identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
03C2A127FFAFF34F6380FF658243EFF9.text	03C2A127FFAFF34F6380FF658243EFF9.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Choroterpes (Dilatognathus) bogori	<div><p>Choroterpes (Dilatognathus) bogori sp.n.</p><p>(Figs 16–24, 29, 31, 34, 35, 38, 39–43)</p><p>= " Choroterpides exiguus " sensu Ulmer 1939 (non Thraulus exiguus Eaton 1884); = Dilatognathus/g(1) sp.2: Kluge 2012.</p><p>Material. Holotype: L-S-I♂ {No. [XXIII] (5)}, INDONESIA, JAVA, Bogor, river Ci Sadane, 3.IX.2012, coll. N. Kluge. Paratypes: the same locality and date: 4 L-S-I♂, 1 L-S♂, 4 L-S-I♀, 98 larvae: 20 L n♂, 38 L n♀, 9 L n- 1♂, 20 L n-1♀, 4 L n- 2♂, 5 L n-2♀, 1 L n- 3♂, 1 L n-3♀; river Ci Liwung in botanic garden of Bogor, 2.IX.2012: 1 L-S-I♂, 19 larvae: 5 L n♂, 9 L n♀, 1 L n- 1♂, 2 L n-1♀, 2 L n-2♀; the same locality, 24.II.2008, at light, coll. V. Ivanov: 2 I ♂, 4 S♂, 1 S♀.</p><p>Descriptions. Larva. CUTICULAR COLORATION: Cuticle of head, pronotum and mesonotum brown, with diffusive markings; cuticle of legs and abdomen light brownish, nearly unicolorous, without contrasting markings.</p><p>HYPODERMAL COLORATION: General color light brown. Head and thorax with dark maculae. Legs light ocher; each femur with large brown macula or band at middle, sometimes with smaller brown macula at apex, which is separated from middle macula (as in imago, Fig. 31). Abdominal terga with brown pigmentation, median-anterior part of each tergum ocher (as in imago, Fig. 29). Dorsal lamella of each tergalius light brown, with main trachea bordered by dark brown and side tracheae poorly visible; ventral lamella lighter brownish, with lighter trachea.</p><p>SHAPE AND SETATION: Labrum as in Ch. (D.) cataractae (Kluge 2012: Fig. 133): median incision very shallow, with shallow convexity in middle and 5 shallow denticles ventrad of it (Fig. 24); proximal transverse row of setae (shifted proximally, as in all Dilatognathus) interrupted medially; instead of distal transverse row, there is a wide stripe of irregularly situated setae, smaller than setae of the proximal transverse row; most part of dorsal surface with irregularly situated long and thin setae, somewhat smaller than setae of the distal stripe. Maxilla without dentiseta at least beginning from instar L n-3; with maxillary tusk at least beginning from instar L n-3; without ventro-apical flange beginning from instar L n-2. In both sexes maxilla of pen-pen-penultimolarva (L n-3) with wellexpressed maxillary tusk; ventro-apical flange either small (Fig. 20) or smoothed out (Fig. 19). In next instars (from L n-2 to L n) ventro-apical flange not expressed, and maxillary tusk becomes progressively longer at each instar (Fig. 18). Three vestiges of pectinate setae present in all instars. Legs as in Ch. (D.) cataractae (Kluge 2012: Figs 147–151): Femur of fore leg widest proximally, femora of middle and hind legs widest at middle. Femora with irregularly situated stout setae of various lengths; dorsal side of fore femur with few setae, dorsal side of middle and hind femora with many setae. Ventral side of hind femur with irregular row of curved bipectinate setae. Stout pointed bipectinate setae on inner side of fore tibia situated densely and irregularly. Inner side of middle and hind tibia with sparse row of stout setae; dorsal side of middle tibia with row of longer setae; outer side of hind tibia with stout setae of variable length, as on outer side of femur. Outer sides of all tibiae with irregularly situated thin hairs, whose length exceed tibia width. Abdominal segments VIII and IX with posterolateral spines short and obtuse. In all instars including ultimolarva, hind margins of abdominal terga I–II without regular row of denticles (Fig. 34); terga III–VII with row of very small, thin, spine-like denticles only in middle part (Fig. 35); terga VIII–X with row of somewhat longer denticles; sterna lack denticles. Tergalii II–VII with apical processes gradually narrowed toward apex (Fig. 23); tergalii II–V of equal size and shape, tergalius VI smaller, tergalius VII smallest and lacks costal process of dorsal lamella; anal-proximal projection of ventral lamella poorly expressed on tergalius II, better developed on tergalii III–VII. Protogonostyli of male larva short and narrowly separated; protopenes convergent, completely hidden under sternum XI; female larva has hind margin of abdominal sternum IX slightly incised [as in Ch. (D.) cataractae] (Kluge 2012: Figs 143, 144).</p><p>Subimago. CUTICULAR COLORATION: Cuticle of thorax at most light with brown sclerites; mesonotum at most brown; most part of medioscutum and submedioscutum brown, posterior scutal protuberance brown (Fig. 38). Cuticle of legs light, with diffusive brownish longitudinal stripes. Wings light brownish. Cuticle of all abdominal terga and sterna colorless; cuticle of gonostyli light brownish.</p><p>HYPODERMAL COLORATION: Abdomen and femora with brown coloration as in imago.</p><p>TEXTURE: On each leg 1st tarsomere (shortened and fused with tibia) covered by microtrichiae of the same shape as microtrichiae on tibia; 2nd–5th tarsomeres covered by pointed microlepides.</p><p>Imago, male. Head light brown with ocher; dorsal eyes not elevated, brownish-orange. Thorax light brown with ocher. Fore and hind wings with membrane nearly colorless, evenly tinged with light ocher; veins from light brown to colorless; pterostigma with 9–15 simple cross veins. Hind wing with blunt costal projection. Femora of all legs light ocher-brown with contrasting dark brown band in middle (Fig. 31). On fore leg tibia and tarsus entirely light brown. On middle and hind leg tibia lighter, with brown base (Fig. 31). Abdominal terga II–VII in anterior-median part light ocher, in posterior and lateral parts brown with ocher blanks (Fig. 29). Abdominal sterna I–VIII light. Abdominal sternum IX and styliger ocher with brown, gonostyli light brown. Styliger with nearly straight posterior margin. 1st+2nd segments of gonostylus with inner margin angulated. Penis lobes apically pressed together; each lobe with apical projection moderately long, conic, slightly bent medially-dorsally, rounded apically; denticles absent, instead of them few round concavities on dorsal side of penis close to apex (Fig. 21). Caudalii light brown, with dark brown ring at each joining.</p><p>Imago, female. Coloration as in male. Posterior plate of abdominal sternum IX apically slightly incised.</p><p>Egg. Oval, 0.15 mm length. Chorion with evenly dispersed papillae and ridges going from one papilla to another; papillae small, on one pole larger (Fig. 39–43).</p><p>Dimension. Size of both sexes greatly varies: fore wing length 5–8 mm.</p><p>Distribution. Java.</p><p>Habitat. Larvae were collected in rivers Ci Sadane and Ci Liwung at places with stony bottom and fast current. This species is most abundant upstream of the city, where water pollution is moderate; but is also present in center of Bogor (Ci Liwung in botanic garden), where the river is extremely polluted. In this respect, Ch. (D.) bogori markedly differs from Ch. (D.) cataractae and Ch. (D.) major, whose larvae were found only in selected areas of clean rivers under certain waterfalls.</p><p>Diagnosis. Male and female imagoes differ from Ch. (D.) cataractae by absence of brown coloration in proximal part of subcostal field of fore wing. Larva of penultimate and previous instars differs from Ch. (D.) cataractae by structure of maxilla (compare Figs 19, 20 and 6–13): pen-pen-penultimolarva of both sexes differs by presence of maxillary tusk and absence of dentiseta; pen-penultimolarva of both sexes differs by absence of dentiseta and ventro-apical flange; male penultimolarva differs by absence of dentiseta and ventro-apical flange. Imagoes, subimagoes and larvae differ from Ch. (D.) major by light femora with well-expressed dark band at middle (compare Figs 31 and 33). Imagoes, subimagoes and larvae differ from Ch. (D.) major and Ch. (D.) cataractae by light median part of abdominal terga (compare Figs 29 and 30). Larva of last instar differs from Ch. (D.) cataractae and Ch. (D.) major by absence of regular row of spines on posterior margins of abdominal terga I and II and by smaller spines on abdominal terga III–IX (compare Figs 34–35 and 36–37). Larva differs from Ch. (D.) nigella in shape of labrum, and from Ch. (D.) minor in shape of tergalii. Structure of egg (Fig. 39) is the same as in Ch. (D.) cataractae and differs from Ch. (D.) nigella, whose papillae have equal size on all egg surface (Kang &amp; Yang 1994: Fig. 12); eggs of other species are unknown.</p></div>	https://treatment.plazi.org/id/03C2A127FFAFF34F6380FF658243EFF9	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Kluge, Nikita J.	Kluge, Nikita J. (2014): Indonesian species of Dilatognathus Kluge 2012 (Ephemeroptera, Leptophlebiidae, Choroterpes s. l.) and species-specific sexual dimorphism in development of maxilla. Zootaxa 3786 (1), DOI: 10.11646/zootaxa.3786.1.2
03C2A127FFA8F3446380FB418312E9BE.text	03C2A127FFA8F3446380FB418312E9BE.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Choroterpes (Dilatognathus) cataractae Kluge 2012	<div><p>Choroterpes (Dilatognathus) cataractae Kluge 2012</p><p>(Figs 1–15, 30, 32, 36, 37)</p><p>Material (holotype and paratypes). INDONESIA, LOMBOK, Senaru, under Sendang Gila waterfall, 19– 26.IX.2009, coll. N. Kluge &amp; L. Sheyko: 4 L-S-I♂, 10 L-S-I♀, 1 L-S♂, 1 L-S♀, 52 larvae and larval exuviae: 12 L n♂, 22 L n♀, 1 L n- 1♂ (at molting to L n), 11 L n-1♀, 1 L n- 2♂, 4 L n-2♀, 1 L n-3♀.</p><p>Additional characteristics. Larva, subimago, imago and eggs are described by Kluge (2012). Apex of penis, hypodermal coloration of abdomen and legs and denticles on posterior margin of larval terga are figured here (Figs 1, 30, 32).</p><p>Structure of maxillary apex in penultimolarva and previous larval instars differs in males and females (Figs 6– 13). In pen-pen-penultimolarva (L n-3) ventro-apical flange and dentiseta are present, and tusk is not expressed (Fig. 13); this structure is observed on a single female larva, and probably is equal in both sexes. In male, larva of next two instars (L n-2 and L n-1) gets a short tusk and retains ventro-apical flange and dentiseta (Figs 7, 8) (this was observed on one larva of each instar).When male larva molts from penultimolarva (L n-1) to ultimolarva (L n), its maxillary structure is greatly changed (Fig. 14). In female, such change takes place earlier. In female penpenultimolarva (L n-2) maxilla has a long tusk and diminished ventro-apical flange; dentiseta is present (Fig. 12), but can be vestigial and nearly indistinguishable among setae (Fig. 11). In female penultimolarva (L n-1) maxilla has no ventro-apical flange and no dentiseta (Fig. 10); it is similar to maxilla of ultimolarva, but shorter (Fig. 15).</p><p>Dimension. Fore wing length 7.5– 8 mm.</p><p>Distribution. Lombok Island.</p><p>Habitat. All larvae were collected in a single place, under Sendang Gila waterfall. We did not find larvae of this species in other parts of the same river.</p></div>	https://treatment.plazi.org/id/03C2A127FFA8F3446380FB418312E9BE	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Kluge, Nikita J.	Kluge, Nikita J. (2014): Indonesian species of Dilatognathus Kluge 2012 (Ephemeroptera, Leptophlebiidae, Choroterpes s. l.) and species-specific sexual dimorphism in development of maxilla. Zootaxa 3786 (1), DOI: 10.11646/zootaxa.3786.1.2
03C2A127FFA2F34D6380FB0A808FEDF2.text	03C2A127FFA2F34D6380FB0A808FEDF2.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Choroterpes (Dilatognathus) major (Ulmer 1939) Ulmer 1939	<div><p>Choroterpes (Dilatognathus) major (Ulmer 1939)</p><p>(Figs 27, 33)</p><p>= Choroterpides major Ulmer 1939</p><p>= Dilatognathus/g(1) sp.1: Kluge 2012</p><p>Material. INDONESIA, JAVA, Cibodas (= Tjibodas), pool under waterfall at southern side of golf field, 6– 11.VIII. 2009, coll. N. Kluge &amp; L. Sheyko: 25 larvae: 2 L/S♂, 1 L n♂, 6 L n♀, 2 L n- 1♂, 5 L n-1♀, 2 L n-2♀, 3 L n-3♀, 4 L n- 4 ♀.</p><p>Additional descriptions. Larva. Described by Ulmer (1939). CUTICULAR COLORATION: Cuticle of head, pronotum and mesonotum brown, with diffusive markings; cuticle of legs and abdomen light brownish, nearly unicolorous, without contrasting markings.</p><p>HYPODERMAL COLORATION: General color dark blackish-brown. Head and thorax with dark maculae; labrum and exposed areas of mandibles dark brown; palps and other mouth parts ocher. Each leg with coxa, trochanter, femur and tibia dark brown; tarsus and extreme apex of tibia ocher (Fig. 33). Abdominal terga nearly entirely dark brown, with pair of small submedian blanks adjacent to anterior margin; abdominal sterna ocher with brown, variable. Tergalii dark brown; dorsal lamella darker, with main trachea of the same color and side tracheae nearly invisible; ventral lamella lighter, with main trachea of the same color and side tracheae poorly visible.</p><p>SHAPE AND SETATION. Maxilla without dentiseta at least beginning from instar L n-4; with maxillary tusk at least beginning from instar L n-4; without ventro-apical flange beginning from instar L n-3. At least female maxilla of penpen-pen-penultimolarva (L n-4) with well-expressed maxillary tusk; ventro-apical flange small (similar to pen-penpenultimolava of Ch. bogori: Fig. 20). In next instars (from L n-3 to L n) ventro-apical flange not expressed, and maxillary tusk becomes progressively longer at each instar. Three vestiges of pectinate setae present in all instars. Other characters as in Ch. (D.) cataractae (Kluge 2012) . In contrast to Ch. (D.) bogori, tergalii II–VII with apical processes small and thread-like (Fig. 27).</p><p>Subimago (extracted from mature larva). CUTICULAR COLORATION: Cuticle of thorax at most light with brown sclerites; mesonotum at most brown; most part of medioscutum and submedioscutum brown, posterior scutal protuberance brown (as in Fig. 38). Wings brown.</p><p>HYPODERMAL COLORATION: General color dark blackish-brown. Femora entirely dark brown; tibiae dark brown with light apex; fore tarsus dark brown with light articulations; middle and hind tarsi light, shaded with brown. Abdominal terga and sterna brown with small paired blanks.</p><p>TEXTURE: On each leg 1st tarsomere (shortened and fused with tibia) covered by microtrichiae of the same shape as microtrichiae on tibia; 2nd–5th tarsomeres covered by pointed microlepides.</p><p>Imago, male. Described by Ulmer (1939). Not examined.</p><p>Imago, female. Unknown.</p><p>Dimension. Length of ultimolarva 7–9 mm.</p><p>Distribution. Java; reported also from Sumatra (Ulmer 1939).</p><p>Habitat. All larvae examined by me, were collected in a single place, under waterfall which falls from the forest in national park. We did not find larvae of this species in other places, including neighboring waterfalls in the national park and the botanic garden. The place of larval inhabitance was so small, that we were unable to collect enough specimens to rear imagoes. In this respect Ch. (D.) major markedly differs from Ch. (D.) bogori, whose larvae inhabit polluted rivers.</p></div>	https://treatment.plazi.org/id/03C2A127FFA2F34D6380FB0A808FEDF2	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Kluge, Nikita J.	Kluge, Nikita J. (2014): Indonesian species of Dilatognathus Kluge 2012 (Ephemeroptera, Leptophlebiidae, Choroterpes s. l.) and species-specific sexual dimorphism in development of maxilla. Zootaxa 3786 (1), DOI: 10.11646/zootaxa.3786.1.2
03C2A127FFA1F34D6380FC2482F0EB90.text	03C2A127FFA1F34D6380FC2482F0EB90.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Choroterpes (Dilatognathus) minor (Dang 1967) Dang 1967	<div><p>Choroterpes (Dilatognathus) minor (Dang 1967)</p><p>= Choroterpides minor Dang 1967</p><p>= Dilatognathus/g(1) sp.4: Kluge 2012</p><p>Material. CHINA, HAINAN, Miao-Li, 19.XII. 1957, coll. N. Andrianova: 4 larvae: L/S♂, L n♂, L n♀, L n- 1 ♀ (ready to molt to L n).</p><p>Additional characteristics. Larva. Differs from all other species of Dilatognathus by shape of tergalii, which are relatively long and narrow (Fig. 28). Median incision of labrum shallow, with shallow median convexity and five or three shallow denticles [similar to Ch. (D.) cataractae, Ch. (D.) major and Ch. (D.) bogori]. Structure of maxillary apex in penultimolarva the same as in ultimolarva: with long tusk bearing 3 vestiges of ventral pectinate setae, without ventro-apical flange and without dentiseta.</p><p>Imago. Unknown.</p><p>Dimension. Length of ultimolarva 6–7 mm.</p><p>Distribution. Indochina Peninsula (Vietnam) and Hainan Island.</p></div>	https://treatment.plazi.org/id/03C2A127FFA1F34D6380FC2482F0EB90	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Kluge, Nikita J.	Kluge, Nikita J. (2014): Indonesian species of Dilatognathus Kluge 2012 (Ephemeroptera, Leptophlebiidae, Choroterpes s. l.) and species-specific sexual dimorphism in development of maxilla. Zootaxa 3786 (1), DOI: 10.11646/zootaxa.3786.1.2
03C2A127FFA1F34C6380F9C1802FEC69.text	03C2A127FFA1F34C6380F9C1802FEC69.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Choroterpes (Dilatognathus) nigella (Kang & Yang 1994) Kang & Yang 1994	<div><p>Choroterpes (Dilatognathus) nigella (Kang &amp; Yang 1994)</p><p>= Choroterpides nigella Kang &amp; Yang 1994 = Dilatognathus/g(1) sp.3: Kluge 2012</p><p>Material. THAILAND: province Mae Hong Son, river Namlang near Lot Cave, 12.I.1998, coll. V. Ivanov: 17 larvae; province Muang Pai, river Pai, 11.I.1998, coll. V. Ivanov: 7 larvae; SW Chiang Mai, Doi-Ithanon, river Klang, 17.I.1998, coll. Grigorenko: 1 larva; national park Kaeng Krachan, river Pchetchabun, 26.II.2006, coll. M. Chertoprud: 4 larvae. CHINA, HAINAN, Miao-Li, 19.XII.1957, coll. N. Andrianova: 2 larvae (one of them with developing cuticle of next instar).</p><p>Additional characteristics. Larva. Described by Kang &amp; Yang (1994). Differs from all other species of Dilatognathus by shape of labrum, whose median incision is deep, semicircular and bordered by sharp semicircular impression on dorsal surface (Kang &amp; Yang 1994: Fig. 3A). Structure of maxillary apex in penultimolarva is the same as in ultimolarva: with long tusk bearing 3 vestiges of ventral pectinate setae, without ventro-apical flange and without dentiseta.</p><p>Imago. Unknown.</p><p>Egg. Described by Kang &amp; Yang (1994).</p><p>Dimension. Length of ultimolarva 6–10 mm.</p><p>Distribution. Indochina Peninsula (Thailand), Hainan and Taiwan Islands.</p><p>Discussion. Kang &amp; Yang (1994) described Choroterpides nigella based on larvae and eggs. They did not compare Ch. nigella with Ch. minor . As characters distinguishing Ch. nigella from Ch. major, only two ones are named: (1) transversal curved row of long setae on labrum and (2) tooth-like maxillary tusk with three comb-like setae at its base. Actually, both characters are present in all Dilatognathus, including Ch. (D.) major (Ulmer 1939: Figs 262, 265).</p></div>	https://treatment.plazi.org/id/03C2A127FFA1F34C6380F9C1802FEC69	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Kluge, Nikita J.	Kluge, Nikita J. (2014): Indonesian species of Dilatognathus Kluge 2012 (Ephemeroptera, Leptophlebiidae, Choroterpes s. l.) and species-specific sexual dimorphism in development of maxilla. Zootaxa 3786 (1), DOI: 10.11646/zootaxa.3786.1.2
