taxonID	type	description	language	source
03C2910FFF8DFFCF38DB3E50B7D9F996.taxon	type_taxon	Type species: Clivina physopleura BURGEON, 1935 by original designation.	en	Balkenohl, Michael (2018): The first species of the subgenus Physoclivina K, 1959 outside equatorial Africa (Coleoptera: Carabidae: Clivinini). Linzer biologische Beiträge 50 (1): 189-195, DOI: 10.5281/zenodo.3985398
03C2910FFF8DFFCF38DB3E50B7D9F996.taxon	description	The Subgenus Physoclivina is recognizable by the following characters: Small sized (2.5 – 3.5 mm), conspicuously cylindrical shape of the habitus, labrum six-setose, with carinae on the frons of the head in form of an inverted V, swollen pleura with the tumidity more or less visible in dorsal view, on the elytron striae one and two are free at the base, no dorsal pores on the third interval and no scutellar striole on the disc of the elytron, and at the mesotibia without a subapical seta furnished spur. At the margin of the abdominal sternum the two setigerous punctures are widely separated.	en	Balkenohl, Michael (2018): The first species of the subgenus Physoclivina K, 1959 outside equatorial Africa (Coleoptera: Carabidae: Clivinini). Linzer biologische Beiträge 50 (1): 189-195, DOI: 10.5281/zenodo.3985398
03C2910FFF8EFFCC38DB38A3B41DFAB9.taxon	description	(Fig 6)	en	Balkenohl, Michael (2018): The first species of the subgenus Physoclivina K, 1959 outside equatorial Africa (Coleoptera: Carabidae: Clivinini). Linzer biologische Beiträge 50 (1): 189-195, DOI: 10.5281/zenodo.3985398
03C2910FFF8EFFCC38DB38A3B41DFAB9.taxon	materials_examined	New records: Three specimens: Elfenbeinküste, Kafolo / Comoè 28.4.1988 leg. F. - T. Krell (MFNB / CBB); one specimen: GTZ Camp bei Kafolo / Comoè 28. IV. 1988 9 ° 37 ’ N / 4 ° 20 ’ W (NHMB); two specimen: Côto d’Ivoire, Parc Nat. Comoé, L 204 N 8 ° 43,668 ’, W 3 ° 50,637 ’ 28. VI. 95 leg. Krell, at light, 595 ft. (CBB).	en	Balkenohl, Michael (2018): The first species of the subgenus Physoclivina K, 1959 outside equatorial Africa (Coleoptera: Carabidae: Clivinini). Linzer biologische Beiträge 50 (1): 189-195, DOI: 10.5281/zenodo.3985398
03C2910FFF8EFFCC38DB38A3B41DFAB9.taxon	distribution	Occurrence: Recorded from the South and South-West of Burkina Faso as well as from the North-East of the Républic of Côte d’Ivoire.	en	Balkenohl, Michael (2018): The first species of the subgenus Physoclivina K, 1959 outside equatorial Africa (Coleoptera: Carabidae: Clivinini). Linzer biologische Beiträge 50 (1): 189-195, DOI: 10.5281/zenodo.3985398
03C2910FFF8EFFCC38DB38A3B41DFAB9.taxon	discussion	Remarks: Well described and illustrated by DOSTAL (2015). The paratype investigated exactly fits with the description.	en	Balkenohl, Michael (2018): The first species of the subgenus Physoclivina K, 1959 outside equatorial Africa (Coleoptera: Carabidae: Clivinini). Linzer biologische Beiträge 50 (1): 189-195, DOI: 10.5281/zenodo.3985398
03C2910FFF8EFFCC38DB385BB41DFC0A.taxon	distribution	Occurrence: Several localities in the West and the South-East of Senegal.	en	Balkenohl, Michael (2018): The first species of the subgenus Physoclivina K, 1959 outside equatorial Africa (Coleoptera: Carabidae: Clivinini). Linzer biologische Beiträge 50 (1): 189-195, DOI: 10.5281/zenodo.3985398
03C2910FFF8EFFCC38DB385BB41DFC0A.taxon	discussion	Remarks: Well described and illustrated by DOSTAL (2015). The paratype investigated exactly fits with the description.	en	Balkenohl, Michael (2018): The first species of the subgenus Physoclivina K, 1959 outside equatorial Africa (Coleoptera: Carabidae: Clivinini). Linzer biologische Beiträge 50 (1): 189-195, DOI: 10.5281/zenodo.3985398
03C2910FFF8EFFCC38DB3B9BB421FD42.taxon	description	(Figs 1, 5)	en	Balkenohl, Michael (2018): The first species of the subgenus Physoclivina K, 1959 outside equatorial Africa (Coleoptera: Carabidae: Clivinini). Linzer biologische Beiträge 50 (1): 189-195, DOI: 10.5281/zenodo.3985398
03C2910FFF8EFFCC38DB3B9BB421FD42.taxon	distribution	Occurrence: Confirmed with many records from the Democratic Republic of the Congo but also from the South of Sudan and Zambia.	en	Balkenohl, Michael (2018): The first species of the subgenus Physoclivina K, 1959 outside equatorial Africa (Coleoptera: Carabidae: Clivinini). Linzer biologische Beiträge 50 (1): 189-195, DOI: 10.5281/zenodo.3985398
03C2910FFF8EFFCC38DB3B9BB421FD42.taxon	discussion	Remarks: Two times the subgenus was treated as genus (BASILEWSKY 1967, 1969) but without genus description or taxonomic statement. Physoclivina might indeed deserve genus level due to some very specific characters. But at the current point in time I refrain from taxonomic action. The holotype and 19 paratypes located in MRACT have been investigated as well as specimens from the author’s collection. The holotype served for the pictures provided. To complete the series of habitus-pictures for all member of the subgenus, a picture has been added to this contribution as well.	en	Balkenohl, Michael (2018): The first species of the subgenus Physoclivina K, 1959 outside equatorial Africa (Coleoptera: Carabidae: Clivinini). Linzer biologische Beiträge 50 (1): 189-195, DOI: 10.5281/zenodo.3985398
03C2910FFF8EFFCB38DB3E6EB209FEA6.taxon	description	(Figs 2 - 4, 7 - 9)	en	Balkenohl, Michael (2018): The first species of the subgenus Physoclivina K, 1959 outside equatorial Africa (Coleoptera: Carabidae: Clivinini). Linzer biologische Beiträge 50 (1): 189-195, DOI: 10.5281/zenodo.3985398
03C2910FFF8EFFCB38DB3E6EB209FEA6.taxon	materials_examined	Type material: Holotype: ♂, with labels and data: blue, printed " NAMIBIA-Exp. ZMB 1992 Kavango: Gelukkie Kavango-Ufer, 18 ° 03 ’ S / 21 ° 08 ’ E. 1. III. 92 leg. M. Uhlig " (MFNB). Paratypes: 5 ♀♀: same data as holotype (MFNB, CBB).	en	Balkenohl, Michael (2018): The first species of the subgenus Physoclivina K, 1959 outside equatorial Africa (Coleoptera: Carabidae: Clivinini). Linzer biologische Beiträge 50 (1): 189-195, DOI: 10.5281/zenodo.3985398
03C2910FFF8EFFCB38DB3E6EB209FEA6.taxon	etymology	Etymology: The species is dedicated to Dr. Alexander Dostal (Vienna) and is expressed as Latinised adjective.	en	Balkenohl, Michael (2018): The first species of the subgenus Physoclivina K, 1959 outside equatorial Africa (Coleoptera: Carabidae: Clivinini). Linzer biologische Beiträge 50 (1): 189-195, DOI: 10.5281/zenodo.3985398
03C2910FFF8EFFCB38DB3E6EB209FEA6.taxon	diagnosis	Diagnosis: A medium sized, cylindriform species, with a row of six short but big and a second row with smaller setae arising from tubercle-like punctures at the mesotibia, and small clypeal wings separated from middle part by small obtuse notches. Distinguished from the other species of the subgenus by the flattened eyes (dorsal view) and the reduced shape of the eye in lateral view, and by the supraantennal plates which are as broad as the eyes. Moreover, the shape of the median lobe of the aedeagus and the structure of the internal sac is different.	en	Balkenohl, Michael (2018): The first species of the subgenus Physoclivina K, 1959 outside equatorial Africa (Coleoptera: Carabidae: Clivinini). Linzer biologische Beiträge 50 (1): 189-195, DOI: 10.5281/zenodo.3985398
03C2910FFF8EFFCB38DB3E6EB209FEA6.taxon	description	Description: Measurements: Body length 2.85 – 2.95 mm (ẍ = 2.92 mm *), width 0.7 – 0.72 mm (ẍ = 0.71 mm *), ratio length / width of pronotum 1.19 – 1.24 (ẍ = 1.21 *), ratio length / width of elytra 2.26 – 2.42 (ẍ = 2.32 *); (* n = 6). Colour: Shiny. Red-brown. Legs, antennae and mouthparts yellow, front femur brown, apex and margin of mandible darker, supraantennal plates translucent yellowish. Head (Figs 3, 4): Less than a quarter narrower than pronotum. Clypeus straight anteriorly, wings small, not as advanced as clypeus, separated from clypeus by small obtuse notches; supraantennal plates wide, margins convex; clypeus, wings, and supraantennal plates with small reflexed margin. Supraantennal plates with carina posteriorly extending up to mideye level, anteriorly separated from wings by obtuse-angled notches. Frons with distinct carina in form of an inverted " V ", prolonged anteriorly as keel, with second but curved carina anteriorly, with some longitudinal wide rugae between carinae and on posterior part of frons. Supraorbital carinae relatively short, separated from eyes by supraorbital furrows, each with two supraorbital setae. Neck laterally with few punctures of moderate size. Eyes distinctly flattened, as high as supraantennal plates, shape in lateral view is remindful of a truncated pyramid. Genae indistinct. Antenna short, just reaching mid-length of pronotum, segments five to ten wider than long. Labrum straight anteriorly, with irregular reticulation, five-setose. Mandible moderately curved, also towards apex, arcuate at apex, finely margined in apical half. Apical segment of maxillary palpomere securiform, pointed at apex, terminal segment of labial palpomere bottle like, slender apically. Mentum with median tooth developed a pointed slender knob, not as projecting anteriorly as lateral lobes. Lateral lobes finely carinate, surface with irregular reticulation. Pronotum: Slightly convex on disc (lateral view). Cylindrical, longer than wide. Lateral margin nearly straight in dorsal view, sigmoid in lateral view. Anterior angles rounded off by reflexed lateral margin. Reflexed lateral margin smooth. Lateral channel narrow, slightly widened at level of anterior setigerous punctures. Anterior and posterior setigerous punctures situated in marginal channel. Posterior angle not visible. Pleura visible in dorsal view. Anterior margin excised. Anterior transverse line distinct, not reaching anterior angles. Median line sharp, running over level of anterior transverse line but not joining, nearly reaching anterior margin. Distinctly joining basal constriction. Basal constriction moderately wide, flange with sharp carina. Disc usually without bigger punctures (in two paratypes unilaterally with few paralateral punctures), with indistinct longitudinal basal impression bilaterally. Surface with transverse wrinkles. Elytra: Cylindrical, two and a third times longer than wide. Marginal channel running over rounded humerus up to level of third stria without joining striae at base, with umbilical setigerous punctures widely interrupted at middle, thought two to three punctures are missing. Reflexed margin smooth. No humeral tooth. Striole missing, basal tubercle with setigerous puncture distinct, situated at declivity of first stria, Striae one and two free at base, all striae deep, punctuate-striate, stria one joining stria seven at apex, stria two ending free at apex, striae three and four and five and six joining at apex. Intervals distinctly convex, seventh carinate apically. No setigerous punctures on interval three. Surface shiny, with some indistinct wrinkles. Hind wings: Fully developed. Lower surface: Pleura distinctly swollen postero – laterally, with transverse wrinkles and isodiametric reticulation (160 times). Proepisternum and abdominal sternite covered with isodiametric to imbricate reticulation. Basal abdominal sternites with isodiametric reticulation, last three sternites shiny, with few microscopic punctures. Abdominal sternum with the two marginal setae widely separated. No differences observed among sexes. Legs: Protibia with strong digitation, with two big, one smaller and one minute lateral preapical denticles, surface with longitudinal reticulation, apical spine as long as spur, movable spur ensiform, acute at apex, tarsomeres sturdy. Mesotibia without setae on the inner side, dorso-laterally with row of six big and short setae arising from tubercle-like punctures, preapical seta furnished spine missing. Male genitalia (Fig 7, 8): Median lobe moderately slender, moderately arcuate in dorsal view, slightly more curved at middle, nearly straight in lateral view, tip of apex rounded like an elongated spoon. Surface of apical third dorsally with over a dozen of fine striae pointing apically. Internal sac at the beginning of apical third with 6 teeth pointing basally, basal sclerite distinct, V-shaped. Parameres conspicuously asymmetrical, somewhat distorted, each with one short preapical seta. Female genitalia (Fig 9): Coxostyli thin, hyaline. Terminal one like a flattened club, with two setae in the middle part. Basal one broad, velum-like. Variation: Variation noted on the pronotum which exhibits in two of the paratypes unilaterally few paralateral punctures. In other specimens the basal impression is more or less well visible, and the length of the median line on the pronotum differs after passing through the anterior transverse line.	en	Balkenohl, Michael (2018): The first species of the subgenus Physoclivina K, 1959 outside equatorial Africa (Coleoptera: Carabidae: Clivinini). Linzer biologische Beiträge 50 (1): 189-195, DOI: 10.5281/zenodo.3985398
03C2910FFF8EFFCB38DB3E6EB209FEA6.taxon	distribution	Distribution: Known from the banks of the Kavango River in the North of Namibia.	en	Balkenohl, Michael (2018): The first species of the subgenus Physoclivina K, 1959 outside equatorial Africa (Coleoptera: Carabidae: Clivinini). Linzer biologische Beiträge 50 (1): 189-195, DOI: 10.5281/zenodo.3985398
