identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
03C087F9FFF37F124EB7FBC4FD795281.text	03C087F9FFF37F124EB7FBC4FD795281.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Danaea acuminata	<div><p>Danaea acuminata clade</p><p>This is a well-supported clade (BP = 100) of five species, all of which were described in 2001–2010: Danaea acuminata, D. falcata, D. lucens, D. vivax, and D. xenium . The species are relatively similar in general appearance: creeping rhizomes, concolorous laminae that dry dark brown, and serrate pinna apices. The other clades in Holodanaea are more heterogeneous in appearance. Geographically this clade is concentrated to the Amazonian lowlands adjacent to the Andes in Peru and Ecuador, with one species ( D. lucens) between the Cordilleras in Colombia.</p></div>	https://treatment.plazi.org/id/03C087F9FFF37F124EB7FBC4FD795281	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Keskiniva, Venni;Tuomisto, Hanna;Lehtonen, Samuli	Keskiniva, Venni, Tuomisto, Hanna, Lehtonen, Samuli (2024): Danaea (Marattiaceae) keeps diversifying, part 2: phylogeny and identification key for 81 taxa. Willdenowia 53 (3): 229-255, DOI: 10.3372/wi.53.53304, URL: https://www.mendeley.com/catalogue/e926cf5a-1684-366f-8354-946d48c7e673/
03C087F9FFF37F124EB7FA44FE7C50C1.text	03C087F9FFF37F124EB7FA44FE7C50C1.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Danaea alata Sm. Christenhusz 2711	<div><p>Danaea alata clade</p><p>This is a moderately supported clade (BP = 72) that contains species from all corners of the wet American tropics: the Atlantic coastal forests of Brazil ( Danaea ubatubensis), the Lesser Antilles and the Venezuelan coast ( D. alata), Central America ( D. wendlandii), the Pacific coast of Colombia and Ecuador ( D. gracilis), and western Amazonia ( D. lanceolata).</p><p>Danaea wendlandii, D. gracilis, and D. lanceolata form a well-supported clade (BP = 100) and are rather uniform in appearance, being rather small and having lanceolate laminae with narrow pinnae and terminal buds. Danaea alata and D. ubatubensis are clearly larg- er species, although D. ubatubensis shares the propensity to produce terminal buds. Three of the species were strongly supported as monophyletic, but D. gracilis was resolved as paraphyletic.</p></div>	https://treatment.plazi.org/id/03C087F9FFF37F124EB7FA44FE7C50C1	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Keskiniva, Venni;Tuomisto, Hanna;Lehtonen, Samuli	Keskiniva, Venni, Tuomisto, Hanna, Lehtonen, Samuli (2024): Danaea (Marattiaceae) keeps diversifying, part 2: phylogeny and identification key for 81 taxa. Willdenowia 53 (3): 229-255, DOI: 10.3372/wi.53.53304, URL: https://www.mendeley.com/catalogue/e926cf5a-1684-366f-8354-946d48c7e673/
03C087F9FFF37F124D4AFDC4FB5D54E1.text	03C087F9FFF37F124D4AFDC4FB5D54E1.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Danaea crispa	<div><p>Danaea crispa clade</p><p>This is a well-supported (BP = 100) clade containing three small-statured species from Central America that we would never have grouped together based on their morphology. One of them is unique in having simple, bicolorous laminae ( Danaea carillensis), one resembles D. humilis of the D. cuspidata clade in having many small, bicolorous pinnae ( D. pumila), and the third one is similar to D. gracilis of the D. alata clade in having dark green laminae that are so thin as to be translucent ( D. crispa). The D. crispa clade forms a well-supported clade (BP = 93) with the D. moritziana clade.</p></div>	https://treatment.plazi.org/id/03C087F9FFF37F124D4AFDC4FB5D54E1	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Keskiniva, Venni;Tuomisto, Hanna;Lehtonen, Samuli	Keskiniva, Venni, Tuomisto, Hanna, Lehtonen, Samuli (2024): Danaea (Marattiaceae) keeps diversifying, part 2: phylogeny and identification key for 81 taxa. Willdenowia 53 (3): 229-255, DOI: 10.3372/wi.53.53304, URL: https://www.mendeley.com/catalogue/e926cf5a-1684-366f-8354-946d48c7e673/
03C087F9FFF37F124D4AFF48FB6A5501.text	03C087F9FFF37F124D4AFF48FB6A5501.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Danaea excurrens	<div><p>Danaea excurrens clade</p><p>Danaea excurrens was described from the Atlantic rainforests of Brazil, and our DNA samples from that area form a strongly supported clade (BP = 100) with similarlooking material from Bolivia. The DNA samples from Brazil represent both individuals with spathulate pinnae that conform with the type of the species and individuals that have a more classic Holodanaea appearance. Therefore, we have concluded that both forms belong to the same species, even though we have not discovered any obvious reason for such heterophylly .</p></div>	https://treatment.plazi.org/id/03C087F9FFF37F124D4AFF48FB6A5501	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Keskiniva, Venni;Tuomisto, Hanna;Lehtonen, Samuli	Keskiniva, Venni, Tuomisto, Hanna, Lehtonen, Samuli (2024): Danaea (Marattiaceae) keeps diversifying, part 2: phylogeny and identification key for 81 taxa. Willdenowia 53 (3): 229-255, DOI: 10.3372/wi.53.53304, URL: https://www.mendeley.com/catalogue/e926cf5a-1684-366f-8354-946d48c7e673/
03C087F9FFF37F144D4AFC64FD5C5581.text	03C087F9FFF37F144D4AFC64FD5C5581.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Danaea moritziana	<div><p>Danaea moritziana clade</p><p>This clade consists of six species from the Caribbean region that are similar in having bicolorous laminae, large number of pinnae and clearly serrate pinna apices: Danaea moritziana (Coastal Venezuela and adjacent Colombia), D. mazeana (Lesser Antilles), D. urbanii (Greater Antilles), D. cuspidopsis (Costa Rica to northern Andes), D. jamaicensis (Greater Antilles), and D. jenmanii (Greater Antilles). They form a well-supported clade (BP = 98) with relatively small interspecific genetic differences.</p><p>The name Danaea moritziana C. Presl has traditionally been applied in a very wide sense. Our phylogenetic results suggest that the species should be circumscribed more narrowly. Our interpretation of the phylogenetic position of D. moritziana is based on material from the Caribbean coast of Colombia that morphologically matches the type of D. moritziana from Venezuela. Unfortunately, we do not have DNA from the type locality, but this interpretation leads to a morphologically coherent species whose distribution is limited to the northernmost parts of Venezuela and Colombia. Notably, none of the Mexican Holodanaea samples were resolved to the D. moritziana clade, strongly supporting the recognition of D. cuspidata as a distinct species. The situation was complicated by the fact that some Costa Rican and Colombian specimens were deeply embedded in the D. moritziana clade even though they were morphologically very similar to D. cuspidata . These have now been described as D. cuspidopsis (Keskiniva &amp; Tuomisto 2024) .</p><p>Christenhusz (2010) synonymized Danaea jamaicensis under D. mazeana, but the phylogeny shows that D. mazeana forms a clade of its own at the base of the D. moritziana clade, whereas D. jamaicensis groups with D. jenmanii . The latter two species are intermingled in the phylogeny, but we decided to keep them as separate species because of their morphological differences: D. jenmanii is generally smaller and has fewer pairs of shorter pinnae, shorter pinna apices, more scaly petioles and rachises, and the terminal pinna (or a part thereof) is replaced by a bud (vs terminal pinna usually present in D. jamaicensis). In our interpretation, D. mazeana is restricted to the Lesser Antilles, whereas D. jamaicensis and D. jenmanii occur in the Greater Antilles. The material from Lesser Antilles that we identified to D. mazeana has generally narrower pinnae and broader pinna apices, and they dry to a darker colour than D. jamaicensis .</p></div>	https://treatment.plazi.org/id/03C087F9FFF37F144D4AFC64FD5C5581	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Keskiniva, Venni;Tuomisto, Hanna;Lehtonen, Samuli	Keskiniva, Venni, Tuomisto, Hanna, Lehtonen, Samuli (2024): Danaea (Marattiaceae) keeps diversifying, part 2: phylogeny and identification key for 81 taxa. Willdenowia 53 (3): 229-255, DOI: 10.3372/wi.53.53304, URL: https://www.mendeley.com/catalogue/e926cf5a-1684-366f-8354-946d48c7e673/
03C087F9FFF57F144EB7FD44FE9D5181.text	03C087F9FFF57F144EB7FD44FE9D5181.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Danaea cuspidata	<div><p>Danaea cuspidata clade</p><p>This moderately supported (BP = 77) clade is a mixture of large and small plants, translucent and thick laminae and distributions from Mexico to Bolivia and the Guianas (including Danaea ypori, the only species in D. subg. Holodanaea with a Guianan distribution) but no species from the Caribbean islands.</p><p>Both Danaea cuspidata (from Mexico) and the newly described D. andina (from Ecuador) formed strongly supported clades within the D. cuspidata clade, confirming their distinctness from D. moritziana . However, their positions within the D. cuspidata clade varied between analyses done with different alignments and was not well resolved in any of them. In general, the species in this clade were supported by the phylogenetic results, although most of them had too few samples for their coherence to be properly tested. The morphologically relatively similar D. imbricata and D. trichomanoides were resolved to different subclades, supporting their recognition as distinct species.</p><p>Species incertae sedis</p><p>Danaea augusti H. Karst. ex Kunze D. intermedia J. Sm.</p><p>D. nodosa var. intermedia J. Sm. ex Hassl.</p><p>Excluded species</p><p>Danaea evecta Spreng. = Angiopteris evecta (G. Forst.) Hoffm.</p><p>D. paleacea Raddi = Bolbitis serratifolia (Mertens) Schott</p></div>	https://treatment.plazi.org/id/03C087F9FFF57F144EB7FD44FE9D5181	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Keskiniva, Venni;Tuomisto, Hanna;Lehtonen, Samuli	Keskiniva, Venni, Tuomisto, Hanna, Lehtonen, Samuli (2024): Danaea (Marattiaceae) keeps diversifying, part 2: phylogeny and identification key for 81 taxa. Willdenowia 53 (3): 229-255, DOI: 10.3372/wi.53.53304, URL: https://www.mendeley.com/catalogue/e926cf5a-1684-366f-8354-946d48c7e673/
