taxonID	type	description	language	source
03C3879CFF8BFF8F0484E367FBD7C5F6.taxon	materials_examined	Type species. Ceradocus orchestiipes Costa, 1853, monotypy. Emended diagnosis. Body, pleonites with dorsal teeth or smooth, pleonal epimeron 3 usually serrate, urosomites usually with dorsal teeth or processes. Head, rostrum small to absent, lateral cephalic lobe rounded or extended, antennal sinus often forming notch; eyes oval. Antennae elongate; antenna 1, peduncular articles 1 – 2 longer than article 3, primary flagellum as long as peduncle, accessory flagellum long or short; antenna 2 shorter than antenna 1, gland cone very large. Mandible, palp article 1 with acute distal tooth, article 2 longest, article 3 very short, not falcate. Lower lip with inner lobes. Maxilla 1, inner plate triangular, with medial setae; palp 2 - articulate. Maxilla 2, inner plate with medial setae and dense oblique row of facial setae. Coxae ordinary length to short, poorly setose; coxa 1 produced anteroventrally, posteroventral margin without teeth or processes; coxae 5 – 6 lobate; gills present on coxae 2 – 6, oostegites of female present on coxae 2 – 5. Gnathopod 1 small, subchelate; carpus elongate, unlobed; propodus ovate to rectangular, palm oblique to transverse, short. Gnathopod 2 subchelate; male gnathopod 2 enlarged, symmetrical or asymmetrical, carpus short, lobate, propodus large, ovate to subrectangular or trapezoidal, palm oblique to transverse, short to long, smooth or sculptured, dactylus outer margin with 0 – 1 seta; female gnathopod 2 generally smaller, symmetrical, carpus slightly elongate, palm oblique and poorly defined, unsculptured. Pereopods, dactyli simple; bases of pereopods 5 – 7 scarcely expanded, lobate, often sharply, posterior margins weakly serrate. Uropods biramous; uropod 1 peduncle with basofacial robust seta, rami of uropods 1 – 2 extending evenly, with marginal robust setae; uropod 3 extending well beyond tips of uropods 1 – 2, peduncle slightly elongate, rami long, subequal in length, lanceolate to weakly paddle-shaped, with marginal robust setae. Telson deeply cleft, both lobes pointed distally, lateral margins straight to slightly convex, usually with 1 – 2 or more long subdistal robust setae. Included taxa (43 species and 2 subspecies). Ceradocus adangensis Wongkamhaeng & Boonyanusith, 2016; C. alama Myers & Nithyanandan, 2016; C. andamanensis Wongkamhaeng, Coleman & Pholpunthin, 2013; C. baudini Hughes, 2016; C. breweri (Kunkel, 1910);? C. capensis Sheard, 1939; C. chevreuxi Sheard, 1939; C. chiltoni Sheard, 1939; C. circe Lowry & Springthorpe, 2005; C. cotonensis Appadoo & Myers, 2006; C. crenatipalma Ledoyer, 1979; C. dooliba J. L. Barnard, 1972 a; C. greeni Appadoo & Myers, 2005; C. hawaiiensis J. L. Barnard, 1955; C. inermis Hirayama, 1986; C. isimangaliso Milne & Griffiths, 2013; C. kiiensis sp. nov.; C. koreanus Kim & Kim, 1989; C. laevis Oleröd, 1970; C. mahafalensis Ledoyer, 1979; C. mahafalensis incisa Ledoyer, 1978; C. mizani Lim, Azman & Othman, 2010; C. multidentatus Dang & Le, 2011; C. nanhaiensis Ren, 2012; C. natalensis Griffiths, 1974; C. nghisonensis Dang & Le, 2011; C. oliveri Appadoo & Myers, 2006; C. orchestiipes Costa, 1853; C. oxyodus Berents, 1983; C. paucidentatus J. L. Barnard, 1952; C. ramsayi (Haswell, 1879); C. rubromaculatus (Stimpson, 1856); C. rubromaculatus haumuri J. L. Barnard, 1972 b; C. sellickensis Sheard, 1939; C. serratus (Bate, 1862); C. setosus Dang & Le, 2011; C. sheardi Shoemaker, 1948; C. shoalsi Appadoo & Myers, 2006; C. shoemakeri Fox, 1973; C. spinicauda (Holmes, 1908); C. spinifer Ledoyer, 1973; C. tattersalli Ledoyer, 1983; C. vaderi Alves, Johnsson & Senna, 2019; C. wooree Berents, 1983; C. yandala Berents, 1983.	en	Ariyama, Hiroyuki (2019): Two species of Ceradocus collected from coastal areas in Japan, with description of a new species (Crustacea: Amphipoda: Maeridae). Zootaxa 4658 (2): 297-316, DOI: 10.11646/zootaxa.4658.2.5
03C3879CFF8AFF880484E1FFFC7BC5F2.taxon	description	(Figs 2 – 6)	en	Ariyama, Hiroyuki (2019): Two species of Ceradocus collected from coastal areas in Japan, with description of a new species (Crustacea: Amphipoda: Maeridae). Zootaxa 4658 (2): 297-316, DOI: 10.11646/zootaxa.4658.2.5
03C3879CFF8AFF880484E1FFFC7BC5F2.taxon	materials_examined	Type material. Holotype: male, 7.4 mm (OMNH-Ar- 11271), Ena, Yura Town, Wakayama Prefecture, 33 ° 59 ' 36 " N, 135 ° 06 ' 13 " E (Fig. 1 A), 0.5 m depth, stones and coarse sand bottom, 21 August 2017, coll. H. Ariyama. Paratypes: male, 4.3 mm (OMNH-Ar- 11272), same data as holotype; male, 5.6 mm (OMNH-Ar- 11273), Kuroshima Island, Yura Town, Wakayama Prefecture, 34 ° 00 ' 12 " N, 135 ° 05 ' 30 " E (Fig. 1 A), 4 m depth, coarse sand bottom, 2 August 2008, coll. H. Ariyama; female, 6.6 mm (OMNH-Ar- 11274), same data as paratype, male, 5.6 mm (OMNH-Ar- 11273) except for habitat, 1 m depth, under stones. Type locality. Ena, Yura Town in Wakayama Prefecture, Japan. Etymology. Referring to the type locality. “ Kii ” is an old name of Wakayama Prefecture. Diagnosis. Male gnathopod 2 asymmetrical; larger male gnathopod 2 with strongly oblique palm having single shallow excavation and never having defining cusp; smaller male gnathopod 2, palm oblique, smooth, defined by tiny projection. Pereopods 5 – 7, bases with acute posterodistal lobe. Pleonites 1 – 3 with many dorsal teeth, epimera 1 – 3 each bearing posterior, posteroventral and ventral teeth. Urosomites 1 – 3 with 9, 4, 2 dorsal teeth, respectively. Uropod 3 enlarged, both rami wide. Telson, each lobe with 1 long and 1 – 2 short robust setae. Description. Based on holotype, male, 7.4 mm (OMNH-Ar- 11271), and paratype, female, 6.6 mm (OMNH- Ar- 11274) for upper lip. Head (Figs 2 – 3). Eyes relatively large, about 0.3 times as long as head. Antenna 1 about 0.6 times as long as body, poorly setose; peduncle with ratio of lengths of articles 1 – 3 1: 1.15: 0.2, article 1 with 4 posteromedial robust setae, anterodistal corner with bundle of setae; accessory flagellum short, with 5 articles, terminal article minute, length about 0.15 times primary flagellum; primary flagellum long, with 32 articles, terminal article minute. Antenna 2 about half length of antenna 1, weakly setose, with ratio of lengths of peduncular articles 3 – 5 1: 2.2: 1.9; flagellum short, with 15 articles, terminal article minute. Upper lip, ventral margin rounded, with short setae. Mandible, each incisor bearing 4 cusps, left lacinia mobilis 4 - dentate, with 9 accessory setae, right lacinia mobilis bifid, 5 - dentate, with 8 accessory setae, each molar developed; palp articles 1 – 3 length ratio (excluding distal tooth of article 1) 1: 2.05: 0.85, articles 2 – 3 weakly setose. Lower lip, distal margins of inner and outer lobes setose, mandibular processes small. Maxilla 1, inner plate heavily setose on mediodistal margin, outer plate with 9 apical robust setae, palp with many apical setae. Maxilla 2, outer plate same size as inner plate; outer plate bearing many apical setae, inner plate heavily setose on mediodistal margin and dorsomedial surface. Maxilliped, inner plate truncate, distally setose, distomedial corner with 3 robust setae ventrally; outer plate with 13 long-to-short robust setae on distomedial margin; palp 4 - articulate, articles 2 – 3 setose medioventrally, article 4 with large apical robust seta. Pereon (Fig. 4). Gnathopod 1, coxa ventral margin with several short setae; basis short, with 3 long setae on posterior margin, 5 setae on posteromedial surface and 3 thick setae on posterodistal margin; merus setose posteriorly; carpus longish trapezoidal, posterior margin and medial surface heavily setose; propodus longish ovate, slightly shorter than carpus, anterodistal corner and posterior margin setose, palm oblique, posterior margin lined with 4 large and many short robust setae; dactylus slender, slightly curved posteriorly. Gnathopod 2 asymmetrical, carpus – dactylus different between right and left; coxa subquadrate, ventral margin with several setae, gill large, roundish, subequal to basis length; basis posterior margin bearing 2 long and 2 short setae, lateral surface with anterodistal ridge; merus with acute posterodistal spine; right carpus triangular, lobate and setose posterodistally; right propodus longish ovate, about 3.9 times as long as carpus, length about twice of width, palm strongly oblique, posterior margin smooth except for distal shallow excavation, with many marginal and medio-submarginal robust setae; right dactylus stout, curved posteriorly, inner margin with hollow proximally; left carpus longer than right one, heavily setose posterodistally; left propodus slender, about 1.7 and 0.65 times as long as carpus and right propodus, respectively, length about 2.1 times width, anterodistal corner and posterior margin weakly setose, palm oblique, defined by tiny projection with robust seta, posterior margin bearing many marginal and 4 long medio-submarginal robust setae; left dactylus slender. Pereopods 3 – 4 subequal in shape, pereopod 3 about 1.1 times length of pereopod 4; coxae subtrapezoidal, anterior and ventral margins with a few short setae, ventromedial surfaces bearing long seta, gills large; bases rectilinear, posterior margins with 5 long setae; meri – propodi with marginal robust and slender setae; dactyli short. Pereopod 5 about 1.35 times length of pereopod 4; coxa with robust seta on posteroventral margin; basis subrectangular, length about 1.9 times width (excluding posterodistal lobe), posterodistal corner sharply projected, anterior margin bearing long and short robust setae, lateral surface with ridge; merus – propodus with marginal robust setae, dactylus short. Pereopod 6 about 1.3 times as long as pereopod 5; coxa with 4 robust setae on anteroventral margin; basis subrectangular, length about 1.75 times width (excluding posterodistal lobe), posterodistal corner sharply projected, anterior margin bearing long and short robust setae, lateral surface with ridge; merus – propodus with marginal robust setae, dactylus short. Pereopod 7 slightly shorter than pereopod 6; coxa with robust seta on anteroventral margin; basis subrectangular, length about 1.85 times width (excluding posterodistal lobe), posterodistal corner sharply projected, anterior margin bearing short robust setae, lateral surface with ridge; merus – propodus with marginal robust setae, dactylus short. Pleon (Figs 2, 5). Pleonites 1 – 3 with 25, 21, 21 dorsal teeth, respectively; epimeron 1 bearing 4 posterior, 1 posteroventral and 4 ventral teeth, 2 ventral robust setae and lateral ridge; right epimeron 2 (left one abnormal) with 8 posterior, 1 posteroventral and 4 ventral teeth, 2 ventral robust setae and lateral ridge; epimeron 3 with 10 posterior, 1 large posteroventral and 5 ventral teeth and 2 ventral robust setae. Urosomite 1 bearing 9 continuous dorsal teeth, midline with 1 tooth and each dorsolateral margin with 4 teeth. Urosomite 2 with midline gape, each dorsolateral margin with 2 teeth. Urosomite 3 with wide midline gape, each dorsolateral corner bearing tiny tooth. Pleopods 1 – 3, pleopod 3 shortest; peduncles with 2 coupling hooks and 2 short setae distally, peduncles of pleopods 1 – 2 bearing a few and many setae, respectively, peduncle of pleopod 3 with many proximal setae and several lateral and distal robust setae; both rami almost same length, outer rami with 20, 20, 18 articles and inner rami with 18, 19, 17 articles respectively. Uropod 1, peduncle with large basofacial and distolateral robust setae and 8 dorsolateral and 7 dorsomedial robust setae; outer ramus slightly shorter than inner ramus, about 0.8 times as long as peduncle, with 3 lateral, 3 medial and 5 terminal robust setae; inner ramus with 6 lateral, 6 medial and 5 terminal robust setae. Uropod 2 about 0.65 times length of uropod 1; peduncle bearing 3 dorsolateral and 2 dorsomedial robust setae, distolateral corner with 2 dorsal and 1 ventral robust setae, distomedial corner bearing 2 robust setae dorsally; outer ramus about 0.9 times length of inner ramus, about 1.2 times as long as peduncle, with 5 lateral, 1 medial and 5 terminal robust setae; inner ramus with 9 medial and 5 terminal robust setae. Uropod 3 about 1.55 times as long as uropod 2; peduncle projected proximally, lateral surface with 1 dorsal, 3 dorsodistal and 2 ventrodistal robust setae, medial surface bearing 7 dorsal and 3 distal robust setae; outer ramus oblong, slightly widened distally, length about 3.45 times width, about 1.1 times as long as inner ramus, about 1.85 times length of peduncle, ventral and distal margins serrate, with many bundles of robust setae, dorsal margin weakly serrate, bearing several single robust setae; inner ramus situated dorsally on outer ramus, elliptical, length about 2.95 times width, medioproximal surface with small projection for receiving outer ramus, margins weakly serrate, ventral and lateral margins with several and many single robust setae, respectively, distal margin bearing bundle of robust setae. Telson about 1.15 times longer than wide, each medial margin bearing tiny notch in middle; each dorsal surface with 1 long and 1 – 2 short robust setae and 1 long and 1 short sensory setae, each lateral margin also bearing 1 long and 1 short sensory setae. Variation. Gnathopod 2. Female, paratype, 6.6 mm (OMNH-Ar- 11274, Fig. 5 F-G 2): symmetrical, coxa subtrapezoidal, other parts similar to smaller gnathopod 2 of male, holotype; medium-sized male, paratype, 5.6 mm (OMNH-Ar- 11273): asymmetrical, similar to both gnathopods 2 of holotype, right propodus about 0.65 times as long as left propodus; small male, paratype, 4.3 mm (OMNH-Ar- 11272): symmetrical, similar to smaller gnathopod 2 of holotype, but right propodus slightly shorter than left one. Pleonal epimera 1 – 3 (Table 1), numbers of teeth and robust setae varying with body length. Urosomites 1 – 3, numbers of dorsal teeth same as those of holotype except for that on urosomite 1 of small male, 4.3 mm (OMNH-Ar- 11272) (Table 1). Telson. Female, 6.6 mm (OMNH-Ar- 11274), each lobe with 1 long and 2 short robust setae; smaller males, 5.6 and 4.3 mm (OMNH-Ar- 11273, 11272), each lobe with 1 long and 1 short robust setae. Coloration in recently fixed specimen (Fig. 6). Eyes light brown, anterior body faintly orangish white, posterior body white, sparsely with faintly red spots. Remarks. The key in Krapp-Schickel & Vader (2009) does not lead these specimens to any species, because they have 9 dorsal teeth on the urosomite 1 whereas there are only “ more than 10 ” and “ six to eight ” in the couplet 29 of the key. Ceradocus kiiensis sp. nov. has (1) asymmetrical male gnathopod 2, (2) larger male gnathopod 2 with oblique palm having single excavation and never having defining cusp, and (3) pleonites 1 – 3 with many dorsal teeth. These three characters are shared with several Ceradocus species. Several important characters of this new species and the related species having such characters are shown in Table 2. All the related species are different from C. kiiensis in a part of the characters. For example, C. alama from Kuweit, C.? rubromaculatus sensu Lyons & Myers (1993) from Red Sea, and C. rubromaculatus sensu Monod (1937) from the Suez Canal (probably the same species) can be distinguished from C. kiiensis by the anterodistal projection on the basis of the larger male gnathopod 2. Ceradocus inermis from Japan, C. rubromaculatus sensu Ledoyer (1968) and C. tattersalli Ledoyer, 1983 both from Madagascar are different in the 0 – 1 posterior tooth each on the pleonal epimera 1 – 2. Ceradocus mizani from Malaysia and C. multidentatus from Vietnam have the deeper excavation on the palm of the larger male gnathopod 2 (Lim et al. 2010, Dang & Le 2011). The most resembling species is C. rubromaculatus sensu Ren (1998, 2012) from the Nansha Islands, South China Sea; however, this species is distinguishable from C. kiiensis in the number of the dorsal teeth on the urosomite 1 and the number of the robust setae on the telson. In addition, plural species are probably mixed in C. rubromaculatus as Lyons & Myers (1993) stated, therefore, revision of the species complex is needed. Habitat. Coarse sand bottom and under stones, 0.5 – 4 m depth. Distribution. Japan: Yura Town in Wakayama Prefecture (present study).	en	Ariyama, Hiroyuki (2019): Two species of Ceradocus collected from coastal areas in Japan, with description of a new species (Crustacea: Amphipoda: Maeridae). Zootaxa 4658 (2): 297-316, DOI: 10.11646/zootaxa.4658.2.5
03C3879CFF83FF9D0484E5D5FC0BC49D.taxon	description	(Figs 7 – 11)	en	Ariyama, Hiroyuki (2019): Two species of Ceradocus collected from coastal areas in Japan, with description of a new species (Crustacea: Amphipoda: Maeridae). Zootaxa 4658 (2): 297-316, DOI: 10.11646/zootaxa.4658.2.5
03C3879CFF83FF9D0484E5D5FC0BC49D.taxon	materials_examined	Material examined. Two males, 6.9 and 6.1 mm (OMNH-Ar- 11275, 11276), Hinai Beach, Iriomote Island, Okinawa Prefecture, 24 ° 24 ' 49 " N, 123 ° 48 ' 22 " E (Fig. 1 C), 0.5 m depth, dead coral and fine sand bottom, 14 July 2015, coll. H. Ariyama; male, 4.6 mm (OMNH-Ar- 11277) and ovigerous female, 7.4 mm (OMNH-Ar- 11278), Shirahama, Iriomote Island, Okinawa Prefecture, 24 ° 21 ' 44 " N, 123 ° 44 ' 45 " E (Fig. 1 C), 0.05 m depth, dead coral and fine sand bottom, 16 July 2015, coll. H. Ariyama; male, 6.9 mm (OMNH-Ar- 11279) and female with juveniles, 7.0 mm (OMNH-Ar- 11280), Awase, Okinawa Island, Okinawa Prefecture, 26 ° 18 ' 48 " N, 127 ° 50 ' 15 " E (Fig. 1 B), intertidal, dead coral and fine sand bottom, 3 July 2015, coll. H. Ariyama. Type locality. Puerto Garera in Mindoro, Philippines (Oleröd 1970). Diagnosis. Male gnathopod 2 asymmetrical; larger male gnathopod 2 with strongly oblique palm having triangular and small trapezoidal projections distally; smaller male gnathopod 2, palm oblique, smooth, defined by tiny projection. Pereopods 5 – 7, bases with rounded posterodistal lobe. Pleonites 1 – 3 and urosomites 1 – 3 without dorsal teeth; pleonal epimera 1 – 2 each bearing 1 posteroventral and 1 ventral teeth, epimeron 3 with 1 – 2 posterior, 1 posteroventral and 1 – 2 ventral teeth. Uropod 3 enlarged, both rami wide. Telson, each lobe with 1 long and 1 short robust setae. Description. Based on male, 6.9 mm (OMNH-Ar- 11275), and male, 6.9 mm (OMNH-Ar- 11279) for pleonal epimera. Head (Figs 7 – 8). Eyes relatively large, about quarter of head length. Antenna 1 poorly setose; peduncle with ratio of lengths of articles 1 – 3 1: 1.15: 0.2, article 1 with 4 posteromedial robust setae, anterodistal corner with bundle of setae; accessory flagellum short, with 5 articles, terminal article minute; primary flagellum long, with> 16 articles. Antenna 2 relatively short, weakly setose, with ratio of lengths of peduncular articles 3 – 5 1: 2.5: 2.05; flagellum short, with 9 articles, terminal article minute. Upper lip, ventral margin rounded, with short setae. Mandible, each incisor bearing 4 cusps, left lacinia mobilis 4 - dentate, with 7 accessory setae, right lacinia mobilis bifid, 5 - dentate, with 7 accessory setae, each molar developed; palp articles 1 – 3 length ratio (excluding distal tooth of article 1) 1: 2.3: 0.8, article 2 weakly setose, article 3 with several long setae at tip. Lower lip, distal margins of inner and outer lobes setose, mandibular processes small. Maxilla 1, inner plate setose on mediodistal margin, outer plate with 9 apical robust setae, palp with many apical setae. Maxilla 2, outer plate same size as inner plate; both plates bearing many apical setae, inner plate with dorsomedial setae. Maxilliped, inner plate truncate, distally setose, distomedial corner with 3 robust setae ventrally; outer plate with 14 long-to-short robust setae on distomedial margin; palp 4 - articulate, articles 2 – 3 setose medially, article 4 with large apical robust seta. Pereon (Fig. 9). Gnathopod 1 slender; coxa ventral margin with several short setae; basis short, with single long seta on posterior margin and several setae on posterodistal margin; merus setose posteriorly; carpus longish trapezoidal, posterior margin and medial surface heavily setose; propodus longish ovate, about 0.85 times as long as carpus, anterodistal corner and anteromedial surface setose, palm oblique, defined by single robust seta; dactylus slender, slightly curved posteriorly. Gnathopod 2 asymmetrical, carpus – dactylus different between right and left; coxa subtrapezoidal, ventral margin with several setae, gill large, oval, subequal to basis length; basis posterior margin bearing many long or short setae, lateral surface with ridge; merus with acute posterodistal spine; right carpus short, lobate and setose posterodistally; right propodus longish ovate, length about 2.1 times width, palm strongly oblique, posterior margin distally with triangular and small trapezoidal projections bearing 3 lateral and 4 medial, and 3 lateral and 2 medial robust setae, respectively, middle to proximal parts smooth, posteromedial surface with 2 large and many short robust setae; right dactylus stout, strongly curved posteriorly; left carpus much longer than right one, heavily setose posterodistally; left propodus slender, about 1.9 and 0.55 times as long as carpus and right propodus, respectively, length about twice of width, anterodistal corner and posterior margin setose, palm oblique, defined by tiny projection with long robust seta, posterior margin bearing many marginal and 2 medio-submarginal robust setae; left dactylus slender. Pereopods 3 – 4 subequal in shape, pereopod 3 about 1.1 times length of pereopod 4; coxae subtrapezoidal, ventral margins with a few short setae, gills large; bases rectilinear, anterior and posterior margins with 3 and 5 long setae, respectively; meri – propodi with marginal robust setae; dactyli short. Pereopod 5 about 1.3 times length of pereopod 4; coxa with several short setae on ventral margin; basis subrectangular, length about 1.65 times width (excluding posterodistal lobe), posteroproximal corner projected, posterodistal corner roundly lobed, anterior margin bearing short robust setae, anterodistal corner with long robust seta, lateral surface with ridge; merus – propodus with marginal robust setae, dactylus short. Pereopod 6 about 1.3 times as long as pereopod 5; coxa with 2 robust setae on anteroventral margin; basis subrectangular, length about 1.45 times width (excluding posterodistal lobe), posterodistal corner roundly lobed, anterior margin bearing short robust setae, anterodistal corner with long robust seta, lateral surface with ridge; merus – propodus with marginal robust setae, dactylus short. Pereopod 7 slightly shorter than pereopod 6; coxa with robust seta on anteroventral margin; basis subrectangular, length about 1.4 times width (excluding posterodistal lobe), posterodistal corner roundly lobed, anterior margin bearing short robust setae, anterodistal corner with long robust seta, lateral surface with ridge; merus – propodus with marginal robust setae, dactylus short. Pleon (Figs 7, 10). Pleonites 1 – 3 lacking dorsal teeth; epimeron 1 bearing 1 posteroventral and 1 ventral teeth and lateral ridge; epimeron 2 with 1 posteroventral and 1 ventral teeth, 2 ventral robust setae and lateral ridge; epimeron 3 with 1 posterior, 1 large posteroventral and 2 ventral teeth and 4 ventral robust setae. Urosomites 1 – 3 also lacking dorsal teeth. Pleopods 1 – 3, pleopod 3 shortest; peduncles with a few proximal short setae, 2 coupling hooks and 2 short setae distally; outer rami slightly shorter than inner rami, outer rami with 17, 17, 14 articles and inner rami with 16, 16, 14 articles respectively. Uropod 1, peduncle with large basofacial and distolateral robust setae and 6 dorsolateral and 6 dorsomedial (distal one large) robust setae; outer ramus slightly shorter than inner ramus, about 0.65 times as long as peduncle, with 2 lateral, 3 medial and 4? terminal robust setae; inner ramus with 3 lateral, 4 medial and 5 terminal robust setae. Uropod 2 about 0.65 times length of uropod 1; peduncle bearing 4 dorsolateral and 2 dorsomedial robust setae, mediodistal corner with 2 robust setae dorsally; outer ramus about 0.9 times length of inner ramus, slightly shorter than peduncle, with 4 lateral, 3 medial and 5 terminal robust setae; inner ramus with 2 lateral, 6 medial and 5 terminal robust setae. Uropod 3 about 2.35 times as long as uropod 2; peduncle projected proximally, lateral surface with 2 dorsodistal and 3 ventrodistal robust setae, medial surface bearing 4 dorsal and 2 distal robust setae; outer ramus oblong, a little widened distally, length about 3.5 times width, slightly longer than inner ramus, about 2.55 times length of peduncle, ventral margin serrate, with many bundles of robust setae, dorsal margin almost smooth, bearing several single robust setae; inner ramus situated dorsally on outer ramus, longish elliptical, length about 3.0 times width, medioproximal surface with ridge for restraining outer ramus, ventral margin smooth, with a few robust setae distally, dorsal margin also smooth, with several robust setae. Telson about 1.2 times longer than wide; each distal end with 1 long and 1 short robust setae and sensory seta, each lateral margin bearing 2 sensory setae. Variation. Gnathopod 2. Female, 7.4 mm (OMNH-Ar- 11278, Fig. 10 F-G 2): symmetrical, coxa longish subtrapezoidal, other parts similar to smaller gnathopod 2 of male, 6.9 mm (OMNH-Ar- 11275); medium-sized male, 6.1 mm (OMNH-Ar- 11276): asymmetrical, similar to both gnathopods 2 of male, 6.9 mm, but palmar projections of larger gnathopod 2 smaller, left propodus about 0.6 times as long as right propodus; small male, 4.6 mm (OMNH- Ar- 11277; Fig. 10 SM-LG 2, RG 2): shape symmetrical, similar to smaller gnathopod 2 of male, 6.9 mm, but right propodus about 0.8 times longer than left one. Pleonal epimera 1 – 3 (Table 3), numbers of teeth and robust setae with little variation. Telson, number of robust setae with no variation. Coloration in recently fixed specimen (Fig. 11). Eyes dark brown, body and appendages whitish, pale yellow internal organs seen through. Remarks. Morphological characters of the present specimens quite agree with the original description and figures of Ceradocus laevis from Philippines (Oleröd 1970) except for 3 posterior teeth on the epimeron 3 (1 – 2 teeth in the present specimens, see Table 3). This species closely resembles C. woorree from Australia; however, the latter species has the rounded anteroventral corner of the gnathopod 1, the narrower bases of the pereopods 5 – 6, the epimera 1 – 3 without posterior and ventral teeth, and the telson lobes each with 1 long and 2 short robust setae (Berents 1983). Habitat in Japan. Dead coral and fine sand bottom, intertidal to 0.5 m depth. Distribution. Japan: Okinawa Island and Iriomote Island in Okinawa Prefecture (present study); China: Hainan Island (Ren 2012); Philippines: Mindoro (Oleröd 1970); Vietnam: Khánh Hòa, Bà R ịa-Vũng Tàu and Kiên Giang Provinces (Dang & Le 2011); Singapore: Pulau Ubin (White 2015).	en	Ariyama, Hiroyuki (2019): Two species of Ceradocus collected from coastal areas in Japan, with description of a new species (Crustacea: Amphipoda: Maeridae). Zootaxa 4658 (2): 297-316, DOI: 10.11646/zootaxa.4658.2.5
