the basis of body width, color, and punctation of the head. However, body width is a difficult character to evaluate in polymorphic species, especially with sample sizes as small as those available to Forel. In my opinion the structural differences cited by Forel are at least partly allometric in nature and not indicative of a species boundary. Likewise, a distinction based on punctation is artifactual. The cephalic punctation of the two forms is similar, but easier to see in S. transformis solely as a consequence of the slightly darker color. I find no compelling reason to retain S. transformis, so I synonymize it here.	en	Wild, Alexander L. (2007): A catalogue of the ants of Paraguay (Hymenoptera: Formicidae). Zootaxa 1622: 1-55, DOI: 10.5281/zenodo.179222
03C2878BFFF3FFF7F3D4D36C32F3F966.taxon	discussion	Forel (1908 h) distinguished the variety postangustata from H. opaciceps on the basis of having a more convex head shape, but having examined and photographed both Mayr’s and Forel’s types, I can find no discernable difference in shape. Both ants are remarkably similar in most respects except for color- the H. opaciceps type is piceous while H. postangustata is ferruginous. In any case, H. opaciceps is variable in many characters even within nest series. Forel’s H. postangustata type falls well within the bounds of variation of H. opaciceps and I synonomize it here.	en	Wild, Alexander L. (2007): A catalogue of the ants of Paraguay (Hymenoptera: Formicidae). Zootaxa 1622: 1-55, DOI: 10.5281/zenodo.179222
03C2878BFFF3FFF7F3D4D7543269FD53.taxon	discussion	Solenopsis medioclara is commonly encountered in Paraguay in sifted litter samples from tall forest habitats. This species can be recognized by the short antennal scapes (about ½ head length), the lack of punctation on the mesopleuron, and the shape of the petiole, which in profile is considerably bulkier than the post-petiole. I have not examined the type specimens of Emery’s Solenopsis brevicornis, but Santschi’s types of S. medioclara are a close match to numerous recent collections from humid subtropical semi-deciduous forest in Canindeyú. I change the status of Solenopsis medioclara to eliminate the trinomen, but detailed study of Solenopsis (Diplorhoptrum) will be needed to clarify the identity of this species. S. brevicornis may turn out to be the valid name for the Paraguayan populations.	en	Wild, Alexander L. (2007): A catalogue of the ants of Paraguay (Hymenoptera: Formicidae). Zootaxa 1622: 1-55, DOI: 10.5281/zenodo.179222
03C2878BFFF3FFF7F3D4D51B36F3FB5D.taxon	discussion	Solenopsis shiptoni can be recognized by the following combination of character states: strong transverse striae present on propodeal dorsum; compound eyes comprised of <30 ommatidia; postpetiolar node in dorsal view broader than long; head width> 0.4 mm; body color testaceous to light reddish brown. These ants are uncommonly collected in Paraguay, and all records are from the Chaco. The taxonomy of the complex of species associated with S. metanotalis is poorly understood and I change the status of S. shiptoni to eliminate the trinomen. Most of the material I have seen identified as S. metanotalis has less pronounced mesosomal striation, larger eyes, and a narrower postpetiole than S. shiptoni, but the question of whether this variation is intra- or inter-specific will require more detailed study than is possible in the present study.	en	Wild, Alexander L. (2007): A catalogue of the ants of Paraguay (Hymenoptera: Formicidae). Zootaxa 1622: 1-55, DOI: 10.5281/zenodo.179222
03C2878BFFF3FFF4F3D4D11633A4FEEE.taxon	discussion	Pseudomyrmex acanthobius, a small pallidus- group species with large eyes and a relatively swollen fore femur, has been recorded sporadically from Paraguay nesting in Acacia thorns. Emery described both P acanthobius and P. acanthobius var. fuscatus in the same paper (Emery 1896 b), distinguishing the two forms on the basis of color. The collections I have seen of this species can vary from orange-yellow to dark brown. Phil Ward, who is currently revising Pseudomyrmex, has examined Emery’s types and my recent Paraguayan collections and concluded that the color variation is intraspecific. As there are no other characters that reliably differ between the forms, I see no reason to dissent from Ward’s assessment.	en	Wild, Alexander L. (2007): A catalogue of the ants of Paraguay (Hymenoptera: Formicidae). Zootaxa 1622: 1-55, DOI: 10.5281/zenodo.179222
03C2878BFFF0FFF4F3D4D3D434ACF99E.taxon	discussion	I was unable to view Santschi’s material, but it is probable that C. insignis is conspecific with either C. scipio or C. crassus.	en	Wild, Alexander L. (2007): A catalogue of the ants of Paraguay (Hymenoptera: Formicidae). Zootaxa 1622: 1-55, DOI: 10.5281/zenodo.179222
03C2878BFFF0FFF4F3D4D79E3466FC73.taxon	discussion	Ants in the Pseudomyrmex gracilis species complex are ubiquitous in the neotropics, and the species P. gracilis itself presents a confusing array of variation (Ward 1989). Santschi distinguished his variety P. atrinodus on the basis of small size and light coloration on parts of the mesosoma, but I have examined the type material and in both these characters P. atrinodus is well within the normal range of variation of the species. Phil Ward, who is revising Pseudomyrmex, concurs with this synonymy (Ward, pers comm.)	en	Wild, Alexander L. (2007): A catalogue of the ants of Paraguay (Hymenoptera: Formicidae). Zootaxa 1622: 1-55, DOI: 10.5281/zenodo.179222
03C2878BFFF0FFF4F3D4D23B3704FAF5.taxon	discussion	Pseudomyrmex lizeri is known from a pair of localities in eastern Paraguay where it is locally sympatric with the common P. flavidulus. There is no reason to consider the two forms conspecific. In Paraguay, Pseudomyrmex lizeri is a noticeably larger ant than P. flavidulus and has a disproportionately broader head. I have not examined the type material of this species, but Phil Ward identified the Paraguayan specimens and concurs with a change in status for P. lizeri (Ward, pers comm.).	en	Wild, Alexander L. (2007): A catalogue of the ants of Paraguay (Hymenoptera: Formicidae). Zootaxa 1622: 1-55, DOI: 10.5281/zenodo.179222
